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<document id="931573AADF554716EE90DE52060C5C6E" ID-CLB-Dataset="27316" ID-DOI="10.11646/zootaxa.4583.1.1" ID-GBIF-Dataset="027ed27d-5035-4646-93a6-c23b565f3d20" ID-ISSN="1175-5326" ID-Zenodo-Dep="2637603" ID-ZooBank="27F24995-359E-46F6-AB22-75568BACFDCF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555054242242" checkinUser="plazi" docAuthor="Zhang, - H." docDate="2019" docId="724987F6FF942409FF7EB9F4FCA7384D" docLanguage="en" docName="zootaxa.4583.1.1.pdf" docOrigin="Zootaxa 4583 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Eulodrobia carinata Zhang 2019, n. sp." docType="treatment" docVersion="15" lastPageNumber="21" masterDocId="8E70FF8EFF86241DFFE9BE79FFE73928" masterDocTitle="New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages" masterLastPageNumber="67" masterPageNumber="1" pageNumber="19" updateTime="1698720421338" updateUser="ExternalLinkService">
<mods:mods id="DEE6FEFB43AED482B0A75A276CB36849" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="07D4A3A7CE2986A2B99458C2021DBB1E">
<mods:title id="5BEA25327158819280216C94C23AB9B4">New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages</mods:title>
</mods:titleInfo>
<mods:name id="8510A87A1CFCCD4FE86341EFC247F710" type="personal">
<mods:role id="93BFBDD01F2EB43DE4207C6C11FD1111">
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</mods:role>
<mods:namePart id="2D6427789D803B4EBFDB3C44C170868A">Zhang, - H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7ED790A8A7B9A2AD1C7657DFD86BB057">text</mods:typeOfResource>
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<mods:title id="0932BD93EE3367060552685E07A0BF2F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7210D20F7D8FEF673BE693745519F59F">
<mods:date id="F1753194995F6368C4E2DE46DA01B8D9">2019</mods:date>
<mods:detail id="790A4563ABBE4E1474FF4DEF9EC8E1E5" type="pubDate">
<mods:number id="ECCD4A6AF57F5C52C774C4EC98D7C1D9">2019-04-10</mods:number>
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<mods:detail id="FA98D4B1FB2694C6C41D919287A354C7" type="volume">
<mods:number id="F38CC109F1BA96B4487C9DB2D4CA773A">4583</mods:number>
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<mods:detail id="6B18A06710295F06C7D75EF5AD9E1E8F" type="issue">
<mods:number id="6A79F28DC73B06FA160D55A33DAD8CB6">1</mods:number>
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<mods:classification id="276F101B5C4142A9908EDDE1A01CE2C0">journal article</mods:classification>
<mods:identifier id="358F4044C35366A3253F014E6492142A" type="CLB-Dataset">27316</mods:identifier>
<mods:identifier id="C4AE5EEF57D614ECAC32D7E96A1B67DA" type="DOI">10.11646/zootaxa.4583.1.1</mods:identifier>
<mods:identifier id="DC8E00CEAC764E605980E9BB491D2D59" type="GBIF-Dataset">027ed27d-5035-4646-93a6-c23b565f3d20</mods:identifier>
<mods:identifier id="B70A3F9E1AA8F7A089D699B0DD5E418F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="23799F3860C24E96B8D218CF35EE8456" type="Zenodo-Dep">2637603</mods:identifier>
<mods:identifier id="5E78EDF0E31635C4FC41414826CE34FA" type="ZooBank">27F24995-359E-46F6-AB22-75568BACFDCF</mods:identifier>
</mods:mods>
<treatment id="724987F6FF942409FF7EB9F4FCA7384D" ID-GBIF-Taxon="156198442" LSID="urn:lsid:plazi:treatment:724987F6FF942409FF7EB9F4FCA7384D" httpUri="http://treatment.plazi.org/id/724987F6FF942409FF7EB9F4FCA7384D" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection id="B2FA656BFF94240FFF7EB9F4FE323E8F" box="[151,469,1933,1959]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="FA5F36E0FF94240FFF7EB9F4FE323E8F" blockId="18.[151,469,1933,1959]" box="[151,469,1933,1959]" pageId="18" pageNumber="19">
<heading id="A117818CFF94240FFF7EB9F4FE323E8F" bold="true" box="[151,469,1933,1959]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="C894EAF2FF94240FFF7EB9F4FE323E8F" bold="true" box="[151,469,1933,1959]" pageId="18" pageNumber="19">
<taxonomicName id="3DE04D63FF94240FFF7EB9F4FE6B3E8F" ID-CoL="7R9CT" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[151,396,1933,1959]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="carinata" status="sp. nov.">
<emphasis id="C894EAF2FF94240FFF7EB9F4FE6B3E8F" bold="true" box="[151,396,1933,1959]" italics="true" pageId="18" pageNumber="19">Eulodrobia carinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF94240FFE7AB9F4FE323E8F" box="[403,469,1933,1959]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF94240EFF7EB9ADFAAF3BAD" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="FA5F36E0FF94240EFF7EB9ADFBBE38A0" blockId="18.[151,1045,2004,2026]" lastBlockId="19.[151,1437,151,645]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="C894EAF2FF94240FFF7EB9ADFF193EC2" box="[151,254,2004,2026]" italics="true" pageId="18" pageNumber="19">Jardinella</emphasis>
n. sp. keel
<bibRefCitation id="9E714B11FF94240FFE98B9ACFDF83EC2" author="Perez, K. E. &amp; Ponder, W. F. &amp; Colgan, D. J. &amp; Clark, S. A. &amp; Lydeard, C." box="[369,543,2004,2026]" pageId="18" pageNumber="19" pagination="545 - 556" refId="ref35351" refString="Perez, K. E., Ponder, W. F., Colgan, D. J., Clark, S. A. &amp; Lydeard, C. (2005) Molecular phylogeny and biogeography of springassociated hydrobiid snails of the Great Artesian Basin, Australia. Molecular Phylogenetics and Evolution, 34, 545 - 556. https: // doi. org / 10.1016 / j. ympev. 2004.11.020" type="journal article" year="2005">
Perez
<emphasis id="C894EAF2FF94240FFE59B9ACFE043EC2" box="[432,483,2004,2026]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2005
</bibRefCitation>
: 547 (incorrectly cited as C.400132 = C.400131).
<emphasis id="C894EAF2FF95240EFF7EBEEEFE663998" bold="true" box="[151,385,151,176]" pageId="19" pageNumber="20">Material examined.</emphasis>
<materialsCitation id="4A883CBDFF95240EFE60BEEEFBB238A0" ID-GBIF-Occurrence="2599922915" collectingDate="2001-05-02" collectingDateMax="2002-04-04" collectingDateMin="2001-05-02" collectionCode="QM" collectorName="W. F. Ponder &amp; J. H. Waterhouse &amp; A. C. Miller &amp; J. H. Waterhouse &amp; A. C. Miller" county="Same" latitude="-27.951944" location="Bundoona Station" longLatPrecision="20" longitude="144.76889" municipality="Yowah Creek Springs" pageId="19" pageNumber="20" specimenCode="MO85764, 5" specimenCount="3" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="255B8842FF95240EFE60BEEEFE123998" box="[393,501,151,176]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
:
<collectingRegion id="3824F802FF95240EFDE8BEEEFD6F3998" box="[513,648,151,176]" country="Australia" name="Queensland" pageId="19" pageNumber="20">Queensland</collectingRegion>
, Yowah Creek Springs, on Bundoona Station,
<geoCoordinate id="9FD45027FF95240EFB7EBEE1FAD73998" box="[1175,1328,151,176]" degrees="27" direction="south" minutes="57" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="08" value="-27.952223">27° 57' 08&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF95240EFAD4BEE1FF3F39FC" degrees="144" direction="east" minutes="46" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="12" value="144.76999">144° 46' 12&quot; E</geoCoordinate>
, in flowing water, at spring head and outflow, up to
<quantity id="3D189B05FF95240EFCC9BEC5FC6839FC" box="[800,911,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="19" pageNumber="20" unit="cm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 cm</quantity>
from spring head,
<collectingRegion id="3824F802FF95240EFB8DBEC5FB0C39FD" box="[1124,1259,188,213]" country="Australia" name="Queensland" pageId="19" pageNumber="20">Queensland</collectingRegion>
, W.F. Ponder &amp; C. Lydeard,
<date id="8E5E1020FF95240EFEC3BE99FE5A39D0" box="[298,445,224,248]" pageId="19" pageNumber="20" value="2001-05-02">
<collectingDate id="9E1AE9C8FF95240EFEC3BE99FE5A39D0" box="[298,445,224,248]" pageId="19" pageNumber="20" value="2001-05-02">2 May 2001</collectingDate>
</date>
, C.479942.
<typeStatus id="255B8842FF95240EFDA4BE99FD5939D0" box="[589,702,224,248]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
:
<collectingCounty id="133E4E6CFF95240EFD24BE99FCEC39D0" box="[717,779,224,248]" pageId="19" pageNumber="20">Same</collectingCounty>
data, C.400131, 50+;
<collectionCode id="9CF1AE25FF95240EFBFFBE99FBA439D0" box="[1046,1091,224,248]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" pageId="19" pageNumber="20">QM</collectionCode>
<specimenCode id="AA469E9BFF95240EFBA6BEA6FB2239D0" box="[1103,1221,223,248]" collectionCode="QM" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="19" pageNumber="20" type="Museum">MO85764</specimenCode>
,
<specimenCode id="AA469E9BFF95240EFB3CBEA6FB0139D0" box="[1237,1254,223,248]" collectionCode="QM" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="19" pageNumber="20" type="Museum">5</specimenCode>
. Other material examined:
<collectingRegion id="3824F802FF95240EFEFDBF7DFE7C3835" box="[276,411,260,285]" country="Australia" name="Queensland" pageId="19" pageNumber="20">Queensland</collectingRegion>
,
<collectingMunicipality id="1A3BAC9AFF95240EFE41BF7DFD443835" box="[424,675,260,285]" pageId="19" pageNumber="20">Yowah Creek Springs</collectingMunicipality>
, on
<location id="FF3F603BFF95240EFD3FBF7DFC463835" LSID="urn:lsid:plazi:treatment:724987F6FF942409FF7EB9F4FCA7384D:FF3F603BFF95240EFD3FBF7DFC463835" box="[726,929,260,285]" county="Same" latitude="-27.951944" longLatPrecision="20" longitude="144.76889" municipality="Yowah Creek Springs" name="Bundoona Station" pageId="19" pageNumber="20" stateProvince="Queensland">Bundoona Station</location>
, main spring, head of spring,
<geoCoordinate id="9FD45027FF95240EFB12BF7DFA703834" box="[1275,1431,260,285]" degrees="27" direction="south" minutes="57" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="07" value="-27.951944">27° 57' 07&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF95240EFF7EBF51FEA73817" box="[151,320,295,320]" degrees="144" direction="east" minutes="46" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="08" value="144.76889">144° 46' 08&quot; E</geoCoordinate>
, on wet sand, mostly out of water flow,
<collectorName id="57155336FF95240EFCE2BF51FC703868" box="[779,919,295,320]" pageId="19" pageNumber="20">W.F. Ponder</collectorName>
,
<collectorName id="57155336FF95240EFC4DBF51FBBA3868" box="[932,1117,295,320]" pageId="19" pageNumber="20">J.H. Waterhouse</collectorName>
&amp;
<collectorName id="57155336FF95240EFB6BBF51FAE03868" box="[1154,1287,295,320]" pageId="19" pageNumber="20">A.C. Miller</collectorName>
,
<collectingDate id="9E1AE9C8FF95240EFAFABF51FA7F3868" box="[1299,1432,296,320]" pageId="19" pageNumber="20" value="2002-04-04">
<date id="8E5E1020FF95240EFAFABF51FA7F3868" box="[1299,1432,296,320]" pageId="19" pageNumber="20" value="2002-04-04">4 Apr 2002</date>
</collectingDate>
, C.421805, 2; main spring, area around head of spring,
<geoCoordinate id="9FD45027FF95240EFD14BF35FC74384C" box="[765,915,332,357]" degrees="27" direction="south" minutes="57" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="07" value="-27.951944">27° 57' 07&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF95240EFC49BF35FBA1384C" box="[928,1094,332,357]" degrees="144" direction="east" minutes="46" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="08" value="144.76889">144° 46' 08&quot; E</geoCoordinate>
, mostly on thick wet mud, out of water flow,
<collectorName id="57155336FF95240EFED3BF09FE2238A0" box="[314,453,367,392]" pageId="19" pageNumber="20">W.F. Ponder</collectorName>
,
<collectorName id="57155336FF95240EFE39BF09FCD738A0" box="[464,816,367,392]" pageId="19" pageNumber="20">J.H. Waterhouse &amp; A.C. Miller</collectorName>
,
<date id="8E5E1020FF95240EFCD3BF09FC5A38A0" box="[826,957,368,392]" pageId="19" pageNumber="20" value="2002-04-04">
<collectingDate id="9E1AE9C8FF95240EFCD3BF09FC5A38A0" box="[826,957,368,392]" pageId="19" pageNumber="20" value="2002-04-04">4 Apr 2002</collectingDate>
</date>
, C.421810, 4
</materialsCitation>
.
</paragraph>
<paragraph id="FA5F36E0FF95240EFF2EBFEDFAAF3BAD" blockId="19.[151,1437,151,645]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EBFEDFEE43885" bold="true" box="[199,259,404,429]" pageId="19" pageNumber="20">Shell</emphasis>
(
<figureCitation id="62DB2A65FF95240EFEFABFEDFE803884" box="[275,359,404,429]" captionStart="FIGURE 2" captionStartId="8.[151,250,1857,1879]" captionTargetBox="[185,1418,742,1819]" captionTargetId="figure@8.[150,1436,697,1836]" captionTargetPageId="8" captionText="FIGURE 2. Shells of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., holotype, spring at Town Common, Eulo (C.479950). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia cf. eulo (Ponder &amp; Clark, 1990), Tunga Spring (C.410643). D. Eulodrobia ovata n. sp., holotype, Yowah Springs, Bundoona Station (C.479944). E. Eulodrobia bundoona n. sp., holotype, Yowah Springs, Bundoona Station (C.479943). F. Eulodrobia carinata n. sp., holotype, Yowah Springs, Bundoona Station (C.479942). G–I. Eulodrobia spirula n. sp., Yowah Springs, Bundoona Station. G. dorsal view of holotype (479945). H, I. apertural view of paratypes (C.400133). Scale bars 1 mm." httpUri="https://zenodo.org/record/2637607/files/figure.png" pageId="19" pageNumber="20">Fig. 2F</figureCitation>
,
<figureCitation id="62DB2A65FF95240EFE9CBFEDFE7F3884" box="[373,408,404,428]" captionStart="FIGURE 3" captionStartId="9.[151,250,1476,1498]" captionTargetBox="[221,1370,184,1448]" captionTargetId="figure@9.[204,1382,169,1464]" captionTargetPageId="9" captionText="FIGURE 3. Protoconchs and protoconch microsculpture of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). D. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). E. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). F. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 100 µm (lowpowered images [left]), 10 µm (high-powered images [right])." httpUri="https://zenodo.org/record/2637609/files/figure.png" pageId="19" pageNumber="20">3E</figureCitation>
). Depressed-ovate, spire outline slightly convex, normally coiled, opaque. Length
<quantity id="3D189B05FF95240EFAA1BFEDFF2438E7" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="19" pageNumber="20" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3–1.6 mm</quantity>
(mean
<quantity id="3D189B05FF95240EFEF1BFCEFE9438F8" box="[280,371,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="20" unit="mm" value="1.5">1.5 mm</quantity>
), width
<quantity id="3D189B05FF95240EFE26BFC1FDB038E7" box="[463,599,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.9" metricValueMin="2.3" pageId="19" pageNumber="20" unit="mm" value="2.5999999999999996" valueMax="2.9" valueMin="2.3">2.3–2.9 mm</quantity>
(mean
<quantity id="3D189B05FF95240EFD45BFCEFCE038F8" box="[684,775,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="20" unit="mm" value="2.6">2.6 mm</quantity>
). Protoconch of about 1.3 whorls, with most of first whorl distinctly pitted and with irregular axial threads; last third whorl with fine pitting and distinct axial ridges (
<figureCitation id="62DB2A65FF95240EFAD0BFA5FA6838DC" box="[1337,1423,476,501]" captionStart="FIGURE 3" captionStartId="9.[151,250,1476,1498]" captionTargetBox="[221,1370,184,1448]" captionTargetId="figure@9.[204,1382,169,1464]" captionTargetPageId="9" captionText="FIGURE 3. Protoconchs and protoconch microsculpture of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). D. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). E. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). F. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 100 µm (lowpowered images [left]), 10 µm (high-powered images [right])." httpUri="https://zenodo.org/record/2637609/files/figure.png" pageId="19" pageNumber="20">Fig. 3E</figureCitation>
). Teleoconch whorls weakly convex, strongly keeled in middle of periphery, total number 3.0–3.5 (mean 3.3). Umbilicus wide, open. Sculpture of growth lines and some weak spiral threads on peripheral area. Aperture circular, inner lip narrow, thin, firmly attached to parietal wall or slightly to moderately separated along whole length of parietal wall, outer lip thin. Periostracum moderately developed, yellow-brown or reddish-brown.
</paragraph>
</subSubSection>
<caption id="AE9F6668FF95240EFF7EBCCDFDB73BC3" ID-Table-UUID="AE9F6668FF95240EFF7EBCCDFDB73BC3" httpUri="http://table.plazi.org/id/AE9F6668FF95240EFF7EBCCDFDB73BC3" pageId="19" pageNumber="20" targetBox="[159,1333,774,1044]" targetIsTable="true" targetPageId="19">
<paragraph id="FA5F36E0FF95240EFF7EBCCDFDB73BC3" blockId="19.[151,1435,691,747]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF7EBCCDFEEA3BE3" bold="true" box="[151,269,692,715]" pageId="19" pageNumber="20">TABLE 5.</emphasis>
Shell measurements (in mm) of type material of
<taxonomicName id="3DE04D63FF95240EFCD6BCCAFBFA3BE3" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[831,1053,691,715]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="carinata" status="sp. nov.">
<emphasis id="C894EAF2FF95240EFCD6BCCAFBFA3BE3" box="[831,1053,691,715]" italics="true" pageId="19" pageNumber="20">Eulodrobia carinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF95240EFBCEBCCDFB803BE3" box="[1063,1127,692,715]" pageId="19" pageNumber="20" rank="species">
<emphasis id="C894EAF2FF95240EFBCEBCCDFB803BE3" bold="true" box="[1063,1127,692,715]" pageId="19" pageNumber="20">n. sp.</emphasis>
</taxonomicNameLabel>
(paratypes, C.400131). See Materials and Methods for abbreviations.
</paragraph>
</caption>
<paragraph id="FA5F36E0FF95240EFE3BBD7FFAD23D3C" pageId="19" pageNumber="20">
<table id="88E0C440FF95DBE2FF76BD7FFAD23D3C" box="[159,1333,774,1044]" gridcols="7" gridrows="7" pageId="19" pageNumber="20">
<tr id="44D034A2FF95DBE2FF76BD7FFAD23A34" box="[159,1333,774,796]" gridrow="0" pageId="19" pageNumber="20" rowspan-0="1">
<th id="07015DDEFF95DBE2FE3BBD7FFE193A34" box="[466,510,774,796]" gridcol="1" gridrow="0" pageId="19" pageNumber="20">SH</th>
<th id="07015DDEFF95DBE2FD86BD7FFD7C3A34" box="[623,667,774,796]" gridcol="2" gridrow="0" pageId="19" pageNumber="20">SW</th>
<th id="07015DDEFF95DBE2FCE0BD7FFCD23A34" box="[777,821,774,796]" gridcol="3" gridrow="0" pageId="19" pageNumber="20">AH</th>
<th id="07015DDEFF95DBE2FC55BD7FFC0F3A34" box="[956,1000,774,796]" gridcol="4" gridrow="0" pageId="19" pageNumber="20">AW</th>
<th id="07015DDEFF95DBE2FB86BD7FFB7C3A34" box="[1135,1179,774,796]" gridcol="5" gridrow="0" pageId="19" pageNumber="20">BW</th>
<th id="07015DDEFF95DBE2FAE0BD7FFAD23A34" box="[1289,1333,774,796]" gridcol="6" gridrow="0" pageId="19" pageNumber="20">TW</th>
</tr>
<tr id="44D034A2FF95DBE2FF76BD49FAD23A6E" box="[159,1333,816,838]" gridrow="1" pageId="19" pageNumber="20">
<th id="07015DDEFF95DBE2FF76BD49FF1A3A6E" box="[159,253,816,838]" gridcol="0" gridrow="1" pageId="19" pageNumber="20">Holotype</th>
<td id="07015DDEFF95DBE2FE3BBD49FE193A6E" box="[466,510,816,838]" gridcol="1" gridrow="1" pageId="19" pageNumber="20">1.30</td>
<td id="07015DDEFF95DBE2FD86BD49FD7C3A6E" box="[623,667,816,838]" gridcol="2" gridrow="1" pageId="19" pageNumber="20">2.39</td>
<td id="07015DDEFF95DBE2FCE0BD49FCD23A6E" box="[777,821,816,838]" gridcol="3" gridrow="1" pageId="19" pageNumber="20">0.97</td>
<td id="07015DDEFF95DBE2FC55BD49FC0F3A6E" box="[956,1000,816,838]" gridcol="4" gridrow="1" pageId="19" pageNumber="20">0.77</td>
<td id="07015DDEFF95DBE2FB86BD49FB7C3A6E" box="[1135,1179,816,838]" gridcol="5" gridrow="1" pageId="19" pageNumber="20">1.14</td>
<td id="07015DDEFF95DBE2FAE0BD49FAD23A6E" box="[1289,1333,816,838]" gridcol="6" gridrow="1" pageId="19" pageNumber="20">2.75</td>
</tr>
<tr id="44D034A2FF95DBE2FF76BD20FAD23A47" box="[159,1333,857,879]" gridrow="2" pageId="19" pageNumber="20">
<th id="07015DDEFF95DBE2FF76BD20FAD23A47" box="[159,1333,857,879]" colspan="7" colspanRight="6" gridcol="0" gridrow="2" pageId="19" pageNumber="20">Paratypes (20)</th>
</tr>
<tr id="44D034A2FF95DBE2FF76BDFBFAD23AB0" box="[159,1333,898,920]" gridrow="3" pageId="19" pageNumber="20">
<th id="07015DDEFF95DBE2FF76BDFBFF1A3AB0" box="[159,253,898,920]" gridcol="0" gridrow="3" pageId="19" pageNumber="20">Min</th>
<td id="07015DDEFF95DBE2FE3BBDFBFE193AB0" box="[466,510,898,920]" gridcol="1" gridrow="3" pageId="19" pageNumber="20">1.26</td>
<td id="07015DDEFF95DBE2FD86BDFBFD7C3AB0" box="[623,667,898,920]" gridcol="2" gridrow="3" pageId="19" pageNumber="20">2.3</td>
<td id="07015DDEFF95DBE2FCE0BDFBFCD23AB0" box="[777,821,898,920]" gridcol="3" gridrow="3" pageId="19" pageNumber="20">0.81</td>
<td id="07015DDEFF95DBE2FC55BDFBFC0F3AB0" box="[956,1000,898,920]" gridcol="4" gridrow="3" pageId="19" pageNumber="20">0.89</td>
<td id="07015DDEFF95DBE2FB86BDFBFB7C3AB0" box="[1135,1179,898,920]" gridcol="5" gridrow="3" pageId="19" pageNumber="20">1.19</td>
<td id="07015DDEFF95DBE2FAE0BDFBFAD23AB0" box="[1289,1333,898,920]" gridcol="6" gridrow="3" pageId="19" pageNumber="20">2.95</td>
</tr>
<tr id="44D034A2FF95DBE2FF76BDD5FAD23AEA" box="[159,1333,940,962]" gridrow="4" pageId="19" pageNumber="20">
<th id="07015DDEFF95DBE2FF76BDD5FF1A3AEA" box="[159,253,940,962]" gridcol="0" gridrow="4" pageId="19" pageNumber="20">Max</th>
<td id="07015DDEFF95DBE2FE3BBDD5FE193AEA" box="[466,510,940,962]" gridcol="1" gridrow="4" pageId="19" pageNumber="20">1.58</td>
<td id="07015DDEFF95DBE2FD86BDD5FD7C3AEA" box="[623,667,940,962]" gridcol="2" gridrow="4" pageId="19" pageNumber="20">2.88</td>
<td id="07015DDEFF95DBE2FCE0BDD5FCD23AEA" box="[777,821,940,962]" gridcol="3" gridrow="4" pageId="19" pageNumber="20">1.11</td>
<td id="07015DDEFF95DBE2FC55BDD5FC0F3AEA" box="[956,1000,940,962]" gridcol="4" gridrow="4" pageId="19" pageNumber="20">1.15</td>
<td id="07015DDEFF95DBE2FB86BDD5FB7C3AEA" box="[1135,1179,940,962]" gridcol="5" gridrow="4" pageId="19" pageNumber="20">1.48</td>
<td id="07015DDEFF95DBE2FAE0BDD5FAD23AEA" box="[1289,1333,940,962]" gridcol="6" gridrow="4" pageId="19" pageNumber="20">3.5</td>
</tr>
<tr id="44D034A2FF95DBE2FF76BDACFAD23AC3" box="[159,1333,981,1003]" gridrow="5" pageId="19" pageNumber="20">
<th id="07015DDEFF95DBE2FF76BDACFF1A3AC3" box="[159,253,981,1003]" gridcol="0" gridrow="5" pageId="19" pageNumber="20">Mean</th>
<td id="07015DDEFF95DBE2FE3BBDACFE193AC3" box="[466,510,981,1003]" gridcol="1" gridrow="5" pageId="19" pageNumber="20">1.46</td>
<td id="07015DDEFF95DBE2FD86BDACFD7C3AC3" box="[623,667,981,1003]" gridcol="2" gridrow="5" pageId="19" pageNumber="20">2.61</td>
<td id="07015DDEFF95DBE2FCE0BDACFCD23AC3" box="[777,821,981,1003]" gridcol="3" gridrow="5" pageId="19" pageNumber="20">1.02</td>
<td id="07015DDEFF95DBE2FC55BDACFC0F3AC3" box="[956,1000,981,1003]" gridcol="4" gridrow="5" pageId="19" pageNumber="20">1.05</td>
<td id="07015DDEFF95DBE2FB86BDACFB7C3AC3" box="[1135,1179,981,1003]" gridcol="5" gridrow="5" pageId="19" pageNumber="20">1.38</td>
<td id="07015DDEFF95DBE2FAE0BDACFAD23AC3" box="[1289,1333,981,1003]" gridcol="6" gridrow="5" pageId="19" pageNumber="20">3.30</td>
</tr>
<tr id="44D034A2FF95DBE2FF76BD87FAD23D3C" box="[159,1333,1022,1044]" gridrow="6" pageId="19" pageNumber="20">
<th id="07015DDEFF95DBE2FF76BD87FF1A3D3C" box="[159,253,1022,1044]" gridcol="0" gridrow="6" pageId="19" pageNumber="20">SD</th>
<td id="07015DDEFF95DBE2FE3BBD87FE193D3C" box="[466,510,1022,1044]" gridcol="1" gridrow="6" pageId="19" pageNumber="20">0.08</td>
<td id="07015DDEFF95DBE2FD86BD87FD7C3D3C" box="[623,667,1022,1044]" gridcol="2" gridrow="6" pageId="19" pageNumber="20">0.16</td>
<td id="07015DDEFF95DBE2FCE0BD87FCD23D3C" box="[777,821,1022,1044]" gridcol="3" gridrow="6" pageId="19" pageNumber="20">0.06</td>
<td id="07015DDEFF95DBE2FC55BD87FC0F3D3C" box="[956,1000,1022,1044]" gridcol="4" gridrow="6" pageId="19" pageNumber="20">0.06</td>
<td id="07015DDEFF95DBE2FB86BD87FB7C3D3C" box="[1135,1179,1022,1044]" gridcol="5" gridrow="6" pageId="19" pageNumber="20">0.08</td>
<td id="07015DDEFF95DBE2FAE0BD87FAD23D3C" box="[1289,1333,1022,1044]" gridcol="6" gridrow="6" pageId="19" pageNumber="20">0.14</td>
</tr>
</table>
</paragraph>
<subSubSection id="B2FA656BFF95240EFF2EBA2CFB133E82" pageId="19" pageNumber="20" type="description">
<paragraph id="FA5F36E0FF95240EFF2EBA2CFCB93DB9" blockId="19.[151,1437,1109,2034]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EBA2CFEA83D46" bold="true" box="[199,335,1109,1134]" pageId="19" pageNumber="20">Operculum</emphasis>
(
<figureCitation id="62DB2A65FF95240EFEB7BA2CFE233D46" box="[350,452,1109,1134]" captionStart="FIGURE 4" captionStartId="12.[151,250,1521,1543]" captionTargetBox="[196,1406,256,1477]" captionTargetId="figure@12.[151,1435,229,1500]" captionTargetPageId="12" captionText="FIGURE 4. Opercula of species of Eulodrobia n. gen., external and internal views of each species. A, B. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). C, D. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). E, F. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). G, H. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). I, J. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). K, L. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 500 µm." httpUri="https://zenodo.org/record/2637611/files/figure.png" pageId="19" pageNumber="20">
Fig.
<date id="8E5E1020FF95240EFE7BBA2FFE4B3D46" box="[402,428,1110,1134]" pageId="19" pageNumber="20">4I</date>
, J
</figureCitation>
). Translucent, brown, slightly convex, nucleus near central. Inner side with white streak or smear present, with slightly raised whitish pimple near apex.
</paragraph>
<paragraph id="FA5F36E0FF95240EFF2EBAE4FABA3DF2" blockId="19.[151,1437,1109,2034]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EBAE4FDCE3D9E" bold="true" box="[199,553,1181,1206]" pageId="19" pageNumber="20">Head-foot and external body.</emphasis>
Snout, tentacles, neck and opercular lobes pigmented, dorsal and lateral foot unpigmented, mantle roof weakly pigmented to densely pigmented, visceral coil densely pigmented to black.
</paragraph>
<paragraph id="FA5F36E0FF95240EFF2EBA9CFEE83C9A" blockId="19.[151,1437,1109,2034]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EBA9CFE913DD6" bold="true" box="[199,374,1253,1278]" pageId="19" pageNumber="20">Mantle cavity.</emphasis>
Ctenidium well-developed, filaments 19, broadly triangular, apex right edge. Osphradium narrowly oval, towards posterior end of ctenidium or at middle of ctenidium, length relative to gill 0.2–0.31. Hypobranchial gland moderately developed. Rectum with U-shaped bend, anus behind mantle collar. Kidney extends for about quarter of length into mantle cavity roof. Renal gland transverse. Pericardium extends for about quarter of length into mantle cavity roof, overlapping posterior end of ctenidium or abutting posterior end of ctenidium.
</paragraph>
<paragraph id="FA5F36E0FF95240EFF2EBBC4FE843F69" blockId="19.[151,1437,1109,2034]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EBBC4FEF93CFE" bold="true" box="[199,286,1469,1494]" pageId="19" pageNumber="20">Radula</emphasis>
(
<figureCitation id="62DB2A65FF95240EFEDDBBC4FE763CFE" box="[308,401,1469,1494]" captionStart="FIGURE 5" captionStartId="13.[151,250,1812,1834]" captionTargetBox="[269,1322,207,1777]" captionTargetId="figure@13.[269,1322,193,1794]" captionTargetPageId="13" captionText="FIGURE 5. Radulae of Eulodrobia n. gen. A, B. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). A, half rows, B, detail of lateral tooth. C–E. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779), C, half rows, D detail of lateral tooth, E, detail of central tooth. F–H. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). F. Detail of central tooth. G. Detail of inner marginal tooth. H. Detail of outer lateral teeth. I–K. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). I. Part of tooth row. J. Detail of lateral teeth. K. Detail of inner marginal tooth. L.Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131), half rows. M. N. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133), half rows. Scale bars 10 µm." httpUri="https://zenodo.org/record/2637613/files/figure.png" pageId="19" pageNumber="20">Fig. 5L</figureCitation>
). Central teeth with cusp formula 2+1+2, basal cusps 2+2; median cusp spathulate to triangular, about twice to three times as long as adjacent cusps. Lateral teeth with cusp formula 3–4+1+1(–2), main cusp triangular, about twice length of adjacent cusps. Inner marginal teeth with 25–27 cusps. Outer marginal teeth with 22–23 cusps.
</paragraph>
<paragraph id="FA5F36E0FF95240EFF2EB834FDEB3E16" blockId="19.[151,1437,1109,2034]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EB834FDFD3F4E" bold="true" box="[199,538,1613,1638]" pageId="19" pageNumber="20">Female reproductive system</emphasis>
(
<figureCitation id="62DB2A65FF95240EFDC5B834FD433F4E" box="[556,676,1613,1638]" captionStart="FIGURE 6" captionStartId="15.[151,250,1512,1534]" captionTargetBox="[181,1411,211,1442]" captionTargetId="figure@15.[150,1436,193,1490]" captionTargetPageId="15" captionText="FIGURE 6. Female reproductive system of species of Eulodrobia n. gen., shown from left side. A, B. Eulodrobia fenshami n. sp. (C. 410721). C, F. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). D, E. Eulodrobia ovata n. sp. (C. 400130). G, H. Eulodrobia bundoona n. sp. (C.400132). I, L. Eulodrobia spirula n. sp. (C.400133). J, K. Eulodrobia carinata n. sp. (C.400131). Abbreviations: ag—albumen gland; bc—bursa copulatrix; cg—capsule gland; ga—opening of oviduct; pwposterior wall of mantle cavity; ro—renal oviduct; sr—seminal receptacle; v—vestibule; vc—ventral channel. Scale bars 500 µm." httpUri="https://zenodo.org/record/2637615/files/figure.png" pageId="19" pageNumber="20">Fig. 6J, K</figureCitation>
). Ovary weakly lobed; renal oviduct lacking initial U-shaped arch but weak bend may be present; forms near circular loop over bursa copulatrix. Seminal receptacle absent; orientated sperm in renal oviduct. Bursa copulatrix overlapping albumen gland on left, pyriform, shorter than albumen gland, bursal duct enters bursa mid anteriorly, bursal duct joins coiled oviduct little behind posterior mantle cavity wall. Albumen gland behind posterior mantle cavity wall. Capsule gland with two distinct glandular zones, medium thickness in cross section, markedly indented by rectum. Anterior vestibule small, opening subterminal, intermediate in size.
</paragraph>
<paragraph id="FA5F36E0FF95240EFF2EB930FB133E82" blockId="19.[151,1437,1109,2034]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EB930FE1A3E4A" bold="true" box="[199,509,1865,1890]" pageId="19" pageNumber="20">Male reproductive system</emphasis>
(
<figureCitation id="62DB2A65FF95240EFDE5B930FD8A3E49" box="[524,621,1865,1890]" captionStart="FIGURE 7" captionStartId="16.[151,250,1324,1346]" captionTargetBox="[170,1416,747,1275]" captionTargetId="figure@16.[150,1436,735,1299]" captionTargetPageId="16" captionText="FIGURE 7. Prostate and vas deferens of species of Eulodrobia n. gen. Each species is shown with anterior at left and posterior at right, with line indicating position of posterior pallial wall. A. Eulodrobia fenshami n. sp. (C. 410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). C. Eulodrobia ovata n. sp. (C.400130). D. Eulodrobia bundoona n. sp. (C.400132). E, Eulodrobia carinata n. sp. (C. 400131). F. Eulodrobia spirula n. sp. (C.400133). Abbreviations: avd—anterior vas deferens; pw—position of posterior wall of pallial cavity; ve—posterior vas deferens. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637617/files/figure.png" pageId="19" pageNumber="20">Figs. 7E</figureCitation>
,
<figureCitation id="62DB2A65FF95240EFD90B933FD7B3E4A" box="[633,668,1866,1890]" captionStart="FIGURE 8" captionStartId="17.[151,250,1042,1064]" captionTargetBox="[170,1435,206,1008]" captionTargetId="figure@17.[151,1435,195,1020]" captionTargetPageId="17" captionText="FIGURE 8. Penis and vas deferens of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp. (C. 410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). C. Eulodrobia ovata n. sp. (C. 400130). D. Eulodrobia bundoona n. sp. (C.400132). E, Eulodrobia carinata n. sp. (C. 400131). F. Eulodrobia spirula n. sp. (C.400133). Abbreviations: ngl—non-glandular lobe; pd—penial duct; vd—vas deferens. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637619/files/figure.png" pageId="19" pageNumber="20">8E</figureCitation>
). Prostate gland less than half in mantle roof, oval, medium in cross section. Posterior pallial vas deferens straight, anteriorly slightly undulating. Penis towards middle of head, well down neck, distal end tapering, terminal papilla small, with small near-terminal non-glandular lobe.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF95240EFF2EB9CCFD913EE6" box="[199,630,1973,1998]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="FA5F36E0FF95240EFF2EB9CCFD913EE6" blockId="19.[151,1437,1109,2034]" box="[199,630,1973,1998]" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EB9CCFEA93EE6" bold="true" box="[199,334,1973,1998]" pageId="19" pageNumber="20">Etymology.</emphasis>
Carinatus—Latin, keeled.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF952409FF2EB9A0FDEE39FD" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="distribution">
<paragraph id="FA5F36E0FF952409FF2EB9A0FDEE39FD" blockId="19.[151,1437,1109,2034]" lastBlockId="20.[151,1436,151,357]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="C894EAF2FF95240EFF2EB9A0FE153EDA" bold="true" box="[199,498,2009,2034]" pageId="19" pageNumber="20">Distribution and habitat.</emphasis>
Found only in the main spring, Yowah Creek Springs, on Bundoona Station, about
<quantity id="3D189B05FF922409FF7EBEEEFF073998" box="[151,224,151,176]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="20" pageNumber="21" unit="km" value="36.0">36 km</quantity>
NW of Eulo (
<figureCitation id="62DB2A65FF922409FE6EBEEEFE2D3998" box="[391,458,151,176]" captionStart="FIGURE 1" captionStartId="4.[151,250,1844,1866]" captionTargetBox="[151,1435,498,1802]" captionTargetId="figure@4.[151,1435,498,1802]" captionTargetPageId="4" captionText="FIGURE 1. Map of the spring supergroups associated with the Great Artesian Basin (GAB) in Queensland, showing the main non-town locations mentioned in the text (squares) and some of the towns in the area (ovals). The pale grey shading is the discharge part of the GAB, the dark grey the recharge part. The inset map of Australia shows the area illustrated. The approximate location of Conondale is indicated by a triangle and Nundle by a star. The spring supergroups are outlined with dotted lines and are as follows: A, Mulligan River; B, Springvale; C, Flinders River; D, Hughenden; E, Barcaldine (see inset at upper right for details); F, Springsure; G, Eulo." httpUri="https://zenodo.org/record/2637605/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
), where it lives in the very shallow spring outflow. This spring has well oxygenated, clean water and coarse sediment.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF922409FF2EBEA6FCA7384D" pageId="20" pageNumber="21" type="discussion">
<paragraph id="FA5F36E0FF922409FF2EBEA6FCA7384D" blockId="20.[151,1436,151,357]" pageId="20" pageNumber="21">
<emphasis id="C894EAF2FF922409FF2EBEA6FEDC39D0" bold="true" box="[199,315,223,248]" pageId="20" pageNumber="21">Remarks.</emphasis>
This distinctive species lives in the shallow outflow of the main spring together with the species described immediately below. It differs from all other species in the GAB (and in
<collectingCountry id="82F77670FF922409FBC1BF7DFB763835" box="[1064,1169,260,285]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
) in having a prominent, flange-like keel on the periphery. Compared to the other species from the Eulo Supergroup, it is most similar to the species described below and is contrasted under that species.
</paragraph>
</subSubSection>
</treatment>
</document>
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