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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.901.39432" ID-GBIF-Dataset="654566e8-947b-4dfe-8fbc-baea3d254fa8" ID-PMC="PMC6965333" ID-Pensoft-Pub="1313-2970-901-1" ID-Pensoft-UUID="D8C38C1C3844549F8BEC7C1D884CDF8E" ID-PubMed="31975782" ModsDocID="1313-2970-901-1" checkinTime="1578615027095" checkinUser="pensoft" docAuthor="Balkenohl, Michael" docDate="2020" docId="5DEE114B7FB4546593D67C204D8440B6" docLanguage="en" docName="ZooKeys 901: 1-81" docOrigin="ZooKeys 901" docSource="http://dx.doi.org/10.3897/zookeys.901.39432" docTitle="Salcedia schoutedeni Alluaud 1930" docType="treatment" docVersion="4" id="D8C38C1C3844549F8BEC7C1D884CDF8E" lastPageNumber="1" masterDocId="D8C38C1C3844549F8BEC7C1D884CDF8E" masterDocTitle="A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="1" updateTime="1668168267548" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balkenohl, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>901</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>81</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.901.39432</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.901.39432</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-901-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">D8C38C1C3844549F8BEC7C1D884CDF8E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="161671757" LSID="urn:lsid:plazi:treatment:5DEE114B7FB4546593D67C204D8440B6" httpUri="http://treatment.plazi.org/id/5DEE114B7FB4546593D67C204D8440B6" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="5DEE114B-7FB4-5465-93D6-7C204D8440B6" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia schoutedeni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="schoutedeni">Salcedia schoutedeni Alluaud, 1930</taxonomicName>
<figureCitation captionStart="Figures 14–19" captionStartId="F8" captionText="Figures 14 - 19. Salcedia species, habitus, dorsal view, holotype 14 S. schoutedeni Alluaud 15 S. nigeriensis Alluaud (cotype) 16 S. putzeysi (Oberthuer) 17 S. matsumotoi sp. nov. 18 S. procera sp. nov. 19 S. baroensis sp. nov." figureDoi="10.3897/zookeys.901.39432.figures14-19" httpUri="https://binary.pensoft.net/fig/370609" pageId="0" pageNumber="1">Figs 14</figureCitation>
<figureCitation captionStart="Figures 26–35" captionStartId="F10" captionText="Figures 26 - 35. Salcedia species, pronotum, dorsal view 26 S. perrieri Fairmaire 27 S. unifoveata sp. nov. 28 S. faillei sp. nov. 29 S. coquilhati Alluaud 30 S. elongata Alluaud 31 S. africana (Britton) 32 S. schoutedeni Alluaud 33 S. nigeriensis Alluaud 34 S. putzeysi (Oberthuer) 35 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures26-35" httpUri="https://binary.pensoft.net/fig/370611" pageId="0" pageNumber="1">, 32</figureCitation>
<figureCitation captionStart="Figures 44–51" captionStartId="F12" captionText="Figures 44 - 51. Salcedia species, left elytron, basal half, dorsal view 44 S. perrieri Fairmaire 45 S. unifoveata sp. nov. 46 S. elongata Alluaud 47 S. africana (Britton) 48 S. schoutedeni Alluaud 49 S. nigeriensis Alluaud 50 S. baroensis sp. nov. 51 S. miranda (Andrewes)." figureDoi="10.3897/zookeys.901.39432.figures44-51" httpUri="https://binary.pensoft.net/fig/370613" pageId="0" pageNumber="1">, 48</figureCitation>
<figureCitation captionStart="Figures 58–63" captionStartId="F14" captionText="Figures 58 - 63. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 58 S. schoutedeni Alluaud 59 S. nigeriensis Alluaud 60 S. putzeysi (Oberthuer) 61 S. matsumotoi sp. nov. 62 S. procera sp. nov. 63 S. baroensis sp. nov." figureDoi="10.3897/zookeys.901.39432.figures58-63" httpUri="https://binary.pensoft.net/fig/370615" pageId="0" pageNumber="1">, 58</figureCitation>
<figureCitation captionStart="Figures 68–76" captionStartId="F16" captionText="Figures 68 - 76. Salcedia species, female coxostyli (gonopod IX) and laterotergite IX (epipleutite IX) 68 S. perrieri Fairmaire 69 S. unifoveata sp. nov. 70 S. coquilhati Alluaud 71 S. elongata Alluaud 72 S. africana (Britton) 73 S. schoutedeni Alluaud 74 S. nigeriensis Alluaud 75 S. putzeysi (Oberthuer) 76 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures68-76" httpUri="https://binary.pensoft.net/fig/370617" pageId="0" pageNumber="1">, 73</figureCitation>
