132 lines
15 KiB
XML
132 lines
15 KiB
XML
<document id="F8591F923D31ED10D6CDE494D56AF096" ID-CLB-Dataset="181704" ID-DOI="10.11646/zootaxa.5203.1.1" ID-GBIF-Dataset="b6f94c7b-7033-4952-96a7-3db5fb6ad879" ID-ISSN="1175-5326" ID-Zenodo-Dep="7284841" ID-ZooBank="85C20298-5DF9-44DF-8485-3C64CF40CD08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667554441660" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2022" docId="0C081779B9197840C5B84069FAF02377" docLanguage="en" docName="zootaxa.5203.1.1.pdf" docOrigin="Zootaxa 5203 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5203.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena camerojocula Perkins 2022, new species" docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="F0316F01B913784AC52F422FFFE92502" masterDocTitle="Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="11" updateTime="1699409684053" updateUser="ExternalLinkService">
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<mods:titleInfo id="CCC10F8F5D3B2609E6DF9786F29CAC00">
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<mods:title id="F8595ECB856970F7F1C988FF5422E535">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<mods:name id="26C4E91FCB8807EA0F266D9EDA6F7403" type="personal">
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<mods:namePart id="A738245758A36AA17B875AC001CE9B60">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="E225949E9A2ECC996C9722BADE9B7784">text</mods:typeOfResource>
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<mods:title id="4145D12EAF5CA1421974BD7C6BFFC0D6">Zootaxa</mods:title>
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<mods:part id="FF362D890E1D9A86A1D1653A2C724DE1">
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<mods:date id="AE803D15F5AA9327E23ADBC4543AA76C">2022</mods:date>
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<mods:number id="8F1DE6A9D5392BC8F2484E20C1CFF099">2022-11-03</mods:number>
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<mods:number id="653A2628C3824F6C8EEC4D7055CEA61E">5203</mods:number>
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<mods:url id="5C3B9CEEE8122FACB0B61DE792CF8DFF">http://dx.doi.org/10.11646/zootaxa.5203.1.1</mods:url>
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<mods:classification id="36FC80B6798A16C88A85F50EB6680C8C">journal article</mods:classification>
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<mods:identifier id="C5FF06E490333B3AAA68F707309DA6C9" type="CLB-Dataset">181704</mods:identifier>
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<mods:identifier id="30016BA13B0345B72A39EA3F6FF12DF4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0C081779B9197840C5B84069FAF02377" ID-DOI="http://doi.org/10.5281/zenodo.7300186" ID-GBIF-Taxon="204205212" ID-Zenodo-Dep="7300186" LSID="urn:lsid:plazi:treatment:0C081779B9197840C5B84069FAF02377" httpUri="http://treatment.plazi.org/id/0C081779B9197840C5B84069FAF02377" lastPageNumber="11" pageId="10" pageNumber="11">
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<subSubSection id="CCBBF5E4B9197840C5B84069FDB32762" box="[151,602,582,609]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="841EA66FB9197840C5B84069FDB32762" blockId="10.[151,634,582,645]" box="[151,602,582,609]" pageId="10" pageNumber="11">
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<heading id="DF561103B9197840C5B84069FDB32762" bold="true" box="[151,602,582,609]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="4">
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<emphasis id="B6D57A7DB9197840C5B84069FDB32762" bold="true" box="[151,602,582,609]" pageId="10" pageNumber="11">
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<taxonomicName id="43A1DDECB9197840C5B84069FE552763" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,444,582,609]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="camerojocula" status="sp. nov.">
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<emphasis id="B6D57A7DB9197840C5B84069FE552763" bold="true" box="[151,444,582,609]" italics="true" pageId="10" pageNumber="11">Hydraena camerojocula</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="ADE6C706B9197840C4E54068FDB32762" box="[458,602,582,609]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B9197840C5B84044FD932787" box="[151,634,619,645]" pageId="10" pageNumber="11" type="description">
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<paragraph id="841EA66FB9197840C5B84044FD932787" blockId="10.[151,634,582,645]" box="[151,634,619,645]" pageId="10" pageNumber="11">
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<figureCitation id="1C9ABAEAB9197840C5B84044FF0A2787" box="[151,227,619,645]" captionStart="FIGURES 1–2" captionStartId="4.[151,264,1963,1987]" captionTargetBox="[232,1355,525,1939]" captionTargetId="figure-127@4.[232,1355,525,1939]" captionTargetPageId="4" captionText="FIGURES 1–2. 1. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 1; 2. Representative aedeagi of Aedeagal Species Groups 1–4." figureDoi="http://doi.org/10.5281/zenodo.7284846" httpUri="https://zenodo.org/record/7284846/files/figure.png" pageId="10" pageNumber="11">Figs. 1</figureCitation>
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,
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<figureCitation id="1C9ABAEAB9197840C5DE4044FF162787" box="[241,255,619,645]" captionStart="FIGURE 9" captionStartId="34.[151,250,1938,1962]" captionTargetBox="[227,1362,212,1883]" captionTargetId="figure-25@34.[164,1424,181,1914]" captionTargetPageId="34" captionText="FIGURE 9. Hydraena camerojocula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284866" httpUri="https://zenodo.org/record/7284866/files/figure.png" pageId="10" pageNumber="11">9</figureCitation>
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(habitus), 9 (aedeagus), 41 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B9197840C5B8409CFDD627F3" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="841EA66FB9197840C5B8409CFDD627F3" blockId="10.[151,1437,690,1653]" pageId="10" pageNumber="11">
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<emphasis id="B6D57A7DB9197840C5B8409CFEAE27CE" bold="true" box="[151,327,690,717]" pageId="10" pageNumber="11">Type Material.</emphasis>
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<materialsCitation id="34C9AC32B9197840C462409CFDD627F3" ID-GBIF-Occurrence="3962178356" collectingDate="1982-05-22" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Nyong River" pageId="10" pageNumber="11" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5B1A18CDB9197840C462409CFE5C27CF" box="[333,437,691,717]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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(male): “
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<collectingCountry id="FCB6E6FFB9197840C732409CFD7127CF" box="[541,664,691,717]" name="Cameroon" pageId="10" pageNumber="11">Cameroon</collectingCountry>
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:
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<collectorName id="2954C3B9B9197840C78D409CFCF527CF" box="[674,796,691,717]" pageId="10" pageNumber="11">C. S. Prov.</collectorName>
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,
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<quantity id="43590B8AB9197840C60A409DFC8627CF" box="[805,879,690,717]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="km" value="15.0">15 km</quantity>
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.
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<collectorName id="2954C3B9B9197840C656409CFBF827CF" box="[889,1041,691,717]" pageId="10" pageNumber="11">S. Mbalmayo</collectorName>
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,
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<location id="817EF0B4B9197840C133409CFB4727CF" LSID="urn:lsid:plazi:treatment:0C081779B9197840C5B84069FAF02377:817EF0B4B9197840C133409CFB4727CF" box="[1052,1198,691,717]" country="Cameroon" name="Nyong River" pageId="10" pageNumber="11">Nyong River</location>
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, ex. leaves at margin,
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<date id="F01F80AFB9197840C5B840F8FEE127F3" box="[151,264,727,753]" pageId="10" pageNumber="11" value="1982-05-22">
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<collectingDate id="E05B7947B9197840C5B840F8FEE127F3" box="[151,264,727,753]" pageId="10" pageNumber="11" value="1982-05-22">22.v.1982</collectingDate>
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</date>
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, leg.
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<collectorName id="2954C3B9B9197840C46B40F8FE3E27F3" box="[324,471,727,753]" pageId="10" pageNumber="11">P. D. Perkins</collectorName>
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” (
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<collectionCode id="E2B03EAAB9197840C4DF40F8FDDA27F3" box="[496,563,727,753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="10" pageNumber="11" type="Museum">MCZ</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B9197840C5E840D5FABA2683" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="841EA66FB9197840C5E840D5FABA2683" blockId="10.[151,1437,690,1653]" pageId="10" pageNumber="11">
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<emphasis id="B6D57A7DB9197840C5E840D5FE3D2616" bold="true" box="[199,468,762,788]" pageId="10" pageNumber="11">Differential Diagnosis.</emphasis>
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Differentiated from other members of the genus in
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<collectingCountry id="FCB6E6FFB9197840C10B40D4FB702617" box="[1060,1177,763,789]" name="Cameroon" pageId="10" pageNumber="11">Cameroon</collectingCountry>
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by the combination of the posterosubmedian foveae moderately deep, medially confluent forming U-shaped depression, the metaventral plaques moderately narrow posteriorly, tapering to point anteriorly, and the aedeagus (
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<figureCitation id="1C9ABAEAB9197840C17F416CFB7A265F" box="[1104,1171,835,861]" captionStart="FIGURE 9" captionStartId="34.[151,250,1938,1962]" captionTargetBox="[227,1362,212,1883]" captionTargetId="figure-25@34.[164,1424,181,1914]" captionTargetPageId="34" captionText="FIGURE 9. Hydraena camerojocula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284866" httpUri="https://zenodo.org/record/7284866/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
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). Dorsal habitus similar to other species in Aedeagal Group 1; reliable determinations will require examination of the male genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B9197840C5E841A5FD4E232F" pageId="10" pageNumber="11" type="description">
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<paragraph id="841EA66FB9197840C5E841A5FDB221A3" blockId="10.[151,1437,690,1653]" pageId="10" pageNumber="11">
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<emphasis id="B6D57A7DB9197840C5E841A5FEB826A6" bold="true" box="[199,337,906,932]" pageId="10" pageNumber="11">Description</emphasis>
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. Size:
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<typeStatus id="5B1A18CDB9197840C48B41A4FDEF26A7" box="[420,518,907,933]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.35/0.60; head width 0.36; pronotum 0.32/0.46, PA 0.37, PB 0.41; elytra 0.82/0.60. Habitus as illustrated (
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<figureCitation id="1C9ABAEAB9197840C11D4180FB9326CB" box="[1074,1146,943,969]" captionStart="FIGURE 9" captionStartId="34.[151,250,1938,1962]" captionTargetBox="[227,1362,212,1883]" captionTargetId="figure-25@34.[164,1424,181,1914]" captionTargetPageId="34" captionText="FIGURE 9. Hydraena camerojocula, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284866" httpUri="https://zenodo.org/record/7284866/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
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). Dorsum generally dark brown, except labrum, clypeus and anterolateral angles of pronotum lighter, palpi testaceous, tip of last palpomere not darker. Frons moderately finely and moderately densely punctulate, pronotum moderately coarsely, moderately densely punctate, punctures much larger than those of frons, interstices shining, ca. 1–2xpd. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, posterolateral foveae very shallow, posterosubmedian foveae moderately deep, medially confluent forming U-shaped depression.
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</paragraph>
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<paragraph id="841EA66FB9197840C5E84684FD4E232F" blockId="10.[151,1437,690,1653]" pageId="10" pageNumber="11">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or slightly greater, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming moderately strong angle with one another. Venter: Mentum sparsely finely punctulate, shining; postmentum very finely densely micropunctulate, surrounding areas smooth, moderately shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow posteriorly, tapering to point anteriorly, slightly converging toward one another anteriorly, weakly raised, located at sides of shallow median depression. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, straight. Abdominal apex asymmetrical, with deep off-center notch.</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B9197840C5E84419FC212353" box="[199,968,1590,1617]" pageId="10" pageNumber="11" type="distribution">
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<paragraph id="841EA66FB9197840C5E84419FC212353" blockId="10.[151,1437,690,1653]" box="[199,968,1590,1617]" pageId="10" pageNumber="11">
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<emphasis id="B6D57A7DB9197840C5E84419FE882352" bold="true" box="[199,353,1590,1616]" pageId="10" pageNumber="11">Distribution.</emphasis>
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Currently known only from the
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<typeStatus id="5B1A18CDB9197840C7E34418FD152353" box="[716,764,1591,1617]" pageId="10" pageNumber="11">type</typeStatus>
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locality (
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<figureCitation id="1C9ABAEAB9197840C6484418FC532353" box="[871,954,1591,1617]" captionStart="FIGURES 39–41" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 39–41. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="10" pageNumber="11">Fig. 41</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B9197840C5E84474FAF02377" box="[199,1305,1627,1653]" pageId="10" pageNumber="11" type="etymology">
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<paragraph id="841EA66FB9197840C5E84474FAF02377" blockId="10.[151,1437,690,1653]" box="[199,1305,1627,1653]" pageId="10" pageNumber="11">
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<emphasis id="B6D57A7DB9197840C5E84474FEA72377" bold="true" box="[199,334,1627,1653]" pageId="10" pageNumber="11">Etymology.</emphasis>
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Named in reference to the known distribution, and to the smile shaped pronotal fovea.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |