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<document id="AC0CE01EFB2B029033F965D01FF71BEB" ID-CLB-Dataset="27316" ID-DOI="10.11646/zootaxa.4583.1.1" ID-GBIF-Dataset="027ed27d-5035-4646-93a6-c23b565f3d20" ID-ISSN="1175-5326" ID-Zenodo-Dep="2637603" ID-ZooBank="27F24995-359E-46F6-AB22-75568BACFDCF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555054242242" checkinUser="plazi" docAuthor="Zhang, - H." docDate="2019" docId="724987F6FFB12424FF7EBFCEFF07384D" docLanguage="en" docName="zootaxa.4583.1.1.pdf" docOrigin="Zootaxa 4583 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Nundalia secreta Zhang 2019, n. sp." docType="treatment" docVersion="15" lastPageNumber="58" masterDocId="8E70FF8EFF86241DFFE9BE79FFE73928" masterDocTitle="New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages" masterLastPageNumber="67" masterPageNumber="1" pageNumber="56" updateTime="1698720421338" updateUser="ExternalLinkService">
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<mods:titleInfo id="E51F76162A84D4F2E399B87A61F1ECF2">
<mods:title id="4C2FD7051749B12BE7E2D83188B54B25">New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages</mods:title>
</mods:titleInfo>
<mods:name id="DB966F3E0F067D4ED1F8A28E9139EDC2" type="personal">
<mods:role id="6136B6ED4281121DF240DD914C3D1AD7">
<mods:roleTerm id="0D3788C3485817C09A4AA3F4C063BD05">Author</mods:roleTerm>
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<mods:namePart id="425D7944F70D5D8B7F0EF6A2DE680C00">Zhang, - H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="20F5923E623054BA9CEAEE79CBE99ECC">text</mods:typeOfResource>
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<mods:titleInfo id="A738F9EB49162C4833FAA3806B0E78EE">
<mods:title id="D329DC804C89B9BB3999A18A372C719E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="90F900938903434877678370EA69EA26">
<mods:date id="F3542786B183CFA19C8C3FF0D817703E">2019</mods:date>
<mods:detail id="F65DF9D9AA914DA153A98BC5DA42AA79" type="pubDate">
<mods:number id="BF1AE216D584B1025F17438340445BB7">2019-04-10</mods:number>
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<mods:detail id="F1C279E77BBBB68A083274BD4DC50DDE" type="volume">
<mods:number id="0D18AD38ACF0DB694879468E56F1B8C2">4583</mods:number>
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<mods:number id="D380D1D6045B388198722EB24F838479">1</mods:number>
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<mods:classification id="DE6270A42EF7EB9CF86E3B8A47E0321C">journal article</mods:classification>
<mods:identifier id="5096BFDFC046BA6361BAE56CD4D8A6C3" type="CLB-Dataset">27316</mods:identifier>
<mods:identifier id="484F595BD5E3DE2E23B1CD5A7D1FD92F" type="DOI">10.11646/zootaxa.4583.1.1</mods:identifier>
<mods:identifier id="2315CAF910512CD7B6BD2974B8B490AA" type="GBIF-Dataset">027ed27d-5035-4646-93a6-c23b565f3d20</mods:identifier>
<mods:identifier id="DF67101FC2CEC9741137F088D3171972" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BFD1F330BEC12A2A0E573F5676C26EA3" type="Zenodo-Dep">2637603</mods:identifier>
<mods:identifier id="9A89432DF28E990835F784B701914DA0" type="ZooBank">27F24995-359E-46F6-AB22-75568BACFDCF</mods:identifier>
</mods:mods>
<treatment id="724987F6FFB12424FF7EBFCEFF07384D" ID-GBIF-Taxon="156198403" LSID="urn:lsid:plazi:treatment:724987F6FFB12424FF7EBFCEFF07384D" httpUri="http://treatment.plazi.org/id/724987F6FFB12424FF7EBFCEFF07384D" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56">
<subSubSection id="B2FA656BFFB1242AFF7EBFCEFE4938FA" box="[151,430,439,466]" pageId="55" pageNumber="56" type="nomenclature">
<paragraph id="FA5F36E0FFB1242AFF7EBFCEFE4938FA" blockId="55.[151,430,439,466]" box="[151,430,439,466]" pageId="55" pageNumber="56">
<heading id="A117818CFFB1242AFF7EBFCEFE4938FA" bold="true" box="[151,430,439,466]" fontSize="11" level="1" pageId="55" pageNumber="56" reason="1">
<emphasis id="C894EAF2FFB1242AFF7EBFCEFE4938FA" bold="true" box="[151,430,439,466]" pageId="55" pageNumber="56">
<taxonomicName id="3DE04D63FFB1242AFF7EBFCEFE8238F9" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[151,357,439,465]" class="Aves" family="Threskiornithidae" genus="Nundalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelecaniformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="secreta" status="sp. nov.">
<emphasis id="C894EAF2FFB1242AFF7EBFCEFE8238F9" bold="true" box="[151,357,439,465]" italics="true" pageId="55" pageNumber="56">Nundalia secreta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FFB1242AFE85BFC1FE4938FA" box="[364,430,440,466]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFFB1242AFF7EBF87FAE13A8B" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="FA5F36E0FFB1242AFF7EBF87FB833BE3" blockId="55.[151,1437,510,931]" pageId="55" pageNumber="56">
<emphasis id="C894EAF2FFB1242AFF7EBF87FE643B3F" bold="true" box="[151,387,510,535]" pageId="55" pageNumber="56">Material examined.</emphasis>
<materialsCitation id="4A883CBDFFB1242AFE67BF87FCF33B77" ID-GBIF-Occurrence="2235592654" collectingDate="1988-02-23" collectorName="Nundle SF &amp; W. F. Ponder" elevation="500" latitude="-31.448055" location="South Head Creek" longLatPrecision="20" longitude="151.27611" pageId="55" pageNumber="56" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<typeStatus id="255B8842FFB1242AFE67BF87FE1C3B3F" box="[398,507,510,535]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
:
<collectingRegion id="3824F802FFB1242AFDE0BF86FD3F3B3F" box="[521,728,510,535]" country="Australia" name="New South Wales" pageId="55" pageNumber="56">New South Wales</collectingRegion>
,
<location id="FF3F603BFFB1242AFD00BF87FC273B3F" LSID="urn:lsid:plazi:treatment:724987F6FFB12424FF7EBFCEFF07384D:FF3F603BFFB1242AFD00BF87FC273B3F" box="[745,960,510,535]" latitude="-31.448055" longLatPrecision="20" longitude="151.27611" name="South Head Creek" pageId="55" pageNumber="56" stateProvince="New South Wales">South Head Creek</location>
, on track
<elevation id="71CDD1D3FFB1242AFBD1BF87FB653B3E" box="[1080,1154,510,535]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="55" pageNumber="56" unit="m" value="500.0">
<quantity id="3D189B05FFB1242AFBD1BF87FB653B3E" box="[1080,1154,510,535]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="55" pageNumber="56" unit="m" value="500.0">500 m</quantity>
</elevation>
SW from road
<locationDeviation id="EBED2FC2FFB1242AFAD7BF87FD2C3B13" firstLocation="Snowball Road" pageId="55" pageNumber="56" secondLocation="Nundle Forest Way, Nundle">
between Snowball Road and Nundle Forest Way,
<collectorName id="57155336FFB1242AFD91BC5BFD2C3B13" box="[632,715,546,571]" pageId="55" pageNumber="56">Nundle</collectorName>
</locationDeviation>
SF,
<geoCoordinate id="9FD45027FFB1242AFCEFBC5AFC4F3B13" box="[774,936,546,571]" direction="south" orientation="latitude" pageId="55" pageNumber="56" precision="15" value="-31.448055">31° 26' 53&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FFB1242AFC50BC5BFB8B3B12" box="[953,1132,546,571]" direction="east" orientation="longitude" pageId="55" pageNumber="56" precision="15" value="151.27611">151° 16' 34&quot; E</geoCoordinate>
, on weeds and clumps of saturated moss,
<collectorName id="57155336FFB1242AFEBFBC3EFE003B77" box="[342,487,582,607]" pageId="55" pageNumber="56">W.F. Ponder</collectorName>
,
<collectingDate id="9E1AE9C8FFB1242AFE11BC3EFD753B77" box="[504,658,582,607]" pageId="55" pageNumber="56" value="1988-02-23">
<date id="8E5E1020FFB1242AFE11BC3EFD753B77" box="[504,658,582,607]" pageId="55" pageNumber="56" value="1988-02-23">23 Feb 1988</date>
</collectingDate>
, C.479959
</materialsCitation>
.
<typeStatus id="255B8842FFB1242AFCCDBC3EFC723B77" box="[804,917,583,607]" pageId="55" pageNumber="56" type="paratype">Paratypes</typeStatus>
: Same data, C.
<date id="8E5E1020FFB1242AFBA0BC3EFB443B77" box="[1097,1187,583,607]" pageId="55" pageNumber="56" value="00-09-20">200900</date>
,
<materialsCitation id="4A883CBDFFB1242AFB5DBC3EFCC83BE3" ID-GBIF-Occurrence="2235592663" collectingDate="1988-02-23" collectorName="W. F. Ponder" county="Other" elevation="200" latitude="-31.4675" location="Nundle" longLatPrecision="20" longitude="151.2575" municipality="Nundle" pageId="55" pageNumber="56" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
20+.
<collectingCounty id="133E4E6CFFB1242AFB1DBC3FFAD33B77" box="[1268,1332,582,607]" pageId="55" pageNumber="56">Other</collectingCounty>
material examined:
<collectingRegion id="3824F802FFB1242AFEFDBC12FE053BAB" box="[276,482,618,643]" country="Australia" name="New South Wales" pageId="55" pageNumber="56">New South Wales</collectingRegion>
, E of
<collectingMunicipality id="1A3BAC9AFFB1242AFDCEBC13FD9B3BAB" box="[551,636,618,643]" pageId="55" pageNumber="56">Nundle</collectingMunicipality>
,
<location id="FF3F603BFFB1242AFD60BC13FD3B3BAB" LSID="urn:lsid:plazi:treatment:724987F6FFB12424FF7EBFCEFF07384D:FF3F603BFFB1242AFD60BC13FD3B3BAB" box="[649,732,618,643]" county="Other" latitude="-31.4675" longLatPrecision="20" longitude="151.2575" municipality="Nundle" name="Nundle" pageId="55" pageNumber="56" stateProvince="New South Wales">Nundle</location>
S.F,
<location id="FF3F603BFFB1242AFCFEBC13FBFE3BAB" LSID="urn:lsid:plazi:treatment:724987F6FFB12424FF7EBFCEFF07384D:FF3F603BFFB1242AFCFEBC13FBFE3BAB" box="[791,1049,618,643]" county="Other" latitude="-31.4675" longLatPrecision="20" longitude="151.2575" municipality="Nundle" name="Ponderosa Forest Park" pageId="55" pageNumber="56" stateProvince="New South Wales">
<location id="FF3F603BFFB1242AFCFEBC13FC3D3BAA" LSID="urn:lsid:plazi:treatment:724987F6FFB12424FF7EBFCEFF07384D:FF3F603BFFB1242AFCFEBC13FC3D3BAA" box="[791,986,618,643]" county="Other" latitude="-31.4675" longLatPrecision="20" longitude="151.2575" municipality="Nundle" name="Ponderosa Forest" pageId="55" pageNumber="56" stateProvince="New South Wales">Ponderosa Forest</location>
Park
</location>
,
<location id="FF3F603BFFB1242AFBCFBC13FABF3BAA" LSID="urn:lsid:plazi:treatment:724987F6FFB12424FF7EBFCEFF07384D:FF3F603BFFB1242AFBCFBC13FABF3BAA" box="[1062,1368,618,643]" county="Other" latitude="-31.4675" longLatPrecision="20" longitude="151.2575" municipality="Nundle" name="Hanging Rock State Forest" pageId="55" pageNumber="56" stateProvince="New South Wales">Hanging Rock State Forest</location>
,
<location id="FF3F603BFFB1242AFA8EBC12FEF13B8F" LSID="urn:lsid:plazi:treatment:724987F6FFB12424FF7EBFCEFF07384D:FF3F603BFFB1242AFA8EBC12FEF13B8F" county="Other" latitude="-31.4675" longLatPrecision="20" longitude="151.2575" municipality="Nundle" name="Four Mile Creek" pageId="55" pageNumber="56" stateProvince="New South Wales">Four Mile Creek</location>
tributary, ca.
<elevation id="71CDD1D3FFB1242AFE46BCF6FE113B8E" box="[431,502,655,679]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="m" value="200.0">
<quantity id="3D189B05FFB1242AFE46BCF6FE113B8E" box="[431,502,655,679]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="m" value="200.0">200 m</quantity>
</elevation>
E of forestry camp,
<geoCoordinate id="9FD45027FFB1242AFD32BCF6FC973B8F" box="[731,880,655,679]" direction="south" orientation="latitude" pageId="55" pageNumber="56" precision="15" value="-31.4675">31° 28' 03&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FFB1242AFC94BCF7FBC53B8E" box="[893,1058,654,679]" direction="east" orientation="longitude" pageId="55" pageNumber="56" precision="15" value="151.2575">151° 15' 27&quot; E</geoCoordinate>
, small slow-flowing clear stream, in moss and liverworts on rocks,
<collectorName id="57155336FFB1242AFDE3BCCAFD723BE3" box="[522,661,690,715]" pageId="55" pageNumber="56">W.F. Ponder</collectorName>
,
<collectingDate id="9E1AE9C8FFB1242AFD49BCCAFCC83BE3" box="[672,815,690,715]" pageId="55" pageNumber="56" value="1988-02-23">
<date id="8E5E1020FFB1242AFD49BCCAFCC83BE3" box="[672,815,690,715]" pageId="55" pageNumber="56" value="1988-02-23">23 Feb 1988</date>
</collectingDate>
</materialsCitation>
, C.400874, 1; C.410644, 8.
</paragraph>
<paragraph id="FA5F36E0FFB1242AFF2EBCAFFAE13A8B" blockId="55.[151,1437,510,931]" pageId="55" pageNumber="56">
<emphasis id="C894EAF2FFB1242AFF2EBCAFFEE43BC7" bold="true" box="[199,259,726,751]" pageId="55" pageNumber="56">Shell</emphasis>
(
<figureCitation id="62DB2A65FFB1242AFEFABCAFFE7B3BC7" box="[275,412,726,751]" captionStart="FIGURE 28" captionStartId="56.[151,250,638,660]" captionTargetBox="[169,1416,198,616]" captionTargetId="figure@56.[150,1436,193,622]" captionTargetPageId="56" captionText="FIGURE 28. Shell and operculum of Nundalia secreta n. gen. and n. sp. A. Shell of holotype (C. 479959). B. Protoconch (C.200900). C, D. Operculum, outer (C), and inner (D) sides (C.200900)." httpUri="https://zenodo.org/record/2637663/files/figure.png" pageId="55" pageNumber="56">Fig. 28A, B</figureCitation>
). Ovate-conic, spire outline slightly convex, normally coiled, opaque. Length 1.72.0 mm (mean
<quantity id="3D189B05FFB1242AFF0DBC82FEDA3A3B" box="[228,317,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="55" pageNumber="56" unit="mm" value="1.8">1.8 mm</quantity>
), width
<quantity id="3D189B05FFB1242AFE7EBC82FDF93A3A" box="[407,542,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="55" pageNumber="56" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.11.4 mm</quantity>
(mean
<quantity id="3D189B05FFB1242AFD98BC82FD2C3A3B" box="[625,715,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="55" pageNumber="56" unit="mm" value="1.3">1.3 mm</quantity>
). Protoconch of about 1.5 whorls, with irregular transverse folds and scattered rather irregular pits on first half whorl, no folds and weaker on last whorl. Teleoconch whorls moderately convex, evenly rounded, total number 3.63.9 (mean 3.8). Umbilicus represented by chink only. Aperture pyriform, inner lip narrow, medium, firmly attached to parietal wall or slightly separated along its whole length, outer lip medium. Periostracum moderately developed, white or pale brown or yellow-brown.
</paragraph>
</subSubSection>
<caption id="AE9F6668FFB1242AFF7EBDABFDB73D21" ID-Table-UUID="AE9F6668FFB1242AFF7EBDABFDB73D21" httpUri="http://table.plazi.org/id/AE9F6668FFB1242AFF7EBDABFDB73D21" pageId="55" pageNumber="56" targetBox="[159,1322,1061,1331]" targetIsTable="true" targetPageId="55">
<paragraph id="FA5F36E0FFB1242AFF7EBDABFDB73D21" blockId="55.[151,1436,977,1033]" pageId="55" pageNumber="56">
<emphasis id="C894EAF2FFB1242AFF7EBDABFEFB3AC1" bold="true" box="[151,284,978,1001]" pageId="55" pageNumber="56">TABLE 19.</emphasis>
Shell measurements (in mm) of type material of
<taxonomicName id="3DE04D63FFB1242AFCB2BDA8FBF13AC1" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[859,1046,977,1001]" class="Aves" family="Threskiornithidae" genus="Nundalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelecaniformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="secreta" status="sp. nov.">
<emphasis id="C894EAF2FFB1242AFCB2BDA8FBF13AC1" box="[859,1046,977,1001]" italics="true" pageId="55" pageNumber="56">Nundalia secreta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FFB1242AFBCBBDABFB843AC1" box="[1058,1123,978,1001]" pageId="55" pageNumber="56" rank="species">
<emphasis id="C894EAF2FFB1242AFBCBBDABFB843AC1" bold="true" box="[1058,1123,978,1001]" pageId="55" pageNumber="56">n. sp.</emphasis>
</taxonomicNameLabel>
(paratypes, C.200900). See Materials and Methods for abbreviations.
</paragraph>
</caption>
<paragraph id="FA5F36E0FFB1242AFE24BA5CFACE3C1B" pageId="55" pageNumber="56">
<table id="88E0C440FFB1DBE2FF76BA5CFACD3C1B" box="[159,1322,1061,1331]" gridcols="7" gridrows="7" pageId="55" pageNumber="56">
<tr id="44D034A2FFB1DBE2FF76BA5CFACD3D13" box="[159,1322,1061,1083]" gridrow="0" pageId="55" pageNumber="56" rowspan-0="1">
<th id="07015DDEFFB1DBE2FE24BA5CFE1E3D13" box="[461,505,1061,1083]" gridcol="1" gridrow="0" pageId="55" pageNumber="56">SH</th>
<th id="07015DDEFFB1DBE2FD6DBA5CFD573D13" box="[644,688,1061,1083]" gridcol="2" gridrow="0" pageId="55" pageNumber="56">SW</th>
<th id="07015DDEFFB1DBE2FCF6BA5CFCAC3D13" box="[799,843,1061,1083]" gridcol="3" gridrow="0" pageId="55" pageNumber="56">AH</th>
<th id="07015DDEFFB1DBE2FC46BA5CFC3C3D13" box="[943,987,1061,1083]" gridcol="4" gridrow="0" pageId="55" pageNumber="56">AW</th>
<th id="07015DDEFFB1DBE2FBBFBA5CFB653D13" box="[1110,1154,1061,1083]" gridcol="5" gridrow="0" pageId="55" pageNumber="56">BW</th>
<th id="07015DDEFFB1DBE2FB17BA5CFACD3D13" box="[1278,1322,1061,1083]" gridcol="6" gridrow="0" pageId="55" pageNumber="56">TW</th>
</tr>
<tr id="44D034A2FFB1DBE2FF76BA37FACD3D4C" box="[159,1322,1102,1124]" gridrow="1" pageId="55" pageNumber="56">
<th id="07015DDEFFB1DBE2FF76BA37FF1A3D4C" box="[159,253,1102,1124]" gridcol="0" gridrow="1" pageId="55" pageNumber="56">Holotype</th>
<td id="07015DDEFFB1DBE2FE24BA37FE1E3D4C" box="[461,505,1102,1124]" gridcol="1" gridrow="1" pageId="55" pageNumber="56">1.89</td>
<td id="07015DDEFFB1DBE2FD6DBA37FD573D4C" box="[644,688,1102,1124]" gridcol="2" gridrow="1" pageId="55" pageNumber="56">1.34</td>
<td id="07015DDEFFB1DBE2FCF6BA37FCAC3D4C" box="[799,843,1102,1124]" gridcol="3" gridrow="1" pageId="55" pageNumber="56">1.04</td>
<td id="07015DDEFFB1DBE2FC46BA37FC3C3D4C" box="[943,987,1102,1124]" gridcol="4" gridrow="1" pageId="55" pageNumber="56">0.69</td>
<td id="07015DDEFFB1DBE2FBBFBA37FB653D4C" box="[1110,1154,1102,1124]" gridcol="5" gridrow="1" pageId="55" pageNumber="56">1.56</td>
<td id="07015DDEFFB1DBE2FB17BA37FACD3D4C" box="[1278,1322,1102,1124]" gridcol="6" gridrow="1" pageId="55" pageNumber="56">3.5</td>
</tr>
<tr id="44D034A2FFB1DBE2FF76BA01FACD3DA6" box="[159,1322,1144,1166]" gridrow="2" pageId="55" pageNumber="56">
<th id="07015DDEFFB1DBE2FF76BA01FACD3DA6" box="[159,1322,1144,1166]" colspan="7" colspanRight="6" gridcol="0" gridrow="2" pageId="55" pageNumber="56">Paratypes (20)</th>
</tr>
<tr id="44D034A2FFB1DBE2FF76BAD8FACD3D9F" box="[159,1322,1185,1207]" gridrow="3" pageId="55" pageNumber="56">
<th id="07015DDEFFB1DBE2FF76BAD8FF1A3D9F" box="[159,253,1185,1207]" gridcol="0" gridrow="3" pageId="55" pageNumber="56">Min</th>
<td id="07015DDEFFB1DBE2FE24BAD8FE1E3D9F" box="[461,505,1185,1207]" gridcol="1" gridrow="3" pageId="55" pageNumber="56">1.66</td>
<td id="07015DDEFFB1DBE2FD6DBAD8FD573D9F" box="[644,688,1185,1207]" gridcol="2" gridrow="3" pageId="55" pageNumber="56">1.15</td>
<td id="07015DDEFFB1DBE2FCF6BAD8FCAC3D9F" box="[799,843,1185,1207]" gridcol="3" gridrow="3" pageId="55" pageNumber="56">0.76</td>
<td id="07015DDEFFB1DBE2FC46BAD8FC3C3D9F" box="[943,987,1185,1207]" gridcol="4" gridrow="3" pageId="55" pageNumber="56">0.73</td>
<td id="07015DDEFFB1DBE2FBBFBAD8FB653D9F" box="[1110,1154,1185,1207]" gridcol="5" gridrow="3" pageId="55" pageNumber="56">1.39</td>
<td id="07015DDEFFB1DBE2FB17BAD8FACD3D9F" box="[1278,1322,1185,1207]" gridcol="6" gridrow="3" pageId="55" pageNumber="56">3.60</td>
</tr>
<tr id="44D034A2FFB1DBE2FF76BAB3FACD3DC8" box="[159,1322,1226,1248]" gridrow="4" pageId="55" pageNumber="56">
<th id="07015DDEFFB1DBE2FF76BAB3FF1A3DC8" box="[159,253,1226,1248]" gridcol="0" gridrow="4" pageId="55" pageNumber="56">Max</th>
<td id="07015DDEFFB1DBE2FE24BAB3FE1E3DC8" box="[461,505,1226,1248]" gridcol="1" gridrow="4" pageId="55" pageNumber="56">1.98</td>
<td id="07015DDEFFB1DBE2FD6DBAB3FD573DC8" box="[644,688,1226,1248]" gridcol="2" gridrow="4" pageId="55" pageNumber="56">1.41</td>
<td id="07015DDEFFB1DBE2FCF6BAB3FCAC3DC8" box="[799,843,1226,1248]" gridcol="3" gridrow="4" pageId="55" pageNumber="56">0.99</td>
<td id="07015DDEFFB1DBE2FC46BAB3FC3C3DC8" box="[943,987,1226,1248]" gridcol="4" gridrow="4" pageId="55" pageNumber="56">0.90</td>
<td id="07015DDEFFB1DBE2FBBFBAB3FB653DC8" box="[1110,1154,1226,1248]" gridcol="5" gridrow="4" pageId="55" pageNumber="56">1.66</td>
<td id="07015DDEFFB1DBE2FB17BAB3FACD3DC8" box="[1278,1322,1226,1248]" gridcol="6" gridrow="4" pageId="55" pageNumber="56">3.90</td>
</tr>
<tr id="44D034A2FFB1DBE2FF76BA8DFACD3C22" box="[159,1322,1268,1290]" gridrow="5" pageId="55" pageNumber="56">
<th id="07015DDEFFB1DBE2FF76BA8DFF1A3C22" box="[159,253,1268,1290]" gridcol="0" gridrow="5" pageId="55" pageNumber="56">Mean</th>
<td id="07015DDEFFB1DBE2FE24BA8DFE1E3C22" box="[461,505,1268,1290]" gridcol="1" gridrow="5" pageId="55" pageNumber="56">1.82</td>
<td id="07015DDEFFB1DBE2FD6DBA8DFD573C22" box="[644,688,1268,1290]" gridcol="2" gridrow="5" pageId="55" pageNumber="56">1.26</td>
<td id="07015DDEFFB1DBE2FCF6BA8DFCAC3C22" box="[799,843,1268,1290]" gridcol="3" gridrow="5" pageId="55" pageNumber="56">0.88</td>
<td id="07015DDEFFB1DBE2FC46BA8DFC3C3C22" box="[943,987,1268,1290]" gridcol="4" gridrow="5" pageId="55" pageNumber="56">0.82</td>
<td id="07015DDEFFB1DBE2FBBFBA8DFB653C22" box="[1110,1154,1268,1290]" gridcol="5" gridrow="5" pageId="55" pageNumber="56">1.51</td>
<td id="07015DDEFFB1DBE2FB17BA8DFACD3C22" box="[1278,1322,1268,1290]" gridcol="6" gridrow="5" pageId="55" pageNumber="56">3.77</td>
</tr>
<tr id="44D034A2FFB1DBE2FF76BB64FACD3C1B" box="[159,1322,1309,1331]" gridrow="6" pageId="55" pageNumber="56">
<th id="07015DDEFFB1DBE2FF76BB64FF1A3C1B" box="[159,253,1309,1331]" gridcol="0" gridrow="6" pageId="55" pageNumber="56">SD</th>
<td id="07015DDEFFB1DBE2FE24BB64FE1E3C1B" box="[461,505,1309,1331]" gridcol="1" gridrow="6" pageId="55" pageNumber="56">0.09</td>
<td id="07015DDEFFB1DBE2FD6DBB64FD573C1B" box="[644,688,1309,1331]" gridcol="2" gridrow="6" pageId="55" pageNumber="56">0.07</td>
<td id="07015DDEFFB1DBE2FCF6BB64FCAC3C1B" box="[799,843,1309,1331]" gridcol="3" gridrow="6" pageId="55" pageNumber="56">0.07</td>
<td id="07015DDEFFB1DBE2FC46BB64FC3C3C1B" box="[943,987,1309,1331]" gridcol="4" gridrow="6" pageId="55" pageNumber="56">0.05</td>
<td id="07015DDEFFB1DBE2FBBFBB64FB653C1B" box="[1110,1154,1309,1331]" gridcol="5" gridrow="6" pageId="55" pageNumber="56">0.07</td>
<td id="07015DDEFFB1DBE2FB17BB64FACD3C1B" box="[1278,1322,1309,1331]" gridcol="6" gridrow="6" pageId="55" pageNumber="56">0.11</td>
</tr>
</table>
</paragraph>
<subSubSection id="B2FA656BFFB12425FF2EBBE8FD903EFF" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="description">
<paragraph id="FA5F36E0FFB1242AFF2EBBE8FA683C82" blockId="55.[151,1437,1425,2026]" box="[199,1423,1425,1450]" pageId="55" pageNumber="56">
<emphasis id="C894EAF2FFB1242AFF2EBBE8FEAB3C82" bold="true" box="[199,332,1425,1450]" pageId="55" pageNumber="56">Operculum</emphasis>
(
<figureCitation id="62DB2A65FFB1242AFEB2BBE8FE063C82" box="[347,481,1425,1450]" captionStart="FIGURE 28" captionStartId="56.[151,250,638,660]" captionTargetBox="[169,1416,198,616]" captionTargetId="figure@56.[150,1436,193,622]" captionTargetPageId="56" captionText="FIGURE 28. Shell and operculum of Nundalia secreta n. gen. and n. sp. A. Shell of holotype (C. 479959). B. Protoconch (C.200900). C, D. Operculum, outer (C), and inner (D) sides (C.200900)." httpUri="https://zenodo.org/record/2637663/files/figure.png" pageId="55" pageNumber="56">Fig. 28C, D</figureCitation>
). Transparent, pale yellow, slightly concave; with 23 small, white pegs on inner side.
</paragraph>
<paragraph id="FA5F36E0FFB1242AFF2EBBCCFD2D3CE6" blockId="55.[151,1437,1425,2026]" box="[199,714,1461,1486]" pageId="55" pageNumber="56">
<emphasis id="C894EAF2FFB1242AFF2EBBCCFDC43CE6" bold="true" box="[199,547,1461,1486]" pageId="55" pageNumber="56">Head-foot and external body.</emphasis>
Unpigmented.
</paragraph>
<paragraph id="FA5F36E0FFB1242AFF2EBBA0FBAC3FAA" blockId="55.[151,1437,1425,2026]" pageId="55" pageNumber="56">
<emphasis id="C894EAF2FFB1242AFF2EBBA0FE9E3CDA" bold="true" box="[199,377,1497,1522]" pageId="55" pageNumber="56">Mantle cavity.</emphasis>
Ctenidium reduced, filaments 1215, broadly triangular, apex right edge. Osphradium narrowly oval, towards posterior end of ctenidium, length relative to gill 0.210.28. Hypobranchial gland not distinguishable (absent). Rectum with U-shaped bend, faecal pellets longitudinally orientated, anus at or very near mantle collar or behind it. Kidney half or more in roof of mantle cavity. Renal gland transverse. Pericardium half or more in roof of mantle cavity, with short separation from posterior end of ctenidium.
</paragraph>
<paragraph id="FA5F36E0FFB1242AFF2EB8F4FEE73E39" blockId="55.[151,1437,1425,2026]" pageId="55" pageNumber="56">
<emphasis id="C894EAF2FFB1242AFF2EB8F4FEF93F8E" bold="true" box="[199,286,1677,1702]" pageId="55" pageNumber="56">Radula</emphasis>
(
<figureCitation id="62DB2A65FFB1242AFED8B8F4FE6F3F8E" box="[305,392,1677,1702]" captionStart="FIGURE 29" captionStartId="56.[151,250,1114,1136]" captionTargetBox="[371,1216,743,1088]" captionTargetId="figure@56.[367,1220,740,1091]" captionTargetPageId="56" captionText="FIGURE 29. Radula of Nundalia secreta n. gen. and n. sp. (C.200900). A. Half row of radula. B. Detail of central teeth. Scale bar 10 µm." httpUri="https://zenodo.org/record/2637665/files/figure.png" pageId="55" pageNumber="56">Fig. 29</figureCitation>
). Central teeth with cusp formula 36+1+36, basal cusps 2+2; median cusp sharp, about quarter to twice as long as adjacent cusps. Lateral teeth with cusp formula 45+1+56, main cusp narrow, little longer than adjacent cusps. Inner marginal teeth with about 2124 cusps. Outer marginal teeth with approximately 22 cusps.
</paragraph>
<paragraph id="FA5F36E0FFB1242AFF2EB964FC2B3EC2" blockId="55.[151,1437,1425,2026]" pageId="55" pageNumber="56">
<emphasis id="C894EAF2FFB1242AFF2EB964FDF13E1E" bold="true" box="[199,534,1821,1846]" pageId="55" pageNumber="56">Female reproductive system</emphasis>
(
<figureCitation id="62DB2A65FFB1242AFDCFB964FD493E1E" box="[550,686,1821,1846]" captionStart="FIGURE 30" captionStartId="56.[151,250,1679,1701]" captionTargetBox="[188,1418,1235,1645]" captionTargetId="figure@56.[151,1435,1216,1665]" captionTargetPageId="56" captionText="FIGURE 30. Male and female reproductive anatomy of Nundalia secreta n. gen. and n. sp. (C.200900). A. Penis, showing penial duct and vas deferens. B. Female system from left side. C. Bursa copulatrix and seminal receptacle. Abbreviations: agalbumen gland; bc—bursa copulatrix; cg—capsule gland; o—opening of oviduct; p—papilla; pd—penial duct; pw—posterior wall of mantle cavity; ro—renal oviduct; sr—seminal receptacle; v—vestibule; vc—ventral channel. Scale bar 200 µm." httpUri="https://zenodo.org/record/2637667/files/figure.png" pageId="55" pageNumber="56">Fig. 30B, C</figureCitation>
). Ovary simple sac. Renal oviduct with short, anteriorly orientated U-shaped arch. Seminal receptacle near anterior edge of bursa, elongate, duct intermediate. Bursa copulatrix behind albumen gland, elongately-oval or triangular, shorter than albumen gland, bursal duct enters bursa anteroventrally, bursal duct joins coiled oviduct well behind posterior mantle cavity wall. Albumen gland partly in mantle cavity. Capsule gland with two distinct glandular zones, medium thickness in cross section, markedly indented by rectum. Anterior vestibule small, opening anterior to capsule gland, long.
</paragraph>
<caption id="AE9F6668FFBE2425FF7EBC07FC993B9B" httpUri="https://zenodo.org/record/2637663/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,638,660]" targetBox="[169,1416,198,616]" targetPageId="56">
<paragraph id="FA5F36E0FFBE2425FF7EBC07FC993B9B" blockId="56.[151,1436,638,691]" pageId="56" pageNumber="57">
<emphasis id="C894EAF2FFBE2425FF7EBC07FEC53BBC" bold="true" box="[151,290,638,660]" pageId="56" pageNumber="57">FIGURE 28.</emphasis>
Shell and operculum of
<taxonomicName id="3DE04D63FFBE2425FDCFBC07FD323BBC" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[550,725,638,660]" class="Aves" family="Threskiornithidae" genus="Nundalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelecaniformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="secreta" status="n. gen. and n. sp.">
<emphasis id="C894EAF2FFBE2425FDCFBC07FD323BBC" box="[550,725,638,660]" italics="true" pageId="56" pageNumber="57">Nundalia secreta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FFBE2425FD37BC07FC7F3BBC" box="[734,920,638,660]" pageId="56" pageNumber="57" rank="species">
<taxonomicNameLabel id="D3A75789FFBE2425FD37BC07FCC03BBC" box="[734,807,638,660]" pageId="56" pageNumber="57" rank="genus">
<emphasis id="C894EAF2FFBE2425FD37BC07FCC03BBC" bold="true" box="[734,807,638,660]" pageId="56" pageNumber="57">n. gen.</emphasis>
</taxonomicNameLabel>
and
<emphasis id="C894EAF2FFBE2425FCB4BC07FC7F3BBC" bold="true" box="[861,920,638,660]" pageId="56" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
A. Shell of holotype (C. 479959). B. Protoconch (C.200900). C, D. Operculum, outer (C), and inner (D) sides (C.200900).
</paragraph>
</caption>
<caption id="AE9F6668FFBE2425FF7EBA23FEE23DA6" httpUri="https://zenodo.org/record/2637665/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1114,1136]" targetBox="[371,1216,743,1088]" targetPageId="56">
<paragraph id="FA5F36E0FFBE2425FF7EBA23FEE23DA6" blockId="56.[151,1436,1114,1166]" pageId="56" pageNumber="57">
<emphasis id="C894EAF2FFBE2425FF7EBA23FEF83D58" bold="true" box="[151,287,1114,1136]" pageId="56" pageNumber="57">FIGURE 29.</emphasis>
Radula of
<taxonomicName id="3DE04D63FFBE2425FE67BA23FDDC3D58" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[398,571,1114,1136]" class="Aves" family="Threskiornithidae" genus="Nundalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelecaniformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="secreta" status="n. gen. and n. sp.">
<emphasis id="C894EAF2FFBE2425FE67BA23FDDC3D58" box="[398,571,1114,1136]" italics="true" pageId="56" pageNumber="57">Nundalia secreta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FFBE2425FDA8BA23FD153D58" box="[577,754,1114,1136]" pageId="56" pageNumber="57" rank="species">
<taxonomicNameLabel id="D3A75789FFBE2425FDA8BA23FD6E3D58" box="[577,649,1114,1136]" pageId="56" pageNumber="57" rank="genus">
<emphasis id="C894EAF2FFBE2425FDA8BA23FD6E3D58" bold="true" box="[577,649,1114,1136]" pageId="56" pageNumber="57">n. gen.</emphasis>
</taxonomicNameLabel>
and
<emphasis id="C894EAF2FFBE2425FD50BA23FD153D58" bold="true" box="[697,754,1114,1136]" pageId="56" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
(C.200900). A. Half row of radula. B. Detail of central teeth. Scale bar 10 µm.
</paragraph>
</caption>
<caption id="AE9F6668FFBE2425FF7EB8F6FAB73E29" httpUri="https://zenodo.org/record/2637667/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1679,1701]" targetBox="[188,1418,1235,1645]" targetPageId="56">
<paragraph id="FA5F36E0FFBE2425FF7EB8F6FAB73E29" blockId="56.[151,1436,1679,1793]" pageId="56" pageNumber="57">
<emphasis id="C894EAF2FFBE2425FF7EB8F6FEC53F8D" bold="true" box="[151,290,1679,1701]" pageId="56" pageNumber="57">FIGURE 30.</emphasis>
Male and female reproductive anatomy of
<taxonomicName id="3DE04D63FFBE2425FD0CB8F6FC743F8E" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[741,915,1679,1702]" class="Aves" family="Threskiornithidae" genus="Nundalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelecaniformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="secreta" status="n. gen. and n. sp.">
<emphasis id="C894EAF2FFBE2425FD0CB8F6FC743F8E" box="[741,915,1679,1702]" italics="true" pageId="56" pageNumber="57">Nundalia secreta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FFBE2425FC75B8F6FBB23F8D" box="[924,1109,1679,1701]" pageId="56" pageNumber="57" rank="species">
<taxonomicNameLabel id="D3A75789FFBE2425FC75B8F6FC023F8D" box="[924,997,1679,1701]" pageId="56" pageNumber="57" rank="genus">
<emphasis id="C894EAF2FFBE2425FC75B8F6FC023F8D" bold="true" box="[924,997,1679,1701]" pageId="56" pageNumber="57">n. gen.</emphasis>
</taxonomicNameLabel>
and
<emphasis id="C894EAF2FFBE2425FBF3B8F6FBB23F8D" bold="true" box="[1050,1109,1679,1701]" pageId="56" pageNumber="57">n. sp.</emphasis>
</taxonomicNameLabel>
(C.200900). A. Penis, showing penial duct and vas deferens. B. Female system from left side. C. Bursa copulatrix and seminal receptacle. Abbreviations: agalbumen gland; bc—bursa copulatrix; cg—capsule gland; o—opening of oviduct; p—papilla; pd—penial duct; pw—posterior wall of mantle cavity; ro—renal oviduct; sr—seminal receptacle; v—vestibule; vc—ventral channel. Scale bar 200 µm.
</paragraph>
</caption>
<paragraph id="FA5F36E0FFBE2425FF2EB92BFD903EFF" blockId="56.[151,1437,1874,2007]" pageId="56" pageNumber="57">
<emphasis id="C894EAF2FFBE2425FF2EB92BFE183E43" bold="true" box="[199,511,1874,1899]" pageId="56" pageNumber="57">Male reproductive system</emphasis>
(
<figureCitation id="62DB2A65FFBE2425FDF9B92BFD9D3E43" box="[528,634,1874,1899]" captionStart="FIGURE 30" captionStartId="56.[151,250,1679,1701]" captionTargetBox="[188,1418,1235,1645]" captionTargetId="figure@56.[151,1435,1216,1665]" captionTargetPageId="56" captionText="FIGURE 30. Male and female reproductive anatomy of Nundalia secreta n. gen. and n. sp. (C.200900). A. Penis, showing penial duct and vas deferens. B. Female system from left side. C. Bursa copulatrix and seminal receptacle. Abbreviations: agalbumen gland; bc—bursa copulatrix; cg—capsule gland; o—opening of oviduct; p—papilla; pd—penial duct; pw—posterior wall of mantle cavity; ro—renal oviduct; sr—seminal receptacle; v—vestibule; vc—ventral channel. Scale bar 200 µm." httpUri="https://zenodo.org/record/2637667/files/figure.png" pageId="56" pageNumber="57">Fig. 30A</figureCitation>
). Prostate gland mostly in mantle roof, oval, medium in cross section. Posterior pallial vas deferens straight or slightly undulating, anteriorly slightly undulating. Penis towards middle of head, intermediate, distal end tapering or narrow, blunt, terminal papilla absent or small.
<emphasis id="C894EAF2FFBE2425FF2EB9C7FEA93EFF" bold="true" box="[199,334,1982,2007]" pageId="56" pageNumber="57">Etymology.</emphasis>
Secretus—Latin, isolated.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFFBF2424FF2EBEEEFEC13834" pageId="57" pageNumber="58" type="distribution">
<paragraph id="FA5F36E0FFBF2424FF2EBEEEFEC13834" blockId="57.[151,1436,151,357]" pageId="57" pageNumber="58">
<emphasis id="C894EAF2FFBF2424FF2EBEEEFE123998" bold="true" box="[199,501,151,176]" pageId="57" pageNumber="58">Distribution and habitat.</emphasis>
This tiny species is known from only two localities in the Great Dividing Range near Nundle (
<figureCitation id="62DB2A65FFBF2424FED3BEC5FE6739FC" box="[314,384,188,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1844,1866]" captionTargetBox="[151,1435,498,1802]" captionTargetId="figure@4.[151,1435,498,1802]" captionTargetPageId="4" captionText="FIGURE 1. Map of the spring supergroups associated with the Great Artesian Basin (GAB) in Queensland, showing the main non-town locations mentioned in the text (squares) and some of the towns in the area (ovals). The pale grey shading is the discharge part of the GAB, the dark grey the recharge part. The inset map of Australia shows the area illustrated. The approximate location of Conondale is indicated by a triangle and Nundle by a star. The spring supergroups are outlined with dotted lines and are as follows: A, Mulligan River; B, Springvale; C, Flinders River; D, Hughenden; E, Barcaldine (see inset at upper right for details); F, Springsure; G, Eulo." httpUri="https://zenodo.org/record/2637605/files/figure.png" pageId="57" pageNumber="58">Fig. 1</figureCitation>
) in the upper tributaries of the east-flowing Manning River. It is probably more widely distributed in that area than present sampling suggests. It is also possible that other similar species exist in this general area.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFFBF2424FF2EBF5EFF07384D" pageId="57" pageNumber="58" type="discussion">
<paragraph id="FA5F36E0FFBF2424FF2EBF5EFF07384D" blockId="57.[151,1436,151,357]" pageId="57" pageNumber="58">
<emphasis id="C894EAF2FFBF2424FF2EBF5EFEDC3868" bold="true" box="[199,315,295,320]" pageId="57" pageNumber="58">Remarks.</emphasis>
This species differs from other Eastern Australian
<taxonomicName id="3DE04D63FFBF2424FC95BF5EFC3D3868" box="[892,986,295,320]" class="Gastropoda" family="Tateidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="57" pageNumber="58" phylum="Mollusca" rank="family">Tateidae</taxonomicName>
in the characters given above under the genus.
</paragraph>
</subSubSection>
</treatment>
</document>