<figureCitation captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">, 84</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Alluaud" authorityYear="1930" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia schoutedeni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="schoutedeni">Salcedia schoutedeni</taxonomicName>
Alluaud, 1930: 21;
<bibRefCitation author="Alluaud, C" journalOrPublisher="Afra" pageId="0" pageNumber="1" pagination="1 - 6" refId="B3" refString="Alluaud, C, 1932. Carabiques d'Afrique. 1. Nouvelle etude du genre Salcedia. Afra 4: 1 - 6" title="Carabiques d'Afrique. 1. Nouvelle etude du genre Salcedia." volume="4" year="1932">Alluaud 1932</bibRefCitation>
: 2;
<bibRefCitation author="Csiki, E" journalOrPublisher="Entomologist's monthly magazine, May" pageId="0" pageNumber="1" refId="B21" refString="Csiki, E, 1933. Coleopterorum Catalogus auspiciis et auxilio. W. Junk editus a S. Schenkling. 127: Carabidae: Carabinae III, Corrigenda et Addenda. W. Junk, Berlin, 623-641." title="Coleopterorum Catalogus auspiciis et auxilio. W. Junk editus a S. Schenkling. 127: Carabidae: Carabinae III, Corrigenda et Addenda. W. Junk, Berlin, 623 - 641." year="1933">Csiki 1933</bibRefCitation>
: 641;
<bibRefCitation author="Alluaud, C" journalOrPublisher="Afra" pageId="0" pageNumber="1" pagination="19 - 20" refId="B4" refString="Alluaud, C, 1935. Note diverses. Localites nouvelles. Afra 9: 19 - 20" title="Note diverses. Localites nouvelles." volume="9" year="1935">Alluaud 1935</bibRefCitation>
: 19;
<bibRefCitation author="Burgeon, L" journalOrPublisher="Entomologist's monthly magazine, May" pageId="0" pageNumber="1" refId="B19" refString="Burgeon, L, 1935. Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres - Carabides. Annales du Musee du Congo Belge, Zoologie, Serie III. Section II. Tome II: 135-257." title="Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres - Carabides. Annales du Musee du Congo Belge, Zoologie, Serie III. Section II. Tome II: 135 - 257." year="1935">Burgeon 1935</bibRefCitation>
: 159;
<bibRefCitation DOI="https://doi.org/10.1080/00222933608655174" author="Andrewes, HE" journalOrPublisher="The Annals and magazine of Natural History" pageId="0" pageNumber="1" pagination="54 - 65" refId="B8" refString="Andrewes, HE, 1936. Papers on Oriental Carabidae, 30. The Annals and magazine of Natural History 18 (10): 54 - 65, DOI: https://doi.org/10.1080/00222933608655174" title="Papers on Oriental Carabidae, 30." url="https://doi.org/10.1080/00222933608655174" volume="18" year="1936">Andrewes 1936</bibRefCitation>
: 64.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="schoutedeni" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia schoutedeni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="schoutedeni">Salcedia schoutedeni schoutedeni</taxonomicName>
Alluaud, 1930:
<bibRefCitation author="Reichardt, H" journalOrPublisher="Revista Brasileira Entomologia" pageId="0" pageNumber="1" pagination="99 - 110" refId="B38" refString="Reichardt, H, 1975. The genera of Salcediina (Coleoptera, Carabidae, Scaritini). Revista Brasileira Entomologia 19: 99 - 110" title="The genera of Salcediina (Coleoptera, Carabidae, Scaritini)." volume="19" year="1975">Reichardt 1975</bibRefCitation>
: 103;
<bibRefCitation author="Dostal, A" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="1" pagination="117 - 121" refId="B24" refString="Dostal, A, 1993. Beschreibung einer neuen Gattung aus Vietnam: Androzelma gen. nov. (Coleoptera, Carabidae, Scaritinae: Salcediini). Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 45: 117 - 121" title="Beschreibung einer neuen Gattung aus Vietnam: Androzelma gen. nov. (Coleoptera, Carabidae, Scaritinae: Salcediini)." volume="45" year="1993">Dostal 1993</bibRefCitation>
: 121;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Printed by the author, Tutzing" pageId="0" pageNumber="1" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant Ground Beetles of the World (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae), Second Edition. Printed by the author, Tutzing" title="Systematic list of extant Ground Beetles of the World (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae), Second Edition." year="2005">Lorenz 2005</bibRefCitation>
: 155.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♀, with labels and data: white, printed and handwritten
<normalizedToken originalValue="“MUSÉE">&quot;MUSEE</normalizedToken>
DU CONGO Kwamouth 9-VII-1921 Dr H.Schouteden&quot; / white, printed black framed &quot;R.
<normalizedToken originalValue="DÉT">DET</normalizedToken>
. 1783 c&quot; red, black double framed
<normalizedToken originalValue="“HOLOTYPE”">&quot;HOLOTYPE&quot;</normalizedToken>
/ white, black framed, handwritten and printed &quot;
<emphasis italics="true" pageId="0" pageNumber="1">Salcedia Schoutedeni</emphasis>
Type Alluaud, det. 1930&quot; (MRACT).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remark.</paragraph>
<paragraph pageId="0" pageNumber="1">In the holotype both of the front legs are broken and glued.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
1 ♂, 1 ♀, 22 specs.,
<normalizedToken originalValue="Léopoldville">Leopoldville</normalizedToken>
I-1947 Dr E. Dartevelle COLL. MUS. CONGO (MRACT, CBB); 1 ♀, same data (MNHN); 1 spec.,
<normalizedToken originalValue="Léopoldville">Leopoldville</normalizedToken>
-1930 Eg. Devroye
<normalizedToken originalValue="MUSÉE">MUSEE</normalizedToken>
DU CONGO / R.
<normalizedToken originalValue="DÉT">DET</normalizedToken>
. 2019 b /
<taxonomicName class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia schoutedeni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis italics="true" pageId="0" pageNumber="1">Salcedia schoutedeni</emphasis>
</taxonomicName>
All. Alluaud. det. 1931 (MRACT); 1 spec.,
<normalizedToken originalValue="Léopoldville">Leopoldville</normalizedToken>
-1930 E. Devroey
<normalizedToken originalValue="MUSÉE">MUSEE</normalizedToken>
DU CONGO (MRACT); 1 spec., same data but -1931 (MRACT).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A large sized species, with sub-elongate outline of the elytra with maximum width at middle and the pronotum with a rudiment of the outer lateral carina. The pseudohumerus is rectangular with a distinctly laterally projecting tooth. The antennomeres are elongate. Distinguished most clearly from the similar species
<taxonomicName lsidName="S. nigeriensis" pageId="0" pageNumber="1" rank="species" species="nigeriensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. nigeriensis</emphasis>
</taxonomicName>
by the pronotum with the lateral margin converging from middle to the anterior angles, the pseudohumerus of the elytron with a distinct tooth, and the female coxostylus with six large setae and one SSO. In addition,
<taxonomicName lsidName="S. schoutedeni" pageId="0" pageNumber="1" rank="species" species="schoutedeni">
<emphasis italics="true" pageId="0" pageNumber="1">S. schoutedeni</emphasis>
</taxonomicName>
has a distinct keel on the frons of the head.
</paragraph>
<caption doi="10.3897/zookeys.901.39432.figures14-19" httpUri="https://binary.pensoft.net/fig/370609" pageId="0" pageNumber="1" start="Figures 14–19" startId="F8">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 14-19.</emphasis>
<taxonomicName authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Salcedia</emphasis>
</taxonomicName>
species, habitus, dorsal view, holotype
<emphasis bold="true" pageId="0" pageNumber="1">14</emphasis>
<taxonomicName lsidName="S. schoutedeni" pageId="0" pageNumber="1" rank="species" species="schoutedeni">
<emphasis italics="true" pageId="0" pageNumber="1">S. schoutedeni</emphasis>
</taxonomicName>
Alluaud
<emphasis bold="true" pageId="0" pageNumber="1">15</emphasis>
<taxonomicName lsidName="S. nigeriensis" pageId="0" pageNumber="1" rank="species" species="nigeriensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. nigeriensis</emphasis>
</taxonomicName>
Alluaud (cotype)
<emphasis bold="true" pageId="0" pageNumber="1">16</emphasis>
<taxonomicName lsidName="S. putzeysi" pageId="0" pageNumber="1" rank="species" species="putzeysi">
<emphasis italics="true" pageId="0" pageNumber="1">S. putzeysi</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
)
<emphasis bold="true" pageId="0" pageNumber="1">17</emphasis>
<taxonomicName lsidName="S. matsumotoi" pageId="0" pageNumber="1" rank="species" species="matsumotoi">
<emphasis italics="true" pageId="0" pageNumber="1">S. matsumotoi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">18</emphasis>
<taxonomicName lsidName="S. procera" pageId="0" pageNumber="1" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="1">S. procera</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">19</emphasis>
<taxonomicName lsidName="S. baroensis" pageId="0" pageNumber="1" rank="species" species="baroensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. baroensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="redescription">
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
Measurements in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and ratios of Salcedia species. N = number of specimens measured; x ̄ = arithmetic mean; &quot; term used &quot; refers to the shape of the elytra." httpUri="http://table.plazi.org/id/FDDA58A1B347C9F0593A738CF707345F" pageId="0" pageNumber="1" tableUuid="FDDA58A1B347C9F0593A738CF707345F">1</tableCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour and surface:</emphasis>
Anthracite grey, shiny; top of carinae on head, pronotum and elytra as well as margins of pronotum and elytra opaque, covered with pale grey pili; legs piceous, mandibles, antennae and palpi piceous.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head:</emphasis>
Less than two-thirds of width of pronotum. Outline semi-circular. Clypeus wide, convex anteriorly, fused with clypeal wings, separated from supra-antennal plates by distinct notches, with slightly raised pentagon-shaped field at middle, separated from frons by deep transverse furrow. Frons with two raised paramedian carinae, joining anteriorly into a V-like keel, with two small glossy teeth bilaterally anterior to central keel, with two indistinct short parallel carinae paralaterally at base; frons separated from supraorbital plates by deep broad furrows; with conspicuously deep and broad pit at front-eye level; supra-antennal and supraorbital plates margined, margin of supraorbital plate distinctly raised, carina-like, supra-antennal plates vaulted. Base with narrow emargination at middle, sharply rectangular laterally (angle around 109°). Eyes large, convex, genae slightly convex, both of them clearly visible from above, with nearly circular shape in lateral view. Antenna with segments five to ten elongate (L/W 1.25), densely pubescent, segments two to four scarcely pubescent, scapus with irregular reticulation. Labrum nearly straight anteriorly. Mandible moderately short, wide, slightly arcuate at apex. Mentum small, with slight tooth at middle, epilobes wide, projecting and angled anteriorly, margined anteriorly, surface with small pits anteriorly, with isodiametric reticulation basally.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
(Fig.
<figureCitation captionStart="Figures 26–35" captionStartId="F10" captionText="Figures 26 - 35. Salcedia species, pronotum, dorsal view 26 S. perrieri Fairmaire 27 S. unifoveata sp. nov. 28 S. faillei sp. nov. 29 S. coquilhati Alluaud 30 S. elongata Alluaud 31 S. africana (Britton) 32 S. schoutedeni Alluaud 33 S. nigeriensis Alluaud 34 S. putzeysi (Oberthuer) 35 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures26-35" httpUri="https://binary.pensoft.net/fig/370611" pageId="0" pageNumber="1">32</figureCitation>
): Outline rectangular, transverse, two-fifth wider than long. Lateral margin convex, distinctly converging anteriorly, maximum width behind middle. Lateral margin crenulated, with 12-13 small tubercles, tubercles larger basally, with two notches at posterior angles. Base straight laterally, with declining flat keel at middle pointing posteriorly. Disc flattened in lateral view, with two raised paramedian carinae parallel to median line and diverging posteriorly, with median line small, long, and ending in pits anteriorly and posteriorly, with two additional shorter carinae bilaterally, the paralateral one joining with the paramedian carinae at base and forming tooth-like tubercle pointing posteriorly, without anterior extension, without inner lateral carina, with very small outer lateral carinae. All carinae sub-crenulate. Lateral margin broadly wing-like bent up, with six large and deep transverse pits, the basal and/or frontal ones partly separated into two smaller pits. Space between carinae and pits smooth.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Elytron</emphasis>
(Fig.
<figureCitation captionStart="Figures 44–51" captionStartId="F12" captionText="Figures 44 - 51. Salcedia species, left elytron, basal half, dorsal view 44 S. perrieri Fairmaire 45 S. unifoveata sp. nov. 46 S. elongata Alluaud 47 S. africana (Britton) 48 S. schoutedeni Alluaud 49 S. nigeriensis Alluaud 50 S. baroensis sp. nov. 51 S. miranda (Andrewes)." figureDoi="10.3897/zookeys.901.39432.figures44-51" httpUri="https://binary.pensoft.net/fig/370613" pageId="0" pageNumber="1">48</figureCitation>
): Flattened in anterior half (lateral view), convex in frontal view. Sub-elongate, margin straight in anterior half up to pseudohumerus, moderately convex to apex, maximum width at middle, sub-undulate. Pseudohumerus rectangular, tooth distinctly projecting laterally. Apex rounded, with small but acute tooth at suture. Disc with interneur six sub-crenulate, interneur three slightly carinate in basal third, interneur two running up to apex as slightly convex line, conspicuously raised; interneur four running in parallel to interneur six, almost reaching base, shortened at apex. Interneur five and six with two rows of serial pits partly merging transversally.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hind wings:</emphasis>
Fully developed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lower surface:</emphasis>
Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with one row of slightly transverse pits, lateral margin of elytron smooth. Metepisternum elongate, with broad longitudinal groove. Metasternum, abdominal sternites one and five with numerous irregularly situated larger and smaller pits, sternite two to three smooth, with band of small punctures at middle. Last three abdominal sternites laterally with isodiametric reticulation, sternite one with longitudinal reticulation, two with transverse reticulation. Sternum four to six slightly sulcate. Sternum six at apical margin with hollowed out transverse impression at apex.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs:</emphasis>
Profemora with surface indistinctly reticulated. Protibia with short, robust, moderately curved terminal spine, laterally with laterally with two larger and two smaller teeth, dorsally and ventrally with two carinae. First tarsomere distinctly elongated, as long as tarsomeres two to five together, front and intermediate legs with tarsomeres two to four broadened and somewhat flattened.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">External sexual dimorphism:</emphasis>
Sternum six with the hollowed out transverse impression at the apical margin is obtusely longitudinally sulcate in females.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 58–63" captionStartId="F14" captionText="Figures 58 - 63. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 58 S. schoutedeni Alluaud 59 S. nigeriensis Alluaud 60 S. putzeysi (Oberthuer) 61 S. matsumotoi sp. nov. 62 S. procera sp. nov. 63 S. baroensis sp. nov." figureDoi="10.3897/zookeys.901.39432.figures58-63" httpUri="https://binary.pensoft.net/fig/370615" pageId="0" pageNumber="1">58</figureCitation>
): Median lobe stout, in dorsal view distinctly and regularly arcuate, straight to apex, in lateral view straight, apparently no fine pili visible, apical part bent ventrally apex dorsally, in cross section oval at apex. Oroficium small. Endophallus with group of longer microtrichia near oroficium, with additional longer group of brush-like microtrichia basally. Dorsal paramere relatively short, bisinuate, with short apophyses; ventral one shaped like a convex spatula, with fine seta at apex; both parameres slightly distorted.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 68–76" captionStartId="F16" captionText="Figures 68 - 76. Salcedia species, female coxostyli (gonopod IX) and laterotergite IX (epipleutite IX) 68 S. perrieri Fairmaire 69 S. unifoveata sp. nov. 70 S. coquilhati Alluaud 71 S. elongata Alluaud 72 S. africana (Britton) 73 S. schoutedeni Alluaud 74 S. nigeriensis Alluaud 75 S. putzeysi (Oberthuer) 76 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures68-76" httpUri="https://binary.pensoft.net/fig/370617" pageId="0" pageNumber="1">73</figureCitation>
): Coxostylus regularly broadened to base, distinctly curved, acute at apex, with indistinct carina dorsally in apical third, at end of basal third with one strong and five slender nematiform setae laterally, SSO with one fine pilus.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Variation:</emphasis>
The tubercles at the lateral margin of the pronotum are more or less strongly developed. Number and shape of the pits on the ventral surface varies to a small degree.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">84</figureCitation>
). Known from several localities in the west of the Democratic Republic of the Congo along the lower course of the river Congo. The locality Kwamouth refers to the short estuary of the river Kasai into the River Congo. The location
<normalizedToken originalValue="“Zambèze">&quot;Zambeze</normalizedToken>
, Nova Chupanga
<normalizedToken originalValue="près">pres</normalizedToken>
Chemba&quot; quoted in
<bibRefCitation author="Alluaud, C" journalOrPublisher="Afra" pageId="0" pageNumber="1" pagination="19 - 20" refId="B4" refString="Alluaud, C, 1935. Note diverses. Localites nouvelles. Afra 9: 19 - 20" title="Note diverses. Localites nouvelles." volume="9" year="1935">Alluaud (1935)</bibRefCitation>
as new record for
<taxonomicName lsidName="S. schoutedeni" pageId="0" pageNumber="1" rank="species" species="schoutedeni">
<emphasis italics="true" pageId="0" pageNumber="1">S. schoutedeni</emphasis>
</taxonomicName>
refers to
<taxonomicName baseAuthorityName="Britton" baseAuthorityYear="1947" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia africana" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="africana">
<emphasis italics="true" pageId="0" pageNumber="1">Salcedia africana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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