148 lines
17 KiB
XML
148 lines
17 KiB
XML
<document id="FF476983B3AB1F789D6A1E1AFA72F128" ID-CLB-Dataset="181704" ID-DOI="10.11646/zootaxa.5203.1.1" ID-GBIF-Dataset="b6f94c7b-7033-4952-96a7-3db5fb6ad879" ID-ISSN="1175-5326" ID-Zenodo-Dep="7284841" ID-ZooBank="85C20298-5DF9-44DF-8485-3C64CF40CD08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667554441660" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2022" docId="0C081779B91F7847C5B84451FB28263B" docLanguage="en" docName="zootaxa.5203.1.1.pdf" docOrigin="Zootaxa 5203 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5203.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena camerocomplexa Perkins 2022, new species" docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="F0316F01B913784AC52F422FFFE92502" masterDocTitle="Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="13" updateTime="1699409684053" updateUser="ExternalLinkService">
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<mods:titleInfo id="DB71C4CE4773F5F3180FD5DE0D26575C">
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<mods:title id="027B3846318C164DF837CD99AD76B029">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<mods:name id="56B35540BF4C3560C90017266B9D8DAA" type="personal">
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<mods:namePart id="385B2F115313CAEA8B114F833341F6B1">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="DA193E2A7D91D6875ED595C6ED56AC33">text</mods:typeOfResource>
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<mods:title id="14B342F19D5DD84BEDB095EDD12991F5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8824950ED0306AFE9B97626F07D0D009">
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<mods:date id="59BCFFCFD1F240EDB2B0BCDF2CEBF303">2022</mods:date>
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<mods:number id="2692138A273DFA0F4696F81C6F67459E">2022-11-03</mods:number>
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<mods:number id="FF29B61C56887E8384032520B4487327">5203</mods:number>
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<mods:url id="0AD1D2B2EBD5A07FB0C8FCB0609A145C">http://dx.doi.org/10.11646/zootaxa.5203.1.1</mods:url>
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<mods:classification id="EC069A1FAE510C894C1C5184FE2FFF5F">journal article</mods:classification>
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<mods:identifier id="6A6EF64F6AC87C0CC49169A7B8F89180" type="CLB-Dataset">181704</mods:identifier>
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<mods:identifier id="197121F6C1768B57E8ABDE4FE82DE558" type="DOI">10.11646/zootaxa.5203.1.1</mods:identifier>
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<treatment id="0C081779B91F7847C5B84451FB28263B" ID-DOI="http://doi.org/10.5281/zenodo.7300255" ID-GBIF-Taxon="204205205" ID-Zenodo-Dep="7300255" LSID="urn:lsid:plazi:treatment:0C081779B91F7847C5B84451FB28263B" httpUri="http://treatment.plazi.org/id/0C081779B91F7847C5B84451FB28263B" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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<subSubSection id="CCBBF5E4B91F7846C5B84451FD6A239A" box="[151,643,1662,1689]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="841EA66FB91F7846C5B84451FD6A239A" blockId="12.[151,662,1662,1725]" box="[151,643,1662,1689]" pageId="12" pageNumber="13">
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<heading id="DF561103B91F7846C5B84451FD6A239A" bold="true" box="[151,643,1662,1689]" fontSize="11" level="2" pageId="12" pageNumber="13" reason="4">
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<emphasis id="B6D57A7DB91F7846C5B84451FD6A239A" bold="true" box="[151,643,1662,1689]" pageId="12" pageNumber="13">
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<taxonomicName id="43A1DDECB91F7846C5B84451FE0D239B" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,484,1662,1689]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="camerocomplexa" status="sp. nov.">
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<emphasis id="B6D57A7DB91F7846C5B84451FE0D239B" bold="true" box="[151,484,1662,1689]" italics="true" pageId="12" pageNumber="13">Hydraena camerocomplexa</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="ADE6C706B91F7846C4DC4450FD6A239A" box="[499,643,1662,1689]" pageId="12" pageNumber="13" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B91F7846C5B8448CFD7F23BF" box="[151,662,1699,1725]" pageId="12" pageNumber="13" type="description">
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<paragraph id="841EA66FB91F7846C5B8448CFD7F23BF" blockId="12.[151,662,1662,1725]" box="[151,662,1699,1725]" pageId="12" pageNumber="13">
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<figureCitation id="1C9ABAEAB91F7846C5B8448CFF0D23BF" box="[151,228,1699,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1435,633,1769]" captionTargetId="figure-196@5.[151,1435,633,1769]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 2." figureDoi="http://doi.org/10.5281/zenodo.7284849" httpUri="https://zenodo.org/record/7284849/files/figure.png" pageId="12" pageNumber="13">Figs. 3</figureCitation>
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,
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<figureCitation id="1C9ABAEAB91F7846C5DE448CFEE423BF" box="[241,269,1699,1725]" captionStart="FIGURE 13" captionStartId="38.[151,250,1967,1991]" captionTargetBox="[189,1398,188,1918]" captionTargetId="figure-25@38.[151,1436,181,1943]" captionTargetPageId="38" captionText="FIGURE 13. Hydraena camerocomplexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284880" httpUri="https://zenodo.org/record/7284880/files/figure.png" pageId="12" pageNumber="13">13</figureCitation>
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(habitus), 13 (aedeagus), 42 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B91F7846C5B844C4FCA0222B" pageId="12" pageNumber="13" type="materials_examined">
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<paragraph id="841EA66FB91F7846C5B844C4FCA0222B" blockId="12.[151,1436,1770,2013]" pageId="12" pageNumber="13">
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<emphasis id="B6D57A7DB91F7846C5B844C4FEA02206" bold="true" box="[151,329,1770,1797]" pageId="12" pageNumber="13">Type Material.</emphasis>
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<materialsCitation id="34C9AC32B91F7846C47E44C4FEB6222B" ID-GBIF-Occurrence="3962178309" collectingDate="1982-04-10" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="12" pageNumber="13" specimenCount="950" typeStatus="holotype">
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<typeStatus id="5B1A18CDB91F7846C47E44C4FE502207" box="[337,441,1771,1797]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
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(male):
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<emphasis id="B6D57A7DB91F7846C73444C5FDC02206" bold="true" box="[539,553,1770,1796]" pageId="12" pageNumber="13">“</emphasis>
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<collectingCountry id="FCB6E6FFB91F7846C70644C4FD772207" box="[553,670,1771,1797]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
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:
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<location id="817EF0B4B91F7846C78144C4FCFC2207" LSID="urn:lsid:plazi:treatment:0C081779B91F7847C5B84451FB28263B:817EF0B4B91F7846C78144C4FCFC2207" box="[686,789,1771,1797]" country="Cameroon" municipality="Mt. Febe" name="Yaounde" pageId="12" pageNumber="13">Yaounde</location>
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:
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<collectingMunicipality id="647A3C15B91F7846C60F44C4FC602207" box="[800,905,1771,1797]" pageId="12" pageNumber="13">Mt. Febe</collectingMunicipality>
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,
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<specimenCount id="92A76DE6B91F7846C6B944C5FC0E2207" box="[918,999,1770,1797]" pageId="12" pageNumber="13" type="generic">
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<quantity id="43590B8AB91F7846C6B944C5FC0A2207" box="[918,995,1770,1797]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="12" pageNumber="13" unit="m" value="950.0">
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<elevation id="0F8C415CB91F7846C6B944C5FC0A2207" box="[918,995,1770,1797]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="12" pageNumber="13" unit="m" value="950.0">950 m</elevation>
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</quantity>
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.
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</specimenCount>
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, hygropetric,
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<date id="F01F80AFB91F7846C1AB44C4FB152207" box="[1156,1276,1771,1797]" pageId="12" pageNumber="13" value="1982-04-10">
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<collectingDate id="E05B7947B91F7846C1AB44C4FB152207" box="[1156,1276,1771,1797]" pageId="12" pageNumber="13" value="1982-04-10">10.iv.1982</collectingDate>
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</date>
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,
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<collectorName id="2954C3B9B91F7846C02644C4FA752207" box="[1289,1436,1771,1797]" pageId="12" pageNumber="13">P. D. Perkins</collectorName>
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collector” (
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<collectionCode id="E2B03EAAB91F7846C43A4520FEB1222B" box="[277,344,1807,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="13" type="Museum">MCZ</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="34C9AC32B91F7846C4454520FCAC222B" ID-GBIF-Occurrence="3962178342" box="[362,837,1807,1833]" collectingDate="1982-04-10" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" elevation="950" location="Yaounde" municipality="Mt. Febe" pageId="12" pageNumber="13" specimenCount="1" typeStatus="paratype">
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<typeStatus id="5B1A18CDB91F7846C4454520FE39222B" box="[362,464,1807,1833]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
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: Same data as holotype (1
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<collectionCode id="E2B03EAAB91F7846C7D44520FCD4222B" box="[763,829,1807,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="13" type="Museum">MCZ</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B91F7846C5E8451DFD562297" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="841EA66FB91F7846C5E8451DFD562297" blockId="12.[151,1436,1770,2013]" pageId="12" pageNumber="13">
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<emphasis id="B6D57A7DB91F7846C5E8451DFE3D224E" bold="true" box="[199,468,1842,1868]" pageId="12" pageNumber="13">Differential Diagnosis.</emphasis>
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Differentiated from other members of the genus in
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<collectingCountry id="FCB6E6FFB91F7846C10B451CFB70224F" box="[1060,1177,1843,1869]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
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by the combination of the small size (ca.
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<quantity id="43590B8AB91F7846C4474578FE392273" box="[360,464,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="12" pageNumber="13" unit="mm" value="1.16">1.16 mm</quantity>
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), the relatively large and wide pronotum (length/width ratio ca. 17/25), the moderately deep pronotal foveae, and the aedeagus (
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<figureCitation id="1C9ABAEAB91F7846C7704554FD5B2297" box="[607,690,1915,1941]" captionStart="FIGURE 13" captionStartId="38.[151,250,1967,1991]" captionTargetBox="[189,1398,188,1918]" captionTargetId="figure-25@38.[151,1436,181,1943]" captionTargetPageId="38" captionText="FIGURE 13. Hydraena camerocomplexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284880" httpUri="https://zenodo.org/record/7284880/files/figure.png" pageId="12" pageNumber="13">Fig. 13</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B91F7847C5E845B1FCEE27F3" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
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<paragraph id="841EA66FB91F7847C5E845B1FE142467" blockId="12.[151,1436,1770,2013]" lastBlockId="13.[151,1437,151,825]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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<emphasis id="B6D57A7DB91F7846C5E845B1FEB822BA" bold="true" box="[199,337,1950,1976]" pageId="12" pageNumber="13">Description</emphasis>
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. Size:
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<typeStatus id="5B1A18CDB91F7846C48B45B0FDEF22BB" box="[420,518,1951,1977]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.16/0.54; head width 0.33; pronotum 0.31/0.46, PA 0.39, PB 0.41; elytra 0.81/0.54. Habitus as illustrated (
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<figureCitation id="1C9ABAEAB91F7846C12145ECFB8822DF" box="[1038,1121,1987,2013]" captionStart="FIGURE 13" captionStartId="38.[151,250,1967,1991]" captionTargetBox="[189,1398,188,1918]" captionTargetId="figure-25@38.[151,1436,181,1943]" captionTargetPageId="38" captionText="FIGURE 13. Hydraena camerocomplexa, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284880" httpUri="https://zenodo.org/record/7284880/files/figure.png" pageId="12" pageNumber="13">Fig. 13</figureCitation>
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). Dorsum light brown, head and pronotal macula darker (
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<typeStatus id="5B1A18CDB91E7847C4F842B8FDD225B3" box="[471,571,151,177]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
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is teneral), palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum densely moderately finely punctulate, interstices weakly shining; punctures of frons disc and pronotal disc ca. 1xef, interstices ca. 1–2xpd; punctures of clypeus very fine and dense. Labrum apicomedially excised, free margins rounded, weakly upturned. Pronotum relatively large and wide, length/width ratio ca. 17/25. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae shallow, posterosubmedial foveae moderately deep.
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</paragraph>
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<paragraph id="841EA66FB91E7847C5E84340FCEE27F3" blockId="13.[151,1437,151,825]" pageId="13" pageNumber="14">Elytra with posterior declivity very gradual, summit slightly beyond midlength; lateral explanate margins moderately wide; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another. Venter: Mentum and postmentum finely sparsely punctulate, weakly shining. Genae very slightly raised, shining, with weak posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt. Plaques moderately narrow, straight, slightly convergent anteriorly, weakly raised, located at sides of moderately deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 1.5x P2. Pro- meso- and metatibiae moderately stout, straight. Abdominal apex symmetrical, without apicomedian notch.</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B91E7847C5E840D5FC212617" box="[199,968,762,789]" pageId="13" pageNumber="14" type="distribution">
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<paragraph id="841EA66FB91E7847C5E840D5FC212617" blockId="13.[151,1437,151,825]" box="[199,968,762,789]" pageId="13" pageNumber="14">
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<emphasis id="B6D57A7DB91E7847C5E840D5FE882616" bold="true" box="[199,353,762,788]" pageId="13" pageNumber="14">Distribution.</emphasis>
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Currently known only from the
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<typeStatus id="5B1A18CDB91E7847C7E340D4FD152617" box="[716,764,763,789]" pageId="13" pageNumber="14">type</typeStatus>
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locality (
|
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<figureCitation id="1C9ABAEAB91E7847C64840D4FC522617" box="[871,955,763,789]" captionStart="FIGURES 42–43" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 42–43. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="13" pageNumber="14">Fig. 42</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B91E7847C5E84130FB28263B" box="[199,1217,799,825]" pageId="13" pageNumber="14" type="etymology">
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<paragraph id="841EA66FB91E7847C5E84130FB28263B" blockId="13.[151,1437,151,825]" box="[199,1217,799,825]" pageId="13" pageNumber="14">
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<emphasis id="B6D57A7DB91E7847C5E84130FEA7263B" bold="true" box="[199,334,799,825]" pageId="13" pageNumber="14">Etymology.</emphasis>
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Named in reference to the known distribution and the very complex aedeagus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |