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<document id="1BF2E6143118EAC8270C2AA67A65C0A7" ID-CLB-Dataset="181704" ID-DOI="10.11646/zootaxa.5203.1.1" ID-GBIF-Dataset="b6f94c7b-7033-4952-96a7-3db5fb6ad879" ID-ISSN="1175-5326" ID-Zenodo-Dep="7284841" ID-ZooBank="85C20298-5DF9-44DF-8485-3C64CF40CD08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667554441660" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2022" docId="0C081779B90F7856C5B8409DFB1823E3" docLanguage="en" docName="zootaxa.5203.1.1.pdf" docOrigin="Zootaxa 5203 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5203.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena cameroradiata Perkins 2022, new species" docType="treatment" docVersion="7" lastPageNumber="29" masterDocId="F0316F01B913784AC52F422FFFE92502" masterDocTitle="Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="29" updateTime="1699409684053" updateUser="ExternalLinkService">
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<mods:title id="DCF78AD296752C7A410466812C5E8F20">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="2CDAFAE5AF98DA6B3C21B0EA63FD610A">Perkins, Philip D.</mods:namePart>
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<mods:title id="53AFB878617B946AB161BB79F80406A7">Zootaxa</mods:title>
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<mods:date id="AC57FCB33BFDFAD94726FC9B1258A256">2022</mods:date>
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<mods:number id="5715F95775E5D589E1E149CDF9B1E9B1">2022-11-03</mods:number>
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<mods:classification id="291C9D0A25758ADD9FEC93EB8F0DDAA4">journal article</mods:classification>
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<mods:identifier id="BDA02B219F1266CEF9C9CE225BB03E92" type="DOI">10.11646/zootaxa.5203.1.1</mods:identifier>
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<treatment id="0C081779B90F7856C5B8409DFB1823E3" ID-DOI="http://doi.org/10.5281/zenodo.7300284" ID-GBIF-Taxon="204205215" ID-Zenodo-Dep="7300284" LSID="urn:lsid:plazi:treatment:0C081779B90F7856C5B8409DFB1823E3" httpUri="http://treatment.plazi.org/id/0C081779B90F7856C5B8409DFB1823E3" lastPageNumber="29" pageId="28" pageNumber="29">
<subSubSection id="CCBBF5E4B90F7856C5B8409DFD8F27CE" box="[151,614,690,717]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="841EA66FB90F7856C5B8409DFD8F27CE" blockId="28.[151,662,690,753]" box="[151,614,690,717]" pageId="28" pageNumber="29">
<heading id="DF561103B90F7856C5B8409DFD8F27CE" bold="true" box="[151,614,690,717]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="B6D57A7DB90F7856C5B8409DFD8F27CE" bold="true" box="[151,614,690,717]" pageId="28" pageNumber="29">
<taxonomicName id="43A1DDECB90F7856C5B8409DFE2E27CF" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,455,690,717]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cameroradiata" status="sp. nov.">
<emphasis id="B6D57A7DB90F7856C5B8409DFE2E27CF" bold="true" box="[151,455,690,717]" italics="true" pageId="28" pageNumber="29">Hydraena cameroradiata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B90F7856C4F9409CFD8F27CE" box="[470,614,690,717]" pageId="28" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CCBBF5E4B90F7856C5B840F8FD7F27F3" box="[151,662,726,753]" pageId="28" pageNumber="29" type="description">
<paragraph id="841EA66FB90F7856C5B840F8FD7F27F3" blockId="28.[151,662,690,753]" box="[151,662,726,753]" pageId="28" pageNumber="29">
<figureCitation id="1C9ABAEAB90F7856C5B840F8FF0C27F3" box="[151,229,726,753]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="28" pageNumber="29">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB90F7856C5DE40F8FEE427F3" box="[241,269,727,753]" captionStart="FIGURE 37" captionStartId="62.[151,250,1996,2020]" captionTargetBox="[269,1319,181,1973]" captionTargetId="figure-25@62.[269,1319,181,1973]" captionTargetPageId="62" captionText="FIGURE 37. Hydraena cameroradiata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284959" httpUri="https://zenodo.org/record/7284959/files/figure.png" pageId="28" pageNumber="29">37</figureCitation>
(habitus), 37 (aedeagus), 41 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5B84130FD05265F" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="841EA66FB90F7856C5B84130FD05265F" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5B84130FEAE263A" bold="true" box="[151,327,798,825]" pageId="28" pageNumber="29">Type Material.</emphasis>
<materialsCitation id="34C9AC32B90F7856C4624130FF03265F" ID-GBIF-Occurrence="3962178355" collectingDate="1981-12-12" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Yaounde" municipality="Bastos" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB90F7856C4624130FE5C263B" box="[333,437,799,825]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB90F7856C7334130FD7E263B" box="[540,663,799,825]" name="Cameroon" pageId="28" pageNumber="29">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B90F7856C78E4130FCF2263B" box="[673,795,799,825]" pageId="28" pageNumber="29">C. S. Prov.</collectorName>
,
<location id="817EF0B4B90F7856C60B4130FC62263B" LSID="urn:lsid:plazi:treatment:0C081779B90F7856C5B8409DFB1823E3:817EF0B4B90F7856C60B4130FC62263B" box="[804,907,799,825]" country="Cameroon" municipality="Bastos" name="Yaounde" pageId="28" pageNumber="29">Yaounde</location>
:
<collectingMunicipality id="647A3C15B90F7856C6BB4130FC08263B" box="[916,993,799,825]" pageId="28" pageNumber="29">Bastos</collectingMunicipality>
, puddle,
<date id="F01F80AFB90F7856C16A4130FB2E263B" box="[1093,1223,799,825]" pageId="28" pageNumber="29" value="1981-12-12">
<collectingDate id="E05B7947B90F7856C16A4130FB2E263B" box="[1093,1223,799,825]" pageId="28" pageNumber="29" value="1981-12-12">12.xii.1981</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B90F7856C02E4130FA7A263B" box="[1281,1427,799,825]" pageId="28" pageNumber="29">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB90F7856C5B0416CFF0B265F" box="[159,226,835,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="29" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B90F7856C5DA416CFD01265F" ID-GBIF-Occurrence="3962178354" box="[245,744,835,861]" collectingDate="1981-12-12" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Yaounde" municipality="Bastos" pageId="28" pageNumber="29" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB90F7856C5DA416CFE8F265F" box="[245,358,835,861]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
: Same data as holotype (10
<collectionCode id="E2B03EAAB90F7856C7B0416CFD08265F" box="[671,737,835,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="29" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E84149FE2E26EF" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="841EA66FB90F7856C5E84149FE2E26EF" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E84149FE382682" bold="true" box="[199,465,870,896]" pageId="28" pageNumber="29">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB90F7856C1274148FB942683" box="[1032,1149,871,897]" name="Cameroon" pageId="28" pageNumber="29">Cameroon</collectingCountry>
by the combination of the very sparcely and very finely punctulate head and pronotum, interstices strongly shining, the anterolateral pronotal foveae moderately deep, posterolateral and posterosubmedial foveae absent, the absence of metaventral plaques, and the aedeagus (
<figureCitation id="1C9ABAEAB90F7856C44941FCFE5326EF" box="[358,442,979,1005]" captionStart="FIGURE 37" captionStartId="62.[151,250,1996,2020]" captionTargetBox="[269,1319,181,1973]" captionTargetId="figure-25@62.[269,1319,181,1973]" captionTargetPageId="62" captionText="FIGURE 37. Hydraena cameroradiata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284959" httpUri="https://zenodo.org/record/7284959/files/figure.png" pageId="28" pageNumber="29">Fig. 37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E841D9FDE3239B" pageId="28" pageNumber="29" type="description">
<paragraph id="841EA66FB90F7856C5E841D9FDDD2033" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E841D9FEB82112" bold="true" box="[199,337,1014,1040]" pageId="28" pageNumber="29">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB90F7856C48B41D8FDEF2113" box="[420,518,1015,1041]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.33/0.59; head width 0.34; pronotum 0.35/0.47, PA 0.38, PB 0.43; elytra 0.80/0.59. Habitus as illustrated (
<figureCitation id="1C9ABAEAB90F7856C1074634FB962137" box="[1064,1151,1051,1077]" captionStart="FIGURE 37" captionStartId="62.[151,250,1996,2020]" captionTargetBox="[269,1319,181,1973]" captionTargetId="figure-25@62.[269,1319,181,1973]" captionTargