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<document id="7895910560F0FD2F9A571AAAE972DCA6" ID="10.11646/zootaxa.4277.4.4SLASH11421" ID-CLB-Dataset="32818" ID-DOI="10.11646/zootaxa.4277.4.4" ID-GBIF-Dataset="fcd8b13d-0c2c-40ea-9053-8b6c1b3b5a47" ID-ISSN="1175-5326" ID-Zenodo-Dep="814621" ID-ZooBank="A936C241-1B69-411F-873C-39E19341A04A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497946662122" checkinUser="plazi" docAuthor="Negrobov, Oleg P., Grichanov, Igor Ya. &amp; Selivanova, Olga V." docDate="2017" docId="DA22878205048C09FF576B4AFA5AFF58" docLanguage="en" docName="zootaxa.4277.4.4SLASH11421.pdf" docOrigin="Zootaxa 4277 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sympycnus Negrobov &amp; Grichanov &amp; Selivanova 2017" docType="key" docVersion="7" lastPageNumber="547" masterDocId="261BFFFA050B8C19FFC06822FFC6FFB0" masterDocTitle="Review of East Palaearctic species of Sympycnus Loew, 1857, with a key to species" masterLastPageNumber="548" masterPageNumber="531" pageNumber="546" updateTime="1698447347709" updateUser="ExternalLinkService">
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<mods:title id="8DFB184A01A9A68910740A500E9B4CA4">Review of East Palaearctic species of Sympycnus Loew, 1857, with a key to species</mods:title>
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<treatment id="DA22878205048C09FF576B4AFA5AFF58" ID-DOI="http://doi.org/10.5281/zenodo.6040420" ID-GBIF-Taxon="131977672" ID-Zenodo-Dep="6040420" LSID="urn:lsid:plazi:treatment:DA22878205048C09FF576B4AFA5AFF58" httpUri="http://treatment.plazi.org/id/DA22878205048C09FF576B4AFA5AFF58" lastPageId="16" lastPageNumber="547" pageId="15" pageNumber="546">
<subSubSection id="1A91651F05048C16FF576B4AFC62FC32" box="[151,932,871,898]" pageId="15" pageNumber="546" type="nomenclature">
<paragraph id="5234369405048C16FF576B4AFC62FC32" blockId="15.[151,932,871,898]" box="[151,932,871,898]" pageId="15" pageNumber="546">
<heading id="097C81F805048C16FF576B4AFC62FC32" bold="true" box="[151,932,871,898]" fontSize="11" level="1" pageId="15" pageNumber="546" reason="1">
<emphasis id="60FFEA8605048C16FF576B4AFC62FC32" bold="true" box="[151,932,871,898]" pageId="15" pageNumber="546">
Key to the East Palaearctic species of
<taxonomicName id="958B4D1705048C16FDAD6B45FC87FC32" ID-CoL="7QYD" ID-ENA="377530" authority="Loew" authorityName="Negrobov &amp; Grichanov &amp; Selivanova" authorityYear="2017" box="[621,833,871,898]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="genus">
<emphasis id="60FFEA8605048C16FDAD6B45FD30FC31" bold="true" box="[621,758,871,897]" italics="true" pageId="15" pageNumber="546">Sympycnus</emphasis>
Loew
</taxonomicName>
(males)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1A91651F05048C09FF576B8CFA5AFF58" lastPageId="16" lastPageNumber="547" pageId="15" pageNumber="546" type="key">
<keyStep id="E97F2E3105048C16FF576B8CFA5DFC51" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576B8CFA5AFC73" blockId="15.[151,1436,942,2020]" box="[151,1436,942,963]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576B8CFA5AFC73" box="[151,1436,942,963]" pageId="15" pageNumber="546">
1 Hind tarsus without long spine, long bristles or erect setae (
<figureCitation id="CAB02A1105048C16FCDA6B8CFCA7FC73" box="[794,865,942,963]" captionStart="FIGURES 20 25" captionStartId="9.[151,264,1979,2002]" captionTargetBox="[263,1328,987,1960]" captionTargetId="figure@9.[248,1349,965,1969]" captionTargetPageId="9" captionText="FIGURES 20 25. Sympycnus simplicipes Becker, male (Tajikistan). 20, antenna, lateral view; 21, fore leg, lateral view; 22, fore basitarsus, lateral view; 23, hind tarsus, lateral view; 24, hypopygium, left lateral view; 25, hypopygium, ventral view." httpUri="https://zenodo.org/record/814629/files/figure.png" pageId="15" pageNumber="546">Fig. 23</figureCitation>
), rarely with elongate hairs dorsally................. 2
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576BEEFA5DFC51" blockId="15.[151,1436,942,2020]" box="[151,1435,972,993]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576BEEFA5DFC51" box="[151,1435,972,993]" pageId="15" pageNumber="546">
- Hind tarsus with long bristles (
<figureCitation id="CAB02A1105048C16FDC86BEEFD94FC51" box="[520,594,972,993]" captionStart="FIGURES 13 19" captionStartId="6.[153,266,1295,1318]" captionTargetBox="[193,1422,218,1248]" captionTargetId="figure@6.[151,1436,193,1275]" captionTargetPageId="6" captionText="FIGURES 13 19. Sympycnus leleji sp. nov., male holotype. 13, antenna, lateral view; 14, fore leg, lateral view; 15, fore tarsus, dorsal view; 16, hind tarsus, lateroventral view; 17, hypopygium, left lateral view; 18, hypopygium, ventral view; 19, apicoventral epandrial lobe and basoventral appendage of postgonite, dorsal view." httpUri="https://zenodo.org/record/814627/files/figure.png" pageId="15" pageNumber="546">Figs 16</figureCitation>
,
<figureCitation id="CAB02A1105048C16FD9C6BEEFDB3FC51" box="[604,629,972,993]" captionStart="FIGURES 39 44" captionStartId="13.[151,264,1984,2007]" captionTargetBox="[220,1371,1092,1969]" captionTargetId="figure@13.[220,1371,1092,1969]" captionTargetPageId="13" captionText="FIGURES 39 44. Sympycnus yakutensis sp. nov., male holotype. 39, head, anterior view; 40, antenna, lateral view; 41, wing; 42, hind tarsus, dorsolateral view; 43, hypopygium, left lateral view; 44, hypopygium, ventral view." httpUri="https://zenodo.org/record/814635/files/figure.png" pageId="15" pageNumber="546">42</figureCitation>
) or erect setae (
<figureCitation id="CAB02A1105048C16FCCD6BEEFC8DFC51" box="[781,843,972,993]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[151,1435,844,1881]" captionTargetId="figure@2.[151,1436,844,1892]" captionTargetPageId="2" captionText="FIGURES 1 6. Sympycnus changaicus Negrobov, male (Mongolia). 1, head, dorsolateral view; 2, fore tarsus, lateral view; 3, hind tarsus, lateral view, with inset showing erect setae on tarsomeres 3 and 4; 4, wing; 5, hypopygium, left lateral view (holotype); 6, hypopygium, ventral view (holotype)." httpUri="https://zenodo.org/record/814623/files/figure.png" pageId="15" pageNumber="546">Figs 3</figureCitation>
,
<figureCitation id="CAB02A1105048C16FC956BEEFCA8FC51" box="[853,878,972,993]" captionStart="FIGURES 7 12" captionStartId="4.[151,264,1938,1961]" captionTargetBox="[188,1393,999,1904]" captionTargetId="figure@4.[188,1398,993,1915]" captionTargetPageId="4" captionText="FIGURES 7 12. Sympycnus ferganicus sp. nov., male holotype. 7, habitus; 8, head, dorsolateral view; 9, wing; 10, hind tarsus, dorsolateral view; 11, hypopygium, left lateral view; 12, hypopygium, ventral view. Abbreviations: apv lobe = apicoventral epandrial lobe, cer = cercus, ep = epandrium, hyp = hypandrium, ph = phallus, pgt = postgonite, sur = surstylus." httpUri="https://zenodo.org/record/814625/files/figure.png" pageId="15" pageNumber="546">10</figureCitation>
) on some tarsomeres............................ 6
</keyLead>
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<keyStep id="E97F2E3105048C16FF576BCBFA5DFBAB" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576BCBFA5DFC4E" blockId="15.[151,1436,942,2020]" box="[151,1435,1001,1022]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576BCBFA5DFC4E" box="[151,1435,1001,1022]" pageId="15" pageNumber="546">2 At least fore femur entirely and hind femur partly black....................................................... 3</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576C24FA5DFBAB" blockId="15.[151,1436,942,2020]" box="[151,1435,1030,1051]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576C24FA5DFBAB" box="[151,1435,1030,1051]" pageId="15" pageNumber="546">- Fore and hind femora yellow, sometimes brown or black at apex or at base........................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576C06FA5AFB21" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576C06FA5DFBE6" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576C06FA5DFBE6" pageId="15" pageNumber="546">
3 Fore tarsus without claws; hind tarsomere 3 without erect setae......................................................................................................
<taxonomicName id="958B4D1705048C16FC9D6C62FA5DFBE6" ID-CoL="9HTHX" authority="Negrobov, Barkalov, Selivanova &amp; Grichanov" authorityName="Negrobov et al." authorityYear="2016" box="[861,1435,1088,1110]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="taimyricus">
<emphasis id="60FFEA8605048C16FC9D6C62FC1CFBE6" box="[861,986,1088,1110]" italics="true" pageId="15" pageNumber="546">S. taimyricus</emphasis>
Negrobov, Barkalov, Selivanova &amp; Grichanov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576C7CFA5AFB21" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576C7CFA5AFB21" pageId="15" pageNumber="546">
- Fore tarsus with 1 claw (
<figureCitation id="CAB02A1105048C16FE0D6C7CFDD3FBC3" box="[461,533,1118,1139]" captionStart="FIGURES 32 38" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[208,1366,1229,1928]" captionTargetId="figure@11.[196,1382,1226,1933]" captionTargetPageId="11" captionText="FIGURES 32 38. Sympycnus stackelbergi sp. nov., male holotype. 32, antenna, lateral view; 33, fore leg, lateral view; 34, hind tarsus, lateroventral view; 35, hypopygium, left lateral view; 36, hypopygium, ventral view; 37, cerci, dorsal view; 38, apicoventral epandrial lobe, dorsal view." httpUri="https://zenodo.org/record/814633/files/figure.png" pageId="15" pageNumber="546">Fig. 33</figureCitation>
); hind tarsomere 3 (
<figureCitation id="CAB02A1105048C16FD156C7CFCDDFBC3" box="[725,795,1118,1139]" captionStart="FIGURES 32 38" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[208,1366,1229,1928]" captionTargetId="figure@11.[196,1382,1226,1933]" captionTargetPageId="11" captionText="FIGURES 32 38. Sympycnus stackelbergi sp. nov., male holotype. 32, antenna, lateral view; 33, fore leg, lateral view; 34, hind tarsus, lateroventral view; 35, hypopygium, left lateral view; 36, hypopygium, ventral view; 37, cerci, dorsal view; 38, apicoventral epandrial lobe, dorsal view." httpUri="https://zenodo.org/record/814633/files/figure.png" pageId="15" pageNumber="546">Fig. 34</figureCitation>
) with short erect setae, sometimes hardly visible...............................................................................
<taxonomicName id="958B4D1705048C16FC316C59FBBBFB21" ID-CoL="9J25Y" authority="Negrobov &amp; Grichanov &amp; Selivanova, 2017" authorityName="Negrobov &amp; Grichanov &amp; Selivanova" authorityYear="2017" box="[1009,1149,1147,1169]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="stackelbergi" status="sp. nov.">
<emphasis id="60FFEA8605048C16FC316C59FBBBFB21" box="[1009,1149,1147,1169]" italics="true" pageId="15" pageNumber="546">S. stackelbergi</emphasis>
</taxonomicName>
<emphasis id="60FFEA8605048C16FB436C59FB09FB21" bold="true" box="[1155,1231,1147,1169]" pageId="15" pageNumber="546">
<taxonomicNameLabel id="7BCC57FD05048C16FB436C59FB09FB21" box="[1155,1231,1147,1169]" pageId="15" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
[see also couplet 13]
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<keyStep id="E97F2E3105048C16FF576CBBFA5DFB59" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576CBBFA5DFB7B" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576CBBFA5DFB7B" pageId="15" pageNumber="546">
4 Fore and mid tarsomeres 35 with long hairs dorsally, longer on tarsomere 5; mid basitarsus with dorsal row of long bristles.........................................................................
<taxonomicName id="958B4D1705048C16FB856C97FA5DFB7B" authority="Tang, Wang &amp; Yang" authorityName="Tang et al." authorityYear="2015" box="[1093,1435,1205,1227]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="longipilosus">
<emphasis id="60FFEA8605048C16FB856C97FB14FB7B" box="[1093,1234,1205,1227]" italics="true" pageId="15" pageNumber="546">S. longipilosus</emphasis>
Tang, Wang &amp; Yang
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576CF6FA5DFB59" blockId="15.[151,1436,942,2020]" box="[151,1435,1236,1257]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576CF6FA5DFB59" box="[151,1435,1236,1257]" pageId="15" pageNumber="546">
- Fore and mid tarsi without remarkably long hairs and bristles (
<figureCitation id="CAB02A1105048C16FCF86CF6FCB0FB59" box="[824,886,1236,1257]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1915,1938]" captionTargetBox="[151,1435,844,1881]" captionTargetId="figure@2.[151,1436,844,1892]" captionTargetPageId="2" captionText="FIGURES 1 6. Sympycnus changaicus Negrobov, male (Mongolia). 1, head, dorsolateral view; 2, fore tarsus, lateral view; 3, hind tarsus, lateral view, with inset showing erect setae on tarsomeres 3 and 4; 4, wing; 5, hypopygium, left lateral view (holotype); 6, hypopygium, ventral view (holotype)." httpUri="https://zenodo.org/record/814623/files/figure.png" pageId="15" pageNumber="546">Figs 2</figureCitation>
,
<figureCitation id="CAB02A1105048C16FC406CF6FC5CFB59" box="[896,922,1236,1257]" captionStart="FIGURES 13 19" captionStartId="6.[153,266,1295,1318]" captionTargetBox="[193,1422,218,1248]" captionTargetId="figure@6.[151,1436,193,1275]" captionTargetPageId="6" captionText="FIGURES 13 19. Sympycnus leleji sp. nov., male holotype. 13, antenna, lateral view; 14, fore leg, lateral view; 15, fore tarsus, dorsal view; 16, hind tarsus, lateroventral view; 17, hypopygium, left lateral view; 18, hypopygium, ventral view; 19, apicoventral epandrial lobe and basoventral appendage of postgonite, dorsal view." httpUri="https://zenodo.org/record/814627/files/figure.png" pageId="15" pageNumber="546">14</figureCitation>
,
<figureCitation id="CAB02A1105048C16FC646CF6FC78FB59" box="[932,958,1236,1257]" captionStart="FIGURES 26 31" captionStartId="11.[151,264,1123,1146]" captionTargetBox="[226,1362,206,1093]" captionTargetId="figure@11.[215,1383,193,1112]" captionTargetPageId="11" captionText="FIGURES 26 31. Sympycnus simplicitarsis Becker, male syntype. 26, antenna, lateral view; 27, fore tarsus, lateral view; 28, hypopygium, left lateral view; 29, hypopygium, ventral view; 30, basoventral appendage of postgonite, lateral view; 31, apex of phallus, lateral view." httpUri="https://zenodo.org/record/814631/files/figure.png" pageId="15" pageNumber="546">27</figureCitation>
,
<figureCitation id="CAB02A1105048C16FC086CF6FC27FB59" box="[968,993,1236,1257]" captionStart="FIGURES 32 38" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[208,1366,1229,1928]" captionTargetId="figure@11.[196,1382,1226,1933]" captionTargetPageId="11" captionText="FIGURES 32 38. Sympycnus stackelbergi sp. nov., male holotype. 32, antenna, lateral view; 33, fore leg, lateral view; 34, hind tarsus, lateroventral view; 35, hypopygium, left lateral view; 36, hypopygium, ventral view; 37, cerci, dorsal view; 38, apicoventral epandrial lobe, dorsal view." httpUri="https://zenodo.org/record/814633/files/figure.png" pageId="15" pageNumber="546">33</figureCitation>
)................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576CD3FA5DFAF1" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576CD3FA5DFA93" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576CD3FA5DFA93" pageId="15" pageNumber="546">
5 Hind basitarsus shorter than tarsomere 4 (
<figureCitation id="CAB02A1105048C16FDB16CD3FD7EFAB6" box="[625,696,1265,1286]" captionStart="FIGURES 20 25" captionStartId="9.[151,264,1979,2002]" captionTargetBox="[263,1328,987,1960]" captionTargetId="figure@9.[248,1349,965,1969]" captionTargetPageId="9" captionText="FIGURES 20 25. Sympycnus simplicipes Becker, male (Tajikistan). 20, antenna, lateral view; 21, fore leg, lateral view; 22, fore basitarsus, lateral view; 23, hind tarsus, lateral view; 24, hypopygium, left lateral view; 25, hypopygium, ventral view." httpUri="https://zenodo.org/record/814629/files/figure.png" pageId="15" pageNumber="546">Fig. 23</figureCitation>
); fore basitarsus with elongate setae at base and at apex (
<figureCitation id="CAB02A1105048C16FB1B6CD3FAE6FAB6" box="[1243,1312,1265,1286]" captionStart="FIGURES 20 25" captionStartId="9.[151,264,1979,2002]" captionTargetBox="[263,1328,987,1960]" captionTargetId="figure@9.[248,1349,965,1969]" captionTargetPageId="9" captionText="FIGURES 20 25. Sympycnus simplicipes Becker, male (Tajikistan). 20, antenna, lateral view; 21, fore leg, lateral view; 22, fore basitarsus, lateral view; 23, hind tarsus, lateral view; 24, hypopygium, left lateral view; 25, hypopygium, ventral view." httpUri="https://zenodo.org/record/814629/files/figure.png" pageId="15" pageNumber="546">Fig. 22</figureCitation>
).............................................................................................
<taxonomicName id="958B4D1705048C16FB0A6D2FFA5DFA93" ID-CoL="53SNJ" authority="Becker" authorityName="Becker" authorityYear="1908" box="[1226,1435,1293,1315]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="60FFEA8605048C16FB0A6D2FFA94FA93" box="[1226,1362,1293,1315]" italics="true" pageId="15" pageNumber="546">S. simplicipes</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576D0EFA5DFAF1" blockId="15.[151,1436,942,2020]" box="[151,1435,1323,1345]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576D0EFA5DFAF1" box="[151,1435,1323,1345]" pageId="15" pageNumber="546">
- Hind basitarsus longer than tarsomere 4; fore basitarsus with short setulae (
<figureCitation id="CAB02A1105048C16FC726D0EFC3FFAF1" box="[946,1017,1324,1345]" captionStart="FIGURES 26 31" captionStartId="11.[151,264,1123,1146]" captionTargetBox="[226,1362,206,1093]" captionTargetId="figure@11.[215,1383,193,1112]" captionTargetPageId="11" captionText="FIGURES 26 31. Sympycnus simplicitarsis Becker, male syntype. 26, antenna, lateral view; 27, fore tarsus, lateral view; 28, hypopygium, left lateral view; 29, hypopygium, ventral view; 30, basoventral appendage of postgonite, lateral view; 31, apex of phallus, lateral view." httpUri="https://zenodo.org/record/814631/files/figure.png" pageId="15" pageNumber="546">Fig. 27</figureCitation>
)...............
<taxonomicName id="958B4D1705048C16FB796D09FA5DFAF1" ID-CoL="53SNK" authority="Becker" authorityName="Becker" authorityYear="1900" box="[1209,1435,1323,1345]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="simplicitarsis">
<emphasis id="60FFEA8605048C16FB796D09FA94FAF1" box="[1209,1362,1323,1345]" italics="true" pageId="15" pageNumber="546">S. simplicitarsis</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576D6BFA5DFACB" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576D6BFA5DFAEE" blockId="15.[151,1436,942,2020]" box="[151,1435,1353,1374]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576D6BFA5DFAEE" box="[151,1435,1353,1374]" pageId="15" pageNumber="546">
6 Hind tarsomere 3 with two long thin bristles at base, without long spine at apex (
<figureCitation id="CAB02A1105048C16FC276D6BFBEAFAEE" box="[999,1068,1353,1374]" captionStart="FIGURES 39 44" captionStartId="13.[151,264,1984,2007]" captionTargetBox="[220,1371,1092,1969]" captionTargetId="figure@13.[220,1371,1092,1969]" captionTargetPageId="13" captionText="FIGURES 39 44. Sympycnus yakutensis sp. nov., male holotype. 39, head, anterior view; 40, antenna, lateral view; 41, wing; 42, hind tarsus, dorsolateral view; 43, hypopygium, left lateral view; 44, hypopygium, ventral view." httpUri="https://zenodo.org/record/814635/files/figure.png" pageId="15" pageNumber="546">Fig. 42</figureCitation>
)............................. 7
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576D44FA5DFACB" blockId="15.[151,1436,942,2020]" box="[151,1435,1382,1403]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576D44FA5DFACB" box="[151,1435,1382,1403]" pageId="15" pageNumber="546">- Hind tarsomere 3 without two long thin bristles at base, with or without long bristle at apex........................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576DA6FA5AF9BE" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576DA6FA5AFA63" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576DA6FA5AFA63" pageId="15" pageNumber="546">
7 Hind femur pale yellow, with apical 1/3 to 2/3 brown to dark brown; mid tibia usually with posteroventral bristle; hind tarsom- ere 3 distinctly shorter than tarsomere 2; apicoventral lobe of hypandrium subtriangular and pointed at apex (see
<bibRefCitation id="361A4B6505048C16FAEC6D83FED5FA63" author="Pollet" pageId="15" pageNumber="546" refString="Pollet, M., Persson, M., Boggild, E. &amp; Crossley, R. (2015) A long-lasting taxonomic problem in European Sympycnus resolved, with the description of a new species and data on habitat preferences. Zootaxa, 4032 (1), 081 - 102. https: // doi. org / 10.11646 / zootaxa. 4032.1.4" type="journal article" year="2015">
Pollet
<emphasis id="60FFEA8605048C16FAAB6D83FA50FA06" box="[1387,1430,1440,1462]" italics="true" pageId="15" pageNumber="546">et al</emphasis>
. 2015
</bibRefCitation>
, fig. 8)..........................................................................
<taxonomicName id="958B4D1705048C16FB106D9FFA5AFA63" ID-CoL="8BV65" authority="Fallen" authorityName="Fallen" authorityYear="1823" box="[1232,1436,1469,1491]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="pulicarius">
<emphasis id="60FFEA8605048C16FB106D9FFA8CFA63" box="[1232,1354,1469,1491]" italics="true" pageId="15" pageNumber="546">S. pulicarius</emphasis>
(Fallén)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576DFEFA5AF9BE" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576DFEFA5AF9BE" pageId="15" pageNumber="546">
- Hind femur entirely pale yellow; mid tibia without posteroventral bristle; hind tarsomere 3 nearly as long as tarsomere 2 (
<figureCitation id="CAB02A1105048C16FAB46DFEFF3AF9BE" captionStart="FIGURES 39 44" captionStartId="13.[151,264,1984,2007]" captionTargetBox="[220,1371,1092,1969]" captionTargetId="figure@13.[220,1371,1092,1969]" captionTargetPageId="13" captionText="FIGURES 39 44. Sympycnus yakutensis sp. nov., male holotype. 39, head, anterior view; 40, antenna, lateral view; 41, wing; 42, hind tarsus, dorsolateral view; 43, hypopygium, left lateral view; 44, hypopygium, ventral view." httpUri="https://zenodo.org/record/814635/files/figure.png" pageId="15" pageNumber="546">Fig. 42</figureCitation>
); apicoventral lobe of hypandrium semi-rounded (
<figureCitation id="CAB02A1105048C16FD1E6DDBFCE5F9BE" box="[734,803,1529,1550]" captionStart="FIGURES 39 44" captionStartId="13.[151,264,1984,2007]" captionTargetBox="[220,1371,1092,1969]" captionTargetId="figure@13.[220,1371,1092,1969]" captionTargetPageId="13" captionText="FIGURES 39 44. Sympycnus yakutensis sp. nov., male holotype. 39, head, anterior view; 40, antenna, lateral view; 41, wing; 42, hind tarsus, dorsolateral view; 43, hypopygium, left lateral view; 44, hypopygium, ventral view." httpUri="https://zenodo.org/record/814635/files/figure.png" pageId="15" pageNumber="546">Fig. 43</figureCitation>
)...................................
<taxonomicName id="958B4D1705048C16FB096DDAFA8FF9BE" ID-CoL="9J25W" authority="Negrobov &amp; Grichanov &amp; Selivanova, 2017" authorityName="Negrobov &amp; Grichanov &amp; Selivanova" authorityYear="2017" box="[1225,1353,1528,1550]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="yakutensis" status="sp. nov.">
<emphasis id="60FFEA8605048C16FB096DDAFA8FF9BE" box="[1225,1353,1528,1550]" italics="true" pageId="15" pageNumber="546">S. yakutensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCC57FD05048C16FA8F6DDAFA5AF9BE" box="[1359,1436,1528,1550]" pageId="15" pageNumber="546" rank="species">
<emphasis id="60FFEA8605048C16FA8F6DDAFA5AF9BE" bold="true" box="[1359,1436,1528,1550]" pageId="15" pageNumber="546">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576E34FA5DF9F9" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576E34FA5DF99B" blockId="15.[151,1436,942,2020]" box="[151,1435,1558,1579]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576E34FA5DF99B" box="[151,1435,1558,1579]" pageId="15" pageNumber="546">
8 Hind tarsomere 3 with long spine or thick bristle at apex (
<figureCitation id="CAB02A1105048C16FCD06E34FC9FF99B" box="[784,857,1558,1579]" captionStart="FIGURES 13 19" captionStartId="6.[153,266,1295,1318]" captionTargetBox="[193,1422,218,1248]" captionTargetId="figure@6.[151,1436,193,1275]" captionTargetPageId="6" captionText="FIGURES 13 19. Sympycnus leleji sp. nov., male holotype. 13, antenna, lateral view; 14, fore leg, lateral view; 15, fore tarsus, dorsal view; 16, hind tarsus, lateroventral view; 17, hypopygium, left lateral view; 18, hypopygium, ventral view; 19, apicoventral epandrial lobe and basoventral appendage of postgonite, dorsal view." httpUri="https://zenodo.org/record/814627/files/figure.png" pageId="15" pageNumber="546">Fig. 16</figureCitation>
)............................................... 9
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576E16FA5DF9F9" blockId="15.[151,1436,942,2020]" box="[151,1435,1588,1609]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576E16FA5DF9F9" box="[151,1435,1588,1609]" pageId="15" pageNumber="546">- Hind tarsomere 3 without long spine or thick bristle......................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576E73FA5DF90E" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576E73FA5AF934" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576E73FA5AF934" pageId="15" pageNumber="546">
9 Mid femur with almost full posteroventral row of strong bristles, about as long as femur diameter; fore tibia without anterodor- sal row of setae (
<figureCitation id="CAB02A1105048C16FE446E4CFE0DF933" box="[388,459,1646,1667]" captionStart="FIGURES 13 19" captionStartId="6.[153,266,1295,1318]" captionTargetBox="[193,1422,218,1248]" captionTargetId="figure@6.[151,1436,193,1275]" captionTargetPageId="6" captionText="FIGURES 13 19. Sympycnus leleji sp. nov., male holotype. 13, antenna, lateral view; 14, fore leg, lateral view; 15, fore tarsus, dorsal view; 16, hind tarsus, lateroventral view; 17, hypopygium, left lateral view; 18, hypopygium, ventral view; 19, apicoventral epandrial lobe and basoventral appendage of postgonite, dorsal view." httpUri="https://zenodo.org/record/814627/files/figure.png" pageId="15" pageNumber="546">Fig. 14</figureCitation>
); fore tarsus without claws (
<figureCitation id="CAB02A1105048C16FD1A6E4CFCD9F933" box="[730,799,1646,1667]" captionStart="FIGURES 13 19" captionStartId="6.[153,266,1295,1318]" captionTargetBox="[193,1422,218,1248]" captionTargetId="figure@6.[151,1436,193,1275]" captionTargetPageId="6" captionText="FIGURES 13 19. Sympycnus leleji sp. nov., male holotype. 13, antenna, lateral view; 14, fore leg, lateral view; 15, fore tarsus, dorsal view; 16, hind tarsus, lateroventral view; 17, hypopygium, left lateral view; 18, hypopygium, ventral view; 19, apicoventral epandrial lobe and basoventral appendage of postgonite, dorsal view." httpUri="https://zenodo.org/record/814627/files/figure.png" pageId="15" pageNumber="546">Fig. 15</figureCitation>
).......................................
<taxonomicName id="958B4D1705048C16FAC06E4FFA8EF933" ID-CoL="9J25X" authority="Negrobov &amp; Grichanov &amp; Selivanova, 2017" authorityName="Negrobov &amp; Grichanov &amp; Selivanova" authorityYear="2017" box="[1280,1352,1645,1667]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="leleji" status="sp. nov.">
<emphasis id="60FFEA8605048C16FAC06E4FFA8EF933" box="[1280,1352,1645,1667]" italics="true" pageId="15" pageNumber="546">S. leleji</emphasis>
</taxonomicName>
<emphasis id="60FFEA8605048C16FA8F6E4CFA5AF934" bold="true" box="[1359,1436,1646,1668]" pageId="15" pageNumber="546">
<taxonomicNameLabel id="7BCC57FD05048C16FA8F6E4CFA5AF934" box="[1359,1436,1646,1668]" pageId="15" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576EAEFA5DF90E" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576EAEFA5DF90E" pageId="15" pageNumber="546">- Mid femur with at most 23 posteroventral setae at apex; fore tibia with or without anterodorsal row of setae; fore tarsus with or without claws..................................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576EE4FA5DF8A6" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576EE4FA5DF949" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576EE4FA5DF949" pageId="15" pageNumber="546">
10 Fore tibia slightly, but distinctly thickened, without anterodorsal row of setae, with rows of long hairs, about as long as tibia diameter..............................................
<taxonomicName id="958B4D1705048C16FCA26EC1FA5DF949" authority="Negrobov, Barkalov, Selivanova &amp; Grichanov" authorityName="Negrobov et al." authorityYear="2016" box="[866,1435,1763,1785]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="gorodkovi">
<emphasis id="60FFEA8605048C16FCA26EC1FC1DF949" box="[866,987,1763,1785]" italics="true" pageId="15" pageNumber="546">S. gorodkovi</emphasis>
Negrobov, Barkalov, Selivanova &amp; Grichanov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576F23FA5DF8A6" blockId="15.[151,1436,942,2020]" box="[151,1435,1793,1814]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576F23FA5DF8A6" box="[151,1435,1793,1814]" pageId="15" pageNumber="546">
- Fore tibia simple, usually with anterodorsal row of setae (
<figureCitation id="CAB02A1105048C16FCC96F23FC92F8A6" box="[777,852,1793,1814]" captionStart="FIGURES 20 25" captionStartId="9.[151,264,1979,2002]" captionTargetBox="[263,1328,987,1960]" captionTargetId="figure@9.[248,1349,965,1969]" captionTargetPageId="9" captionText="FIGURES 20 25. Sympycnus simplicipes Becker, male (Tajikistan). 20, antenna, lateral view; 21, fore leg, lateral view; 22, fore basitarsus, lateral view; 23, hind tarsus, lateral view; 24, hypopygium, left lateral view; 25, hypopygium, ventral view." httpUri="https://zenodo.org/record/814629/files/figure.png" pageId="15" pageNumber="546">Figs 21</figureCitation>
,
<figureCitation id="CAB02A1105048C16FC9E6F23FCBEF8A6" box="[862,888,1793,1814]" captionStart="FIGURES 32 38" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[208,1366,1229,1928]" captionTargetId="figure@11.[196,1382,1226,1933]" captionTargetPageId="11" captionText="FIGURES 32 38. Sympycnus stackelbergi sp. nov., male holotype. 32, antenna, lateral view; 33, fore leg, lateral view; 34, hind tarsus, lateroventral view; 35, hypopygium, left lateral view; 36, hypopygium, ventral view; 37, cerci, dorsal view; 38, apicoventral epandrial lobe, dorsal view." httpUri="https://zenodo.org/record/814633/files/figure.png" pageId="15" pageNumber="546">33</figureCitation>
), with short hairs............................. 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576F3CFA5DF83B" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576F3CFA5DF8E1" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576F3CFA5DF8E1" pageId="15" pageNumber="546">
11 Hind tarsomere 3 subequal in length to tarsomere 4, with only 1 apical spine......................................................................................................
<taxonomicName id="958B4D1705048C16FC086F19FA5DF8E1" ID-CoL="9HZFN" authority="Negrobov, Barkalov &amp; Selivanova" authorityName="Negrobov et al." authorityYear="2013" box="[968,1435,1851,1873]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="olejnicheki">
<emphasis id="60FFEA8605048C16FC086F19FB8CF8E1" box="[968,1098,1851,1873]" italics="true" pageId="15" pageNumber="546">S. olejnicheki</emphasis>
Negrobov, Barkalov &amp; Selivanova
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576F7BFA5DF83B" blockId="15.[151,1436,942,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576F7BFA5DF83B" pageId="15" pageNumber="546">- Hind tarsomere 3 with 1 short bristle at middle in addition to long apical bristle; tarsomere 4 1.51.8 times longer than tarsom- ere 3............................................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C16FF576FB6FA5DF876" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576FB6FA5DF819" blockId="15.[151,1436,942,2020]" box="[151,1435,1939,1961]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576FB6FA5DF819" box="[151,1435,1939,1961]" pageId="15" pageNumber="546">
12 Apical bristle on hind tarsomere 3 about as long as tarsomere 3; fore and hind femora partly black....
<taxonomicName id="958B4D1705048C16FB0A6FB1FA5DF819" ID-CoL="53SNB" authority="Negrobov" authorityName="Negrobov" authorityYear="1973" box="[1226,1435,1939,1961]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="seticosta">
<emphasis id="60FFEA8605048C16FB0A6FB1FAF0F819" box="[1226,1334,1939,1961]" italics="true" pageId="15" pageNumber="546">S. seticosta</emphasis>
Negrobov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5234369405048C16FF576F93FA5DF876" blockId="15.[151,1436,942,2020]" box="[151,1435,1968,1990]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576F93FA5DF876" box="[151,1435,1968,1990]" pageId="15" pageNumber="546">
- Apical bristle on hind tarsomere 3 about 2 times as long as tarsomere 3; all femora yellow.............
<taxonomicName id="958B4D1705048C16FB1C6F92FA5DF876" ID-CoL="53SPK" authority="Negrobov" authorityName="Negrobov" authorityYear="1973" box="[1244,1435,1968,1990]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="vadimi">
<emphasis id="60FFEA8605048C16FB1C6F92FAF3F876" box="[1244,1333,1968,1990]" italics="true" pageId="15" pageNumber="546">S. vadimi</emphasis>
Negrobov
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E3105048C09FF576FECFA5DFF1C" lastPageId="16" lastPageNumber="547" pageId="15" pageNumber="546">
<paragraph id="5234369405048C16FF576FECFA5AF854" blockId="15.[151,1436,942,2020]" box="[151,1436,1997,2020]" pageId="15" pageNumber="546">
<keyLead id="E97A95A105048C16FF576FECFA5AF854" box="[151,1436,1997,2020]" pageId="15" pageNumber="546">
13 Only hind tarsomere 3 with erect setae (
<figureCitation id="CAB02A1105048C16FDA06FECFD6EF853" box="[608,680,1998,2019]" captionStart="FIGURES 32 38" captionStartId="11.[151,264,1943,1966]" captionTargetBox="[208,1366,1229,1928]" captionTargetId="figure@11.[196,1382,1226,1933]" captionTargetPageId="11" captionText="FIGURES 32 38. Sympycnus stackelbergi sp. nov., male holotype. 32, antenna, lateral view; 33, fore leg, lateral view; 34, hind tarsus, lateroventral view; 35, hypopygium, left lateral view; 36, hypopygium, ventral view; 37, cerci, dorsal view; 38, apicoventral epandrial lobe, dorsal view." httpUri="https://zenodo.org/record/814633/files/figure.png" pageId="15" pageNumber="546">Fig. 34</figureCitation>
); other segments without modified setae...........
<taxonomicName id="958B4D1705048C16FB7D6FEFFA8FF853" ID-CoL="9J25Y" authority="Negrobov &amp; Grichanov &amp; Selivanova, 2017" authorityName="Negrobov &amp; Grichanov &amp; Selivanova" authorityYear="2017" box="[1213,1353,1997,2019]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="546" phylum="Arthropoda" rank="species" species="stackelbergi" status="sp. nov.">
<emphasis id="60FFEA8605048C16FB7D6FEFFA8FF853" box="[1213,1353,1997,2019]" italics="true" pageId="15" pageNumber="546">S. stackelbergi</emphasis>
</taxonomicName>
<emphasis id="60FFEA8605048C16FA8F6FECFA5AF854" bold="true" box="[1359,1436,1998,2020]" pageId="15" pageNumber="546">
<taxonomicNameLabel id="7BCC57FD05048C16FA8F6FECFA5AF854" box="[1359,1436,1998,2020]" pageId="15" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="52343694051B8C09FF5768B5FA5DFF1C" blockId="16.[151,1436,151,232]" box="[151,1435,151,172]" pageId="16" pageNumber="547">
<keyLead id="E97A95A1051B8C09FF5768B5FA5DFF1C" box="[151,1435,151,172]" pageId="16" pageNumber="547">- At least hind tarsomeres 3 and 4 with long bristles or erect setae............................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="E97F2E31051B8C09FF576897FA5AFF58" pageId="16" pageNumber="547">
<paragraph id="52343694051B8C09FF576897FA5DFF7A" blockId="16.[151,1436,151,232]" box="[151,1435,180,202]" pageId="16" pageNumber="547">
<keyLead id="E97A95A1051B8C09FF576897FA5DFF7A" box="[151,1435,180,202]" pageId="16" pageNumber="547">
14 Hind tarsomere 3 enlarged, 1.5 times wider than tarsomere 4 of same tarsus....................
<taxonomicName id="958B4D17051B8C09FB716896FA5DFF7A" ID-CoL="53SFX" authority="Negrobov" authorityName="Negrobov" authorityYear="1973" box="[1201,1435,180,202]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="547" phylum="Arthropoda" rank="species" species="changaicus">
<emphasis id="60FFEA86051B8C09FB716896FAF0FF7A" box="[1201,1334,180,202]" italics="true" pageId="16" pageNumber="547">S. changaicus</emphasis>
Negrobov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="52343694051B8C09FF5768F0FA5AFF58" blockId="16.[151,1436,151,232]" box="[151,1436,209,232]" pageId="16" pageNumber="547">
<keyLead id="E97A95A1051B8C09FF5768F0FA5AFF58" box="[151,1436,209,232]" pageId="16" pageNumber="547">
- Hind tarsomere 3 indistinctly wider than tarsomere 4.........................................
<taxonomicName id="958B4D17051B8C09FB0D68F3FA8FFF57" ID-CoL="9J25V" authority="Negrobov &amp; Grichanov &amp; Selivanova, 2017" authorityName="Negrobov &amp; Grichanov &amp; Selivanova" authorityYear="2017" box="[1229,1353,209,231]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="547" phylum="Arthropoda" rank="species" species="ferganicus" status="sp. nov.">
<emphasis id="60FFEA86051B8C09FB0D68F3FA8FFF57" box="[1229,1353,209,231]" italics="true" pageId="16" pageNumber="547">S. ferganicus</emphasis>
</taxonomicName>
<emphasis id="60FFEA86051B8C09FA8F68F0FA5AFF58" bold="true" box="[1359,1436,210,232]" pageId="16" pageNumber="547">
<taxonomicNameLabel id="7BCC57FD051B8C09FA8F68F0FA5AFF58" box="[1359,1436,210,232]" pageId="16" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>