239 lines
27 KiB
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239 lines
27 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="1E9F3DAD08DA6A93F1D4821C57AD7087" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Coloborhynchus clavirostris Owen 1874" docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="23" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="22" updateTime="1634706563434" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044971" LSID="urn:lsid:plazi:treatment:1E9F3DAD08DA6A93F1D4821C57AD7087" httpUri="http://treatment.plazi.org/id/1E9F3DAD08DA6A93F1D4821C57AD7087" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<subSubSection pageId="21" pageNumber="22" type="nomenclature">
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName LSID="1E9F3DAD-08DA-6A93-F1D4-821C57AD7087" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Coloborhynchus clavirostris" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="clavirostris">Coloborhynchus clavirostris Owen, 1874</taxonomicName>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coloborhynchus clavirostris. Holotype, NHMUK PV R 1822 (late Berriasian / Valanginian, Hastings Group), anterior part of the rostrum A anterior view B respective line drawing C left lateral view D respective line drawing. Abbreviations: dep - depression, m - maxillae, pm - premaxillae, pmcr - premaxillaery crest, prid - palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. Photos courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22075" pageId="21" pageNumber="22">Fig. 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="reference_group">
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="clavirostris">Coloborhynchus clavirostris</taxonomicName>
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Owen:
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<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B67" refString="Owen, R, 1874. Monograph on the fossil Reptilia of the Mesozoic Formations. Palaeontographical Society, London, 14 pp." title="Monograph on the fossil Reptilia of the Mesozoic Formations." year="1874">Owen 1874</bibRefCitation>
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: p. 6, pl. I, fig. 1-4
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Criorhynchidae" genus="Criorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="simus">Criorhynchus simus</taxonomicName>
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(Owen):
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 537
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName class="Reptilia" family="Criorhynchidae" genus="Criorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="clavirostris">Criorhynchus clavirostris</taxonomicName>
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(Owen):
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<bibRefCitation author="Arthaber, G" journalOrPublisher="Palaeontologische Zeitschrift" pageId="64" pageNumber="65" pagination="1 - 47" refId="B2" refString="Arthaber, G, 1922. Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier. Palaeontologische Zeitschrift 4: 1 - 47" title="Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier." volume="4" year="1922">Arthaber 1922</bibRefCitation>
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: p. 18, fig. 7c
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Criorhynchidae" genus="Criorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="simus">Criorhynchus simus</taxonomicName>
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(Owen):
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<bibRefCitation author="Kuhn, O" journalOrPublisher="Verlag Oeben, Krailling bei Muenchen" pageId="67" pageNumber="68" refId="B42" refString="Kuhn, O, 1967. Die fossile Wirbeltierklasse Pterosauria. Verlag Oeben, Krailling bei Muenchen, 52 pp." title="Die fossile Wirbeltierklasse Pterosauria." year="1967">Kuhn 1967</bibRefCitation>
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: 38
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Criorhynchidae" genus="Criorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="simus">Criorhynchus simus</taxonomicName>
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(Owen):
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<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer 1978</bibRefCitation>
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: p. 60
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="clavirostris">Coloborhynchus clavirostris</taxonomicName>
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Owen:
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<bibRefCitation author="Lee, YN" journalOrPublisher="Palaeontology" pageId="67" pageNumber="68" pagination="755 - 763" refId="B43" refString="Lee, YN, 1994. The Early Cretaceous pterodactyloid pterosaur Coloborhynchus from North America. Palaeontology 37: 755 - 763" title="The Early Cretaceous pterodactyloid pterosaur Coloborhynchus from North America." volume="37" year="1994">Lee 1994</bibRefCitation>
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: p. 756, fig. 4
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="clavirostris">Coloborhynchus clavirostris</taxonomicName>
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Owen:
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: p. 206
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="clavirostris">Coloborhynchus clavirostris</taxonomicName>
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Owen:
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<bibRefCitation author="Veldmeijer, AJ" journalOrPublisher="PalArch's Journal of Vertebrate Palaeontology" pageId="70" pageNumber="71" pagination="1 - 14" refId="B92" refString="Veldmeijer, AJ, 2003. Preliminary description of a skull and wing of a Brazilian Cretaceous (Santana Formation; Aptian-Albian) pterosaur (Pterodactyloidea) in the collection of the AMNH. PalArch's Journal of Vertebrate Palaeontology 0: 1 - 14" title="Preliminary description of a skull and wing of a Brazilian Cretaceous (Santana Formation; Aptian-Albian) pterosaur (Pterodactyloidea) in the collection of the AMNH." volume="0" year="2003">Veldmeijer 2003</bibRefCitation>
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: 42
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="clavirostris">Coloborhynchus clavirostris</taxonomicName>
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Owen:
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<bibRefCitation author="Rodrigues, T" journalOrPublisher="Zitteliana B" pageId="68" pageNumber="69" pagination="219 - 228" refId="B73" refString="Rodrigues, T, Kellner, AWA, 2008. Review of the pterodactyloid pterosaur Coloborhynchus. Zitteliana B 28: 219 - 228" title="Review of the pterodactyloid pterosaur Coloborhynchus." volume="28" year="2008">Rodrigues and Kellner 2008</bibRefCitation>
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: p. 220, fig. 1.1, 2.1, 3.1
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="clavirostris">Coloborhynchus clavirostris</taxonomicName>
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Owen:
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<bibRefCitation author="Martill, DM" editor="Batten, DJ" journalOrPublisher="The Palaeontological Association, London" pageId="67" pageNumber="68" pagination="370 - 390" refId="B54" refString="Martill, DM, Sweetman, SC, Witton, MP, 2011. Pterosaurs. In: Batten, DJ, Ed., English Wealden fossils. The Palaeontological Association, London: 370 - 390" title="Pterosaurs." volumeTitle="English Wealden fossils." year="2011">Martill, Sweetman and Witton 2011</bibRefCitation>
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: p. 380, fig. 25.8
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="holotype">
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<paragraph pageId="21" pageNumber="22">Holotype.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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NHMUK PV R 1822, anterior portion of the rostrum (
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coloborhynchus clavirostris. Holotype, NHMUK PV R 1822 (late Berriasian / Valanginian, Hastings Group), anterior part of the rostrum A anterior view B respective line drawing C left lateral view D respective line drawing. Abbreviations: dep - depression, m - maxillae, pm - premaxillae, pmcr - premaxillaery crest, prid - palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. Photos courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22075" pageId="21" pageNumber="22">Fig. 7A-D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="type locality">
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<paragraph pageId="21" pageNumber="22">Type locality.</paragraph>
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<paragraph pageId="21" pageNumber="22">St.-Leonards-on-Sea, East Sussex, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="type horizon">
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<paragraph pageId="21" pageNumber="22">Type horizon.</paragraph>
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<paragraph pageId="21" pageNumber="22">Hastings Group (late Berriasian / Valanginian).</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="diagnosis">
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<paragraph pageId="21" pageNumber="22">Diagnosis.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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Anhanguerid pterosaur with the following combination of characters that distinguishes it from other members of the clade (autapomorphies are marked with an asterisk): oval depression beneath the first pair of alveoli*; second, third and fourth pairs of alveoli located laterally*; fifth and sixth pairs of alveoli located more medially than the preceding alveoli on the base of the palatal ridge*; anterior part of the palatal ridge bordered by two shallow longitudinally elongated depressions* (from
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<bibRefCitation author="Rodrigues, T" journalOrPublisher="Zitteliana B" pageId="68" pageNumber="69" pagination="219 - 228" refId="B73" refString="Rodrigues, T, Kellner, AWA, 2008. Review of the pterodactyloid pterosaur Coloborhynchus. Zitteliana B 28: 219 - 228" title="Review of the pterodactyloid pterosaur Coloborhynchus." volume="28" year="2008">Rodrigues and Kellner 2008</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
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<paragraph pageId="21" pageNumber="22">Description.</paragraph>
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<paragraph lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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The holotype of
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="clavirostris">
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<emphasis italics="true" pageId="22" pageNumber="23">
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<pageBreakToken pageId="22" pageNumber="23" start="start">Coloborhynchus</pageBreakToken>
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clavirostris
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</emphasis>
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</taxonomicName>
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(NHMUK PV R 1822) is a fragment of the premaxillae and maxillae, and has previously been described in detail (
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<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B67" refString="Owen, R, 1874. Monograph on the fossil Reptilia of the Mesozoic Formations. Palaeontographical Society, London, 14 pp." title="Monograph on the fossil Reptilia of the Mesozoic Formations." year="1874">Owen 1874</bibRefCitation>
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;
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<bibRefCitation author="Veldmeijer, AJ" journalOrPublisher="PalArch's Journal of Vertebrate Palaeontology" pageId="70" pageNumber="71" pagination="1 - 14" refId="B92" refString="Veldmeijer, AJ, 2003. Preliminary description of a skull and wing of a Brazilian Cretaceous (Santana Formation; Aptian-Albian) pterosaur (Pterodactyloidea) in the collection of the AMNH. PalArch's Journal of Vertebrate Palaeontology 0: 1 - 14" title="Preliminary description of a skull and wing of a Brazilian Cretaceous (Santana Formation; Aptian-Albian) pterosaur (Pterodactyloidea) in the collection of the AMNH." volume="0" year="2003">Veldmeijer 2003</bibRefCitation>
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;
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<bibRefCitation author="Rodrigues, T" journalOrPublisher="Zitteliana B" pageId="68" pageNumber="69" pagination="219 - 228" refId="B73" refString="Rodrigues, T, Kellner, AWA, 2008. Review of the pterodactyloid pterosaur Coloborhynchus. Zitteliana B 28: 219 - 228" title="Review of the pterodactyloid pterosaur Coloborhynchus." volume="28" year="2008">Rodrigues and Kellner 2008</bibRefCitation>
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). It has a flattened anterior margin, where the first pair of tooth sockets is located. The second, third and fourth pairs of alveoli face laterally, and the fifth and sixth pairs are situated closer to the midline.
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="clavirostris">
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<emphasis italics="true" pageId="22" pageNumber="23">Coloborhynchus clavirostris</emphasis>
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</taxonomicName>
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also has an anterior expansion and a strong palatal ridge in addition a robust premaxillary crest, which begins at the tip of the rostrum.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="remarks">
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<paragraph pageId="22" pageNumber="23">Remarks.</paragraph>
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<paragraph pageId="22" pageNumber="23">
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<bibRefCitation author="Rodrigues, T" journalOrPublisher="Zitteliana B" pageId="68" pageNumber="69" pagination="219 - 228" refId="B73" refString="Rodrigues, T, Kellner, AWA, 2008. Review of the pterodactyloid pterosaur Coloborhynchus. Zitteliana B 28: 219 - 228" title="Review of the pterodactyloid pterosaur Coloborhynchus." volume="28" year="2008">Rodrigues and Kellner (2008)</bibRefCitation>
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reviewed the taxonomic history of the genus
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="22" pageNumber="23">Coloborhynchus</emphasis>
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</taxonomicName>
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and the species
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="clavirostris">
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<emphasis italics="true" pageId="22" pageNumber="23">Coloborhynchus clavirostris</emphasis>
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</taxonomicName>
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, and thus only the main points are repeated here. The genus and species were erected by
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<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B67" refString="Owen, R, 1874. Monograph on the fossil Reptilia of the Mesozoic Formations. Palaeontographical Society, London, 14 pp." title="Monograph on the fossil Reptilia of the Mesozoic Formations." year="1874">Owen (1874)</bibRefCitation>
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. Later authors such as
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley (1914)</bibRefCitation>
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,
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<bibRefCitation author="Kuhn, O" journalOrPublisher="Verlag Oeben, Krailling bei Muenchen" pageId="67" pageNumber="68" refId="B42" refString="Kuhn, O, 1967. Die fossile Wirbeltierklasse Pterosauria. Verlag Oeben, Krailling bei Muenchen, 52 pp." title="Die fossile Wirbeltierklasse Pterosauria." year="1967">Kuhn (1967)</bibRefCitation>
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and
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<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer (1978)</bibRefCitation>
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regarded it synonymous with
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<taxonomicName baseAuthorityName="Owen" baseAuthorityYear="1861" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="simus">
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<emphasis italics="true" pageId="22" pageNumber="23">Ornithocheirus simus</emphasis>
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</taxonomicName>
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[=
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Criorhynchidae" genus="Criorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="simus">
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<emphasis italics="true" pageId="22" pageNumber="23">Criorhynchus simus</emphasis>
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</taxonomicName>
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].
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<bibRefCitation author="Lee, YN" journalOrPublisher="Palaeontology" pageId="67" pageNumber="68" pagination="755 - 763" refId="B43" refString="Lee, YN, 1994. The Early Cretaceous pterodactyloid pterosaur Coloborhynchus from North America. Palaeontology 37: 755 - 763" title="The Early Cretaceous pterodactyloid pterosaur Coloborhynchus from North America." volume="37" year="1994">Lee (1994)</bibRefCitation>
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revalidated
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="22" pageNumber="23">Coloborhynchus</emphasis>
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</taxonomicName>
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, and
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
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not only considered it a valid genus, but also referred additional species to the genus.
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<bibRefCitation author="Rodrigues, T" journalOrPublisher="Zitteliana B" pageId="68" pageNumber="69" pagination="219 - 228" refId="B73" refString="Rodrigues, T, Kellner, AWA, 2008. Review of the pterodactyloid pterosaur Coloborhynchus. Zitteliana B 28: 219 - 228" title="Review of the pterodactyloid pterosaur Coloborhynchus." volume="28" year="2008">Rodrigues and Kellner (2008)</bibRefCitation>
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, noting several unique characters of
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="clavirostris">
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<emphasis italics="true" pageId="22" pageNumber="23">Coloborhynchus clavirostris</emphasis>
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</taxonomicName>
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, restricted the genus to the type species. This view is followed here.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/22075" pageId="22" pageNumber="23" start="Figure 7" startId="F7">
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<paragraph pageId="22" pageNumber="23">
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<emphasis bold="true" pageId="22" pageNumber="23">Figure 7.</emphasis>
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<taxonomicName authorityName="Owen" authorityYear="1874" class="Reptilia" family="Ornithocheiridae" genus="Coloborhynchus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="clavirostris">
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<emphasis italics="true" pageId="22" pageNumber="23">Coloborhynchus clavirostris</emphasis>
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</taxonomicName>
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. Holotype, NHMUK PV R 1822 (late Berriasian / Valanginian, Hastings Group), anterior part of the rostrum
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<emphasis bold="true" pageId="22" pageNumber="23">A</emphasis>
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anterior view
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<emphasis bold="true" pageId="22" pageNumber="23">B</emphasis>
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respective line drawing
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<emphasis bold="true" pageId="22" pageNumber="23">C</emphasis>
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left lateral view
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<emphasis bold="true" pageId="22" pageNumber="23">D</emphasis>
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respective line drawing. Abbreviations:
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<emphasis bold="true" pageId="22" pageNumber="23">dep</emphasis>
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- depression,
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<emphasis bold="true" pageId="22" pageNumber="23">m</emphasis>
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- maxillae,
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<emphasis bold="true" pageId="22" pageNumber="23">pm</emphasis>
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- premaxillae,
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<emphasis bold="true" pageId="22" pageNumber="23">pmcr</emphasis>
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- premaxillaery crest,
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<emphasis bold="true" pageId="22" pageNumber="23">prid</emphasis>
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- palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |