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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.413.7172" ID-GBIF-Dataset="f45c1ebe-039d-4180-bfd0-a9d772f72281" ID-PMC="PMC4086027" ID-Pensoft-Pub="1313-2970-413-1" ID-PubMed="25009414" ID-ZBK="5791CE9C1CC0472095838A585DA79446" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-413-1" ModsDocOrigin="ZooKeys 413" ModsDocTitle="The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups" checkinTime="1451245833965" checkinUser="pensoft" docAuthor="Hita Garcia, Francisco &amp; Fisher, Brian L." docDate="2014" docId="374487C6A19E956204C0760E2886CB99" docLanguage="en" docName="ZooKeys 413: 1-170" docOrigin="ZooKeys 413" docSource="http://dx.doi.org/10.3897/zookeys.413.7172" docTitle="Tetramorium bressleri Hita Garcia &amp; Fisher, sp. n." docType="treatment" docUuid="288AA21B-2B9E-49F1-8C6A-B22E6FD4619E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="29" masterDocId="FFFAFFE7FFB3C93AB161C804401EFFC2" masterDocTitle="The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups" masterLastPageNumber="170" masterPageNumber="1" pageNumber="25" updateTime="1668158573061" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hita Garcia, Francisco</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>413</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>170</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.413.7172</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.413.7172</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-413-1</mods:identifier>
<mods:identifier type="ZBK">5791CE9C1CC0472095838A585DA79446</mods:identifier>
<mods:identifier type="ZooBank">5791CE9C1CC0472095838A585DA79446</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152053316" LSID="urn:lsid:zoobank.org:act:288AA21B-2B9E-49F1-8C6A-B22E6FD4619E" httpUri="http://treatment.plazi.org/id/374487C6A19E956204C0760E2886CB99" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25">
<subSubSection pageId="24" pageNumber="25" type="nomenclature">
<paragraph pageId="24" pageNumber="25">
<taxonomicName LSID="http://zoobank.org/288AA21B-2B9E-49F1-8C6A-B22E6FD4619E" authority="Hita Garcia &amp; Fisher" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri Hita Garcia &amp; Fisher</taxonomicName>
<taxonomicNameLabel pageId="24" pageNumber="25">sp. n.</taxonomicNameLabel>
Figs 10E, 11C, 12C, 13C, 14B, 15, 62
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="type material">
<paragraph pageId="24" pageNumber="25">Type material.</paragraph>
<paragraph pageId="24" pageNumber="25">
Holotype, pinned worker, MADAGASCAR, Mahajanga, Parc National de Namoroka, 9.8 km 300° WNW Vilanandro,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-16.46667">16.46667°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="45.35">45.35°E</geoCoordinate>
, 140 m, tropical dry forest, sifted litter (leaf mold, rotten wood), collection code BLF06446, 4.-8.XI.2002 (B.L. Fisher et al.) (CAS: CASENT0035677). Paratypes, 27 pinned workers with same data as holotype (BMNH: CASENT0035669; CAS: CASENT0035636; CASENT0035640; CASENT0035642; CASENT0035647; CASENT0035648; CASENT0035649; CASENT0035652; CASENT0035653; CASENT0035654; CASENT0035656; CASENT0035658; CASENT0035660; CASENT0035662; CASENT0035666; CASENT0035667; CASENT0035670; CASENT0035674; CASENT0035675; CASENT0035678; CASENT0035681; CASENT0035686; CASENT0035687; CASENT0035689; CASENT0035694; CASENT0035703; MCZ: CASENT0035646).
</paragraph>
<caption pageId="24" pageNumber="25">
<paragraph pageId="24" pageNumber="25">
Figure 15.
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
holotype worker (CASENT0035677). A Body in profile B Body in dorsal view C Head in full-face view.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="non-type material">
<paragraph pageId="24" pageNumber="25">Non-type material.</paragraph>
<paragraph lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
MADAGASCAR: Antananarivo, Reserve Speciale
<normalizedToken originalValue="dAmbohitantely">d'Ambohitantely</normalizedToken>
,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
Ambohitantely, 20.9 km 72°NE Ankazobe,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-18.2253">18.2253°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="47.2868">47.2868°E</geoCoordinate>
, 1410 m, montane rainforest, 17.-22.IV.2001 (
<pageBreakToken pageId="25" pageNumber="26" start="start">B</pageBreakToken>
.L. Fisher et al.); Antsiranana,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAnabohazo">d'Anabohazo</normalizedToken>
, 21.6 km 247° WSW Maromandia,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-14.3089">14.3089°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="47.9143">47.9143°E</geoCoordinate>
, 120 m, tropical dry forest, 11.-16.III.2001 (B.L. Fisher et al.); Fianarantsoa, Ampangabe I Non Protected Area, 21.4 km W Itremo,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-20.6111">20.6111°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="46.6069">46.6069°E</geoCoordinate>
, 1414 m, savannah woodland, 21.-23.III.2010 (A. Ravelomanana); Fianarantsoa, Ampangabe II Non Protected Area, 21.29 km W Itremo,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-20.6114">20.6114°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="46.6081">46.6081°E</geoCoordinate>
, 1402 m, savannah woodland, 21.-23.III.2010 (A. Ravelomanana); Fianarantsoa, Ampangabe VI Non Protected Area, 21.16 km W Itremo,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-20.6144">20.6144°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="46.6104">46.6104°E</geoCoordinate>
, 1379 m, shrubland, 21.-23.III.2010 (A. Ravelomanana); Fianarantsoa, Ampangoabe V Non Protected Area, 21.37 km W Itremo,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-20.6136">20.6136°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="46.608">46.608°E</geoCoordinate>
, 1449 m, shrubland, 22.-23.III.2010 (A. Ravelomanana); Fianarantsoa, Ampotoampoto I National Parc, 8.02 km NW Ilakaka,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.6283">22.6283°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.1886">45.1886°E</geoCoordinate>
, 917 m, savannah woodland, 26.-28.II.2010 (A. Ravelomanana); Fianarantsoa,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAnalalava">d'Analalava</normalizedToken>
, 29.6 km 280° W Ranohira,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.5917">22.5917°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.1283">45.1283°E</geoCoordinate>
, 700 m, 1.-5.II.2003 (B.L. Fisher et al.); Fianarantsoa, Antohatsahomby I Non Protected Area, 22.77 km NW Ambatofinandrahana,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-20.5506">20.5506°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="46.5856">46.5856°E</geoCoordinate>
, 1550 m,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">Uapaca</taxonomicName>
woodland, 15.-17.III.2010 (A. Ravelomanana); Fianarantsoa,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAtsirakambiaty">d'Atsirakambiaty</normalizedToken>
, 7.6 km 285° WNW Itremo,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-20.5933">20.5933°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="46.5633">46.5633°E</geoCoordinate>
, 1550 m, montane rainforest, 22.-26.I.2003 (B.L. Fisher et al.); Fianarantsoa, Parc National
<normalizedToken originalValue="dIsalo">d'Isalo</normalizedToken>
, 9.1 km 354° N Ranohira,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.4817">22.4817°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.4617">45.4617°E</geoCoordinate>
, 725 m, gallery forest, 27.-31.I.2003 (B.L. Fisher et al.); Fianarantsoa, Parc National
<normalizedToken originalValue="dIsalo">d'Isalo</normalizedToken>
, Ambovo Springs, 29.3 km 4° N Ranohira,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.2983">22.2983°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.3517">45.3517°E</geoCoordinate>
, 990 m,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">Uapaca</taxonomicName>
woodland, 9.-14.II.2003 (B.L. Fisher et al.); Fianarantsoa, Mampiarika I Non Protected Area, 28.08 km SW Ambositra,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-20.7344">20.7344°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="47.0835">47.0835°E</geoCoordinate>
, 1480 m,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">Uapaca</taxonomicName>
woodland, 31.I.-2.II.2010 (A. Ravelomanana); Mahajanga,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
Ambohimanga, 26.1 km 314° Mampikony,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-15.9627">15.9627°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="47.4382">47.4382°E</geoCoordinate>
, 250 m, tropical dry forest, 13.-15.XII.2004 (B.L. Fisher); Mahajanga, Boeny Region, District of Soalala, Analamanitra forest, 14 km SW Mitsinjo,
<geoCoordinate direction="south" orientation="latitude" precision="5555" value="-16.7">16.7°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5555" value="45.7">45.7°E</geoCoordinate>
, 19 m, dense dry forest, 26.II.-4.III.2008 (M.
<normalizedToken originalValue="Rinha">Rin'ha</normalizedToken>
); Mahajanga, Parc National
<normalizedToken originalValue="dAnkarafantsika">d'Ankarafantsika</normalizedToken>
, Ampijoroa Station Forestiere, 40 km 306° NW Andranofasika,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-16.3208">16.3208°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="46.8107">46.8107°E</geoCoordinate>
, 130 m, tropical dry forest, 26.III.-1.IV.2001 (B.L. Fisher et al.); Mahajanga, Parc National
<normalizedToken originalValue="dAnkarafantsika">d'Ankarafantsika</normalizedToken>
, Ampijoroa Station Forestiere, 5.4 km 331° NW Andranofasika,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-16.2989">16.2989°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="46.813">46.813°E</geoCoordinate>
, 70 m, tropical dry forest, 26.III.-1.IV.2001 (B.L. Fisher et al.); Mahajanga, Parc National
<normalizedToken originalValue="dAnkarafantsika">d'Ankarafantsika</normalizedToken>
,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
de Tsimaloto, 18.3 km 46° NE de Tsaramandroso,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-16.2281">16.2281°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="47.1436">47.1436°E</geoCoordinate>
, 135 m, tropical dry forest, 2.IV.-8.IV.2001 (B.L. Fisher et al.); Mahajanga, Parc National de Baie de Baly, 12.4 km 337° NNW Soalala,
<geoCoordinate direction="south" orientation="latitude" precision="555" value="-16.01">16.01°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="45.265">45.265°E</geoCoordinate>
, 10 m, tropical dry forest, 26.-30.XI.2002 (B.L. Fisher et al.); Mahajanga, Reserve Forestiere Beanka, 50.2 km E Maintirano,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-18.0265">18.0265°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.0505">44.0505°E</geoCoordinate>
, 250 m, tropical dry forest on tsingy, 19.-26.X.2009 (B.L. Fisher et al.); Mahajanga, Reserve Forestiere Beanka, 52.7 km E Maintirano,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-18.0622">18.0622°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.5259">44.5259°E</geoCoordinate>
, 300 m, tropical dry forest on tsingy, 24.-27.X.2009 (B.L. Fisher et al.); Mahajanga, Reserve Forestiere Beanka, 50.7 km E Maintirano,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-17.8802">17.8802°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.4688">44.4688°E</geoCoordinate>
, 140 m, tropical dry forest on tsingy, 29.X.-1.XI.2009 (B.L. Fisher et al.); Mahajanga, Reserve Forestiere Beanka, 50.2 km E Maintirano,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-17.8876">17.8876°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.4726">44.4726°E</geoCoordinate>
, 153 m, tropical dry forest on tsingy, 31.X.2009 (B.L. Fisher et al.); Mahajanga, Reserve Speciale de Bemarivo, 23.8 km 223° SW Besalampy,
<geoCoordinate direction="south" orientation="latitude" precision="55" value="-16.925">16.925°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.3683">44.3683°E</geoCoordinate>
, 30 m, tropical dry forest, 19.-23.XI.2002 (B.L. Fisher et al.); Mahajanga, Mahavavy River, 6.2 km 145° SE Mitsinjo,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-16.0517">16.0517°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.9083">45.9083°E</geoCoordinate>
, 20 m, gallery forest, 1.-5.XII.2002 (B.L. Fisher et al.); Mahajanga, Parc National de Namoroka, 9.8 km 300° WNW Vilanandro,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-16.4667">16.4667°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="45.35">45.35°E</geoCoordinate>
, 140 m, tropical dry forest, 4.-8.XI.2002 (B.L. Fisher et al.); Mahajanga, Parc National de Namoroka, 17.8 km 329° WNW Vilanandro,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-16.3767">16.3767°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.3267">45.3267°E</geoCoordinate>
, 100 m, tropical dry forest, 8.-12.XI.2002 (B.L. Fisher et al.); Mahajanga, Parc National de Namoroka, 16.9 km 317° NW Vilanandro,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-16.4067">16.4067°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="45.31">45.31°E</geoCoordinate>
, 100 m, tropical dry forest, 12.-16.XI.2002 (B.L. Fisher et al.); Mahajanga, Parc National Tsingy de Bemaraha, 3.4 km 93° E Bekopaka, Tombeau Vazimba,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-19.1419">19.1419°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="44.828">44.828°E</geoCoordinate>
, 50 m, tropical dry forest, 6.-10.XI.2001 (B.L. Fisher et al.); Mahajanga, Parc National Tsingy de Bemaraha, 2.5 km 62° ENE Bekopaka, Ankidrodroa River,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-19.1322">19.1322°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.8147">44.8147°E</geoCoordinate>
, 100 m, 11.-15.XI.2001 (B.L. Fisher et al.); Mahajanga, Parc National Tsingy de Bemaraha, 10.6 km ESE 123° Antsalova,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-18.7094">18.7094°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.7182">44.7182°E</geoCoordinate>
, 150 m, tropical dry forest on tsingy, 16.-20.XI.2001 (B.L. Fisher et al.); Toliara, Parc National
<normalizedToken originalValue="dAndohahela">d'Andohahela</normalizedToken>
,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAmbohibory">d'Ambohibory</normalizedToken>
, 1.7 km 61° ENE Tsimelahy, 36.1 km 308° NW Tolagnaro,
<geoCoordinate direction="south" orientation="latitude" precision="555" value="-24.93">24.93°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="46.6455">46.6455°E</geoCoordinate>
, 300 m, tropical dry forest, 16.-20.I.2002 (B.L. Fisher et al.); Toliara, Andohahela National Park, Tsimelahy,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-24.9368">24.9368°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="46.6267">46.6267°E</geoCoordinate>
, 180 m, transition forest, 16.-26.II.2003 (M.E. Irwin et al.); Toliara, 18 km NNW Betroka,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-23.1633">23.1633°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.9686">45.9686°E</geoCoordinate>
, 825 m, savannah, 24.XI.-4.XII.1994 (M.A. Ivie &amp; D. A. Pollock); Toliara, Atsimo Andrefana Region, District of Betioky, 30 km E Betioky, Beza Mahafaly Special Reserve,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-23.6865">23.6865°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="44.591">44.591°E</geoCoordinate>
, 165 m, gallery dry deciduous forest, 1.-7.I.2002 (M.
<normalizedToken originalValue="Rinha">Rin'ha</normalizedToken>
); Toliara, Beza-Mahafaly, 27 km E Betioky,
<geoCoordinate direction="south" orientation="latitude" precision="555" value="-23.65">23.65°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.6333">44.6333°E</geoCoordinate>
, 135 m, tropical dry forest, 23.IV.1997 (B.L. Fisher); Toliara, Fiherenana,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-23.1769">23.1769°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="43.9608">43.9608°E</geoCoordinate>
, 100 m, gallery forest, 21.-24.X.2002 (Frontier Project); Toliara, Fiherenana,
<geoCoordinate direction="south" orientation="latitude" precision="55" value="-23.177">23.177°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="43.9614">43.9614°E</geoCoordinate>
, gallery forest, 18.-19.VII.2003 (Frontier Wilderness Project); Toliara, southern Isoky-Vohimena Forest, 59 km NE Sakaraha,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.4667">22.4667°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="44.85">44.85°E</geoCoordinate>
, 730 m, tropical dry forest, 21.I.1996 (B.L. Fisher); Toliara, Makay Mts.,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-21.2098">21.2098°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.3418">45.3418°E</geoCoordinate>
, 525 m, gallery forest on sandy soil, 27.XI.-2.XII.2010 (B.L. Fisher et al.); Toliara, Makay Mts.,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-21.2199">21.2199°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="45.324">45.324°E</geoCoordinate>
, 500 m, gallery forest on sandy soil, 24.XI.-12.XII.2010 (B.L. Fisher et al.); Toliara, Makay Mts.,
<geoCoordinate direction="south" orientation="latitude" precision="555" value="-21.31">21.31°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="45.1295">45.1295°E</geoCoordinate>
, 590 m, dry forest on sandy soil, 3.-6.XII.2010 (B.L. Fisher et al.); Toliara, Vohibasia Forest, 59 km NE Sakaraha,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.4667">22.4667°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="44.85">44.85°E</geoCoordinate>
, 780 m, tropical dry forest, 13.I.1996 (B.L. Fisher); Toliara, Parc National de Zombitse, near road,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.8405">22.8405°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.7312">44.7312°E</geoCoordinate>
, 825 m, spiny deciduous forest, 15.X.2001-23.III.2002 (R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
); Toliara, Parc National de Zombitse, near ANGAP office,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.8865">22.8865°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="44.6922">44.6922°E</geoCoordinate>
, 840 m, deciduous spiny forest, 9.XI.2001-26.I.2002 (R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
); Toliara, Parc National de Zombitse, 19.8 km 84° E Sakaraha,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-22.8433">22.8433°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="44.71">44.71°E</geoCoordinate>
, 770 m, tropical dry forest, 5.-9.II.2003 (B.L. Fisher et al.).
</paragraph>
</subSubSection>
<subSubSection lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="diagnosis">
<paragraph pageId="25" pageNumber="26">Diagnosis.</paragraph>
<paragraph pageId="26" pageNumber="27">
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bressleri">
<pageBreakToken pageId="26" pageNumber="27" start="start">Tetramorium</pageBreakToken>
bressleri
</taxonomicName>
can be recognised by the following combination of characters: larger species (HW 0.80-1.00; WL 0.92-1.15); eyes relatively small (OI 18-19); petiolar node high nodiform, not blocky and massively enlarged, anterodorsal and posterodorsal margins at about the same height and equally marginate, anterodorsal margin not protruding anteriorly nor very sharply angled; petiolar node in profile relatively high and thin, between 1.6 to 1.8 times higher than long (LPeI 56-61), in dorsal view between 1.3 to 1.5 times wider than long (DPeI 135-145); gaster never extremely enlarged and swollen; head and mesosoma without strongly developed and conspicuous reticulate-punctate ground sculpture; usually sculpture on the mesopleuron and lateral propodeum mostly absent; basal half of first gastral tergite not strongly reticulate-rugose, only base of tergite weakly sculptured; pilosity on first gastral tergite mostly erect.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="worker measurements">
<paragraph pageId="26" pageNumber="27">Worker measurements</paragraph>
<paragraph pageId="26" pageNumber="27">(N=12). HL 0.81-1.00 (0.94); HW 0.80-1.00 (0.94); SL 0.51-0.64 (0.60); EL 0.15-0.19 (0.18); PH 0.42-0.53 (0.48); PW 0.57-0.73 (0.68); WL 0.92-1.15 (1.07); PSL 0.24-0.32 (0.27); PTL 0.20-0.26 (0.23); PTH 0.35-0.42 (0.40); PTW 0.28-0.36 (0.32); PPL 0.23-0.30 (0.27); PPH 0.32-0.41 (0.38); PPW 0.35-0.45 (0.42); CI 98-102 (100); SI 62-65 (64); OI 18-19 (19); DMI 61-66 (64); LMI 43-47 (45); PSLI 26-32 (28); PeNI 45-49 (47); LPeI 56-61 (58); DPeI 135-145 (138); PpNI 59-63 (61); LPpI 70-75 (73); DPpI 150-158 (152); PPI 125-134 (130).</paragraph>
</subSubSection>
<subSubSection lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="worker description">
<paragraph pageId="26" pageNumber="27">Worker description.</paragraph>
<paragraph lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Head more or less as long as broad (CI 98-102); posterior head margin weakly concave. Anterior clypeal margin with distinct median impression. Frontal carinae strongly developed and forming dorsal margin of very well-developed antennal scrobes, scrobes moderately to very deep and with clearly defined margins all around; median scrobal carina very well developed and distinctly surpassing posterior eye level, usually ending halfway between posterior eye margin and posterior scrobe margin. Antennal scapes short, not reaching posterior head margin (SI 62-65). Eyes relatively small (OI 18-19). Mesosomal outline in profile weakly to moderately convex, rounded and high (LMI 43-47), moderately to strongly marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent. Propodeal spines elongate-triangular to spinose, long, and acute (PSLI 26-32), propodeal lobes short, triangular, and acute, always much shorter than propodeal spines. Petiolar node in profile high, rectangular nodiform with well defined antero- and posterodorsal margins, between 1.6 to 1.8 times higher than long (LPeI 56-61), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins situated at about the same height, petiolar dorsum flat to weakly convex, anterodorsal margin not protruding anteriorly; node in dorsal view between 1.3 to 1.5 times wider than long (DPeI 135-145), in dorsal view pronotum between 2.0 to 2.2 times wider than petiolar node (PeNI 45-49). Postpetiole in profile subglobular and weakly anteroposteriorly compressed, approximately 1.3 to 1.4 times higher than long (LPpI 70-75); in dorsal view around 1.5 to 1.6 times wider than long (DPpI 150-158), pronotum between 1.6 to 1.7 times wider than postpetiole (PpNI 59-63). Postpetiole in profile appearing slightly less voluminous than petiolar node, postpetiole in dorsal view between 1.2 to 1.4 times wider than petiolar node (PPI 125-134). Mandibles variably sculptured, either unsculptured, smooth, and shining, or partly or fully finely rugulose; clypeus longitudinally rugose/rugulose, with five to eight distinct rugae, median ruga always distinct, lateral rugae often weaker and sometimes interrupted; cephalic dorsum between frontal carinae with ten to thirteen longitudinal rugae, rugae running from posterior clypeal margin to posterior head margin, rarely interrupted or with cross-meshes; scrobal area mostly unsculptured, smooth and shiny; lateral head mainly longitudinally rugose. Ground sculpture on head usually weak to absent. Dorsum of mesosoma mostly longitudinally rugose without any distinct ground sculpture, spaces between rugae smooth and shining; lateral pronotum irregularly longitudinally rugose, often with weak to moderate punctate ground sculpture, mesopleuron and lateral pro
<pageBreakToken pageId="27" pageNumber="28" start="start">podeum</pageBreakToken>
usually with very little sculpture, few irregular rugae/rugulae and no ground sculpture. Forecoxae often with weak ground sculpture, but generally very smooth and shining. Petiolar node laterally with conspicuous but relatively weak reticulate-punctate ground sculpture only, appearing weakly matte but still relatively smooth and shiny; dorsum of node medially almost unsculptured, smooth, and shiny, surrounding areas rugulose, ground sculpture on petiolar dorsum neglectable. Postpetiole laterally and dorsally weakly to moderately rugose/rugulose and with conspicuous, moderate reticulate-punctate ground sculpture, appearing relatively matte; base of first gastral tergite usually weakly punctate and/or costulate and/or shagreened, remainder of tergite unsculptured, smooth, and shining. Whole body with abundant, long, and fine standing hairs; first gastral tergite with abundant, long, erect hairs and much scarcer, shorter, decumbent to subdecumbent hairs. Anterior edges of antennal scapes and dorsal (outer) surfaces of hind tibiae with decumbent to suberect hairs. Body usually of uniform brown, appendages often lighter.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="etymology">
<paragraph pageId="27" pageNumber="28">Etymology.</paragraph>
<paragraph pageId="27" pageNumber="28">The name of the new species is a patronym dedicated to Dr. Barry Lee Bressler, retired physicist, former adjunct professor of physics at Virginia Polytechnic Institute and State University, and amateur naturalist, in recognition of his interest in myrmecology and his support of ant taxonomy.</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="distribution and biology">
<paragraph pageId="27" pageNumber="28">Distribution and biology.</paragraph>
<paragraph pageId="27" pageNumber="28">
As mentioned above, it is surprising that most species in the species group, except the holotype of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
, were previously unknown. This is especially true for
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
. It is by far the most common and abundant species of the
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
group. The material available was sampled in many localities and includes more than 500 mounted specimens with many more in alcohol. The distribution ranges of all group members strongly overlap, but the range of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
is by far the largest; this species is known from many more localities than the other four (Fig. 62). The southernmost localities are Andohahela, Beza Mahafaly, and Fiherenana, and from there the distribution ranges north through much of western Madagascar up to the northernmost known locality Anabohazo. The eastern limit of the range goes in an almost straight line north from Andohahela through Mampiarika, Ambohitantely, and Ambohimanga to Anabohazo. The new species prefers arid habitats such as tropical dry forests, tropical dry forests on tsingy, gallery forests, spiny deciduous forests, savannah woodland,
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uapaca" order="Malpighiales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="genus">Uapaca</taxonomicName>
woodland, and spiny thickets. The elevational range of the species is a relatively broad one, ranging from 10 to 1550 m, but most of the material was collected at low elevations (ca. 420 m on average).
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
was mainly sampled by pitfall trapping and litter sifting, suggesting a ground-active life style.
</paragraph>
</subSubSection>
<subSubSection lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="discussion">
<paragraph pageId="27" pageNumber="28">Discussion.</paragraph>
<paragraph pageId="28" pageNumber="29">
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">
<pageBreakToken pageId="28" pageNumber="29" start="start">Tetramorium</pageBreakToken>
bressleri
</taxonomicName>
is not likely to be confused with
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium gollum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="gollum">Tetramorium gollum</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium hobbit" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hobbit">Tetramorium hobbit</taxonomicName>
, or
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium mars" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mars">Tetramorium mars</taxonomicName>
, whereas differentiating between
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
and
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
can be challenging at first glance.
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
lacks the enlarged gaster and strong reticulate-rugose sculpture on the basal half of the first gastral tergite seen in
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium gollum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="gollum">Tetramorium gollum</taxonomicName>
, nor does it have the blocky petiolar node shape of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium mars" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mars">Tetramorium mars</taxonomicName>
or the massively enlarged petiolar node of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium hobbit" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hobbit">Tetramorium hobbit</taxonomicName>
. The separation from
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
requires more attention and caution. The main and obvious difference is body size.
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
is usually a much larger species (HW 0.80-1.00; WL 0.92-1.15) than
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
(HW 0.80-1.00; WL 0.92-1.15). There is some overlap, but this is mainly due to a few very small specimens of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
and one very large specimen of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
. Otherwise, both species fall neatly into their respective size ranges. However, body size alone should not be used as a primary diagnostic character, and in this case, we provide more evidence for their heterospecificity. The eyes of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
(OI 21-23) are larger than in
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
(OI 18-19), although this is often difficult to assess without measuring. In addition, both can be separated by the sculpture on the mesopleuron and lateral propodeum; this sculpture is usually mostly absent in
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
, making the area appear very smooth and shiny, while it is usually longitudinally rugose with reticulate-punctate ground sculpture in
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
. The sculpture on the sides of the petiolar node varies too, since it is much more reticulate-punctate and matte in
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
than in
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
, in which the weak reticulate-punctate ground sculpture looks faintly matte but still relatively smooth and shiny. The shape of the petiolar node in profile is an additional useful character. In
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium plesiarum" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="plesiarum">Tetramorium plesiarum</taxonomicName>
the anterodorsal margin of the node protrudes anteriorly and is a bit more marginate than the more rounded posterodorsal margin, while in
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
both margins are usually equally marginate and the anterodorsal margin does not protrude anteriorly at all.
</paragraph>
<paragraph pageId="28" pageNumber="29">
Considering how common and abundant
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium bressleri" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bressleri">Tetramorium bressleri</taxonomicName>
is in western Madagascar, it is interesting that it shows very little intraspecific variation and remains very stable in its morphology.
</paragraph>
</subSubSection>
</treatment>
</document>