138 lines
17 KiB
XML
138 lines
17 KiB
XML
<document id="86A12E6AE70253E019F875004BE0273F" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A0041FFC4FF182C521682361D" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus panamensis Perkins 2021, new species" docType="treatment" docVersion="7" lastPageNumber="470" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="470" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="52B29EB33501F632D613AEB365597A95">
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<mods:title id="7B69B096C3BA02AA26628F0E4BCE7AA3">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="78A1DC24ABDEAF91B06224159E4D39E4" type="personal">
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<mods:namePart id="6782CE0F6372B204C0EDA7C7CA01B7AF">Perkins, Philip D.</mods:namePart>
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<mods:title id="FBB416BB066F8B69712420312055F954">Zootaxa</mods:title>
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<mods:part id="D110D65EC4C8253B7822C52CE30E8E78">
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<mods:date id="26A4E1E66889EFE403737FD3C478771B">2021</mods:date>
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<mods:number id="52078EFC30ED596AEB176D883BDCF6A6">2021-05-21</mods:number>
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<mods:number id="F00D12E9D293B94ECEB587A4DD5E7F19">4974</mods:number>
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<mods:identifier id="69836CF2BCC05230B3823810AB7D8864" type="CLB-Dataset">6214</mods:identifier>
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<mods:identifier id="1BDF1249E505A7CFF848ECFEC52FF22C" type="DOI">10.11646/zootaxa.4974.3.2</mods:identifier>
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<treatment id="4772526A0041FFC4FF182C521682361D" ID-DOI="http://doi.org/10.5281/zenodo.4777889" ID-GBIF-Taxon="182642574" ID-Zenodo-Dep="4777889" LSID="urn:lsid:plazi:treatment:4772526A0041FFC4FF182C521682361D" httpUri="http://treatment.plazi.org/id/4772526A0041FFC4FF182C521682361D" lastPageNumber="470" pageId="11" pageNumber="470">
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<subSubSection id="87C1B0F70041FFC4FF182C52174F3154" box="[151,594,330,357]" pageId="11" pageNumber="470" type="nomenclature">
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<paragraph id="CF64E37C0041FFC4FF182C52174F3154" blockId="11.[151,594,330,393]" box="[151,594,330,357]" pageId="11" pageNumber="470">
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<heading id="942C54100041FFC4FF182C52174F3154" bold="true" box="[151,594,330,357]" fontSize="11" level="1" pageId="11" pageNumber="470" reason="1">
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<emphasis id="FDAF3F6E0041FFC4FF182C52174F3154" bold="true" box="[151,594,330,357]" pageId="11" pageNumber="470">
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<taxonomicName id="08DB98FF0041FFC4FF182C5214AE3155" ID-CoL="8Y4QX" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,435,330,357]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="470" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
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<emphasis id="FDAF3F6E0041FFC4FF182C5214AE3155" bold="true" box="[151,435,330,357]" italics="true" pageId="11" pageNumber="470">Hydrochus panamensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="E69C82150041FFC4FE4D2C53174F3154" box="[450,594,330,357]" pageId="11" pageNumber="470" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70041FFC4FF182C77142A31B9" box="[151,311,367,393]" pageId="11" pageNumber="470" type="description">
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<paragraph id="CF64E37C0041FFC4FF182C77142A31B9" blockId="11.[151,594,330,393]" box="[151,311,367,393]" pageId="11" pageNumber="470">
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<figureCitation id="57E0FFF90041FFC4FF182C7715F831B9" box="[151,229,367,393]" captionStart="FIGURE 2" captionStartId="5.[151,250,1989,2013]" captionTargetBox="[181,1393,181,1953]" captionTargetId="figure-15@5.[158,1429,181,1965]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777847" httpUri="https://zenodo.org/record/4777847/files/figure.png" pageId="11" pageNumber="470">Figs. 2</figureCitation>
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,
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<figureCitation id="57E0FFF90041FFC4FF7E2C77141231B9" box="[241,271,367,393]" captionStart="FIGURES 11–12" captionStartId="27.[151,264,1944,1968]" captionTargetBox="[169,1429,189,1901]" captionTargetId="figure-15@27.[151,1435,181,1920]" captionTargetPageId="27" captionText="FIGURES 11–12. Habitus and male genitalia of holotypes. 11. Hydrochus perpunctatus n. sp. 12. Hydrochus panamensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777863" httpUri="https://zenodo.org/record/4777863/files/figure.png" pageId="11" pageNumber="470">12</figureCitation>
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,
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<figureCitation id="57E0FFF90041FFC4FE942C77142A31B9" box="[283,311,367,393]" captionStart="FIGURES 28–30" captionStartId="35.[151,264,1944,1968]" captionTargetBox="[158,1429,181,1920]" captionTargetId="figure-15@35.[158,1429,181,1920]" captionTargetPageId="35" captionText="FIGURES 28–30. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777904" httpUri="https://zenodo.org/record/4777904/files/figure.png" pageId="11" pageNumber="470">28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70041FFC4FF182CAF118B31E1" box="[151,1174,438,466]" pageId="11" pageNumber="470" type="materials_examined">
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<paragraph id="CF64E37C0041FFC4FF182CAF118B31E1" blockId="11.[151,1437,438,1581]" box="[151,1174,438,466]" pageId="11" pageNumber="470">
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<emphasis id="FDAF3F6E0041FFC4FF182CAF145831E0" bold="true" box="[151,325,438,465]" pageId="11" pageNumber="470">Type material.</emphasis>
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<materialsCitation id="7FB3E9210041FFC4FEC32CAF118B31E1" ID-GBIF-Occurrence="3128194534" box="[332,1174,439,466]" collectionCode="NMNH" country="Trinidad and Tobago" location="Riv" municipality="August Busck" pageId="11" pageNumber="470" specimenCount="1" typeStatus="holotype">
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<typeStatus id="10605DDE0041FFC4FEC32CAF14A931E1" box="[332,436,439,465]" pageId="11" pageNumber="470" type="holotype">Holotype</typeStatus>
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(male): “
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<collectingCountry id="B7CCA3EC0041FFC4FD932CAF176231E1" box="[540,639,439,465]" name="Trinidad and Tobago" pageId="11" pageNumber="470">Trinidad</collectingCountry>
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<location id="CA04B5A70041FFC4FD092CAF17B231E1" LSID="urn:lsid:plazi:treatment:4772526A0041FFC4FF182C521682361D:CA04B5A70041FFC4FD092CAF17B231E1" box="[646,687,439,465]" country="Trinidad and Tobago" municipality="August Busck" name="Riv" pageId="11" pageNumber="470">Riv</location>
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, Pan. May.21.11,
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<collectingMunicipality id="2F0079060041FFC4FCFA2CAF110B31E1" box="[885,1046,439,465]" pageId="11" pageNumber="470">August Busck</collectingMunicipality>
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” (
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<collectionCode id="A9CA7BB90041FFC4FBA12CAF119731E1" box="[1070,1162,439,465]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="470" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70041FFC4FF482CC211773299" pageId="11" pageNumber="470" type="diagnosis">
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<paragraph id="CF64E37C0041FFC4FF482CC211773299" blockId="11.[151,1437,438,1581]" pageId="11" pageNumber="470">
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<emphasis id="FDAF3F6E0041FFC4FF482CC214CC31C4" bold="true" box="[199,465,474,500]" pageId="11" pageNumber="470">Differential diagnosis.</emphasis>
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Among Central American species, recognized by the combination of large size (ca.
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<quantity id="08234E990041FFC4FF182CE615E33229" box="[151,254,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" pageId="11" pageNumber="470" unit="mm" value="3.95">3.95 mm</quantity>
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), the deep pronotal impressions, the relatively impunctate pronotal reliefs, the elytra with prominent callus on 5
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<superScript id="38AE4E340041FFC4FF462F3A15CB3200" attach="left" box="[201,214,546,560]" fontSize="6" pageId="11" pageNumber="470">th</superScript>
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interval and raised area of 4
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<superScript id="38AE4E340041FFC4FD962F3A173B3200" attach="left" box="[537,550,546,560]" fontSize="6" pageId="11" pageNumber="470">th</superScript>
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interval, and the male genitalia (
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<figureCitation id="57E0FFF90041FFC4FC142F3B16F2320D" box="[923,1007,547,573]" captionStart="FIGURES 11–12" captionStartId="27.[151,264,1944,1968]" captionTargetBox="[169,1429,189,1901]" captionTargetId="figure-15@27.[151,1435,181,1920]" captionTargetPageId="27" captionText="FIGURES 11–12. Habitus and male genitalia of holotypes. 11. Hydrochus perpunctatus n. sp. 12. Hydrochus panamensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777863" httpUri="https://zenodo.org/record/4777863/files/figure.png" pageId="11" pageNumber="470">Fig. 12</figureCitation>
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; described below). The male genitalia have a narrow, parallel-sided basal piece, which is narrower than the width of the genitalia at the proximal part of the parameres; these characters plus the shape of the paramere tips serve to differentiate the form from that of other Central American species. This is the largest known Panamanian species of
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<taxonomicName id="08DB98FF0041FFC4FC662F97117E3299" authorityName="Leach" authorityYear="1817" box="[1001,1123,655,681]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="470" phylum="Arthropoda" rank="genus">
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<emphasis id="FDAF3F6E0041FFC4FC662F97117E3299" box="[1001,1123,655,681]" italics="true" pageId="11" pageNumber="470">Hydrochus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70041FFC4FF482FAA146C3639" pageId="11" pageNumber="470" type="description">
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<paragraph id="CF64E37C0041FFC4FF482FAA146533B1" blockId="11.[151,1437,438,1581]" pageId="11" pageNumber="470">
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<emphasis id="FDAF3F6E0041FFC4FF482FAA144C32FC" bold="true" box="[199,337,690,716]" pageId="11" pageNumber="470">Description</emphasis>
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. Habitus as illustrated (
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<figureCitation id="57E0FFF90041FFC4FDD62FAB17B132FD" box="[601,684,691,717]" captionStart="FIGURES 11–12" captionStartId="27.[151,264,1944,1968]" captionTargetBox="[169,1429,189,1901]" captionTargetId="figure-15@27.[151,1435,181,1920]" captionTargetPageId="27" captionText="FIGURES 11–12. Habitus and male genitalia of holotypes. 11. Hydrochus perpunctatus n. sp. 12. Hydrochus panamensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777863" httpUri="https://zenodo.org/record/4777863/files/figure.png" pageId="11" pageNumber="470">Fig. 12</figureCitation>
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). Size:
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<typeStatus id="10605DDE0041FFC4FD712FAB167D32FD" box="[766,864,691,717]" pageId="11" pageNumber="470" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 3.94/1.42; head width 0.94; pronotum l/w 0.90/0.90; PA 0.87; PB 0.81; elytra 2.52/1.42. Dorsum dark grey, with slight to moderately strong iridescence on head and pronotum, elytra with indistinct black areas. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 1–1.5× strial puncture diameter. Interstria 9
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<superScript id="38AE4E340041FFC4FB302E0611D1331C" attach="left" box="[1215,1228,798,812]" fontSize="6" pageId="11" pageNumber="470">th</superScript>
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more convex than others, overhanging 10
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<superScript id="38AE4E340041FFC4FE162E5A14BB3360" attach="left" box="[409,422,834,848]" fontSize="6" pageId="11" pageNumber="470">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="38AE4E340041FFC4FCB82E5A16593360" attach="left" box="[823,836,834,848]" fontSize="6" pageId="11" pageNumber="470">th</superScript>
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strongly raised, moderately elongate, ca. 5 punctures in row on each side.
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</paragraph>
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<paragraph id="CF64E37C0041FFC4FF482E9316BB33DD" blockId="11.[151,1437,438,1581]" pageId="11" pageNumber="470">Pronotum length and width equal, widest at anterior margin, narrowed at base, sides arcuate, each with unilinear row of minute granules; anterior margin slightly bisinuate; depressions deep, much more densely punctate than distinct reliefs between depressions, latter almost entirely impunctate.</paragraph>
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<paragraph id="CF64E37C0041FFC4FF482EEF165B3469" blockId="11.[151,1437,438,1581]" pageId="11" pageNumber="470">
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Elytra with spaces between strial punctures ca. 0.5–1x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin without subapical angulation. 4
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<superScript id="38AE4E340041FFC4FC54290216F53418" attach="left" box="[987,1000,1050,1064]" fontSize="6" pageId="11" pageNumber="470">th</superScript>
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interval notably raised from about elytra midlength to level of anterior part of callus on 5
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<superScript id="38AE4E340041FFC4FD5A292617FF347C" attach="left" box="[725,738,1086,1100]" fontSize="6" pageId="11" pageNumber="470">th</superScript>
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interval.
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</paragraph>
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<paragraph id="CF64E37C0041FFC4FF48297B17513491" blockId="11.[151,1437,438,1581]" pageId="11" pageNumber="470">Ventral characters: Mentum moderately coarsely punctate, with large, deep median fovea. Submentum with two very shallow foveae. Legs dark brown.</paragraph>
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<paragraph id="CF64E37C0041FFC4FF4829B3174D3549" blockId="11.[151,1437,438,1581]" pageId="11" pageNumber="470">
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Male genitalia (
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<figureCitation id="57E0FFF90041FFC4FEF529B314D134F5" box="[378,460,1195,1221]" captionStart="FIGURES 11–12" captionStartId="27.[151,264,1944,1968]" captionTargetBox="[169,1429,189,1901]" captionTargetId="figure-15@27.[151,1435,181,1920]" captionTargetPageId="27" captionText="FIGURES 11–12. Habitus and male genitalia of holotypes. 11. Hydrochus perpunctatus n. sp. 12. Hydrochus panamensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777863" httpUri="https://zenodo.org/record/4777863/files/figure.png" pageId="11" pageNumber="470">Fig. 12</figureCitation>
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) general characters: paramere tips slender and pointed, much narrower in lateral view than in dorsal/ventral views, in dorsal/ventral views with medial margins at apex parallel or nearly so, then arcuate away from midline; aedeagus lanceolate, moderately slender in dorsal/ventral views; in lateral view slightly wider basally; lateral margins of aedeagus sclerotized; basal piece slightly shorter than parameres, narrow, much narrower than greatest width of distal pieces, in dorsal/ventral views almost parallel sided, slightly arcuate in lateral view, orifice moderately strongly sclerotized.
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</paragraph>
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<paragraph id="CF64E37C0041FFC4FF48289B145835F1" blockId="11.[151,1437,438,1581]" pageId="11" pageNumber="470">Dorsal surface: pdmm sinuate, medial part slightly overlapping aedeagus; adbl moderately large; adtl slightly past midlength.</paragraph>
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<paragraph id="CF64E37C0041FFC4FF4828D3146C3639" blockId="11.[151,1437,438,1581]" pageId="11" pageNumber="470">Ventral surface: pvmm more widely separated than pdmm, approximating lateral margin of aedeagus; avtl very near base; gp large.</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70041FFC4FF482B0B1682361D" box="[199,927,1555,1581]" pageId="11" pageNumber="470" type="etymology">
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<paragraph id="CF64E37C0041FFC4FF482B0B1682361D" blockId="11.[151,1437,438,1581]" box="[199,927,1555,1581]" pageId="11" pageNumber="470">
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<emphasis id="FDAF3F6E0041FFC4FF482B0B1453361D" bold="true" box="[199,334,1555,1581]" pageId="11" pageNumber="470">Etymology.</emphasis>
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Named in reference to the geographical distribution.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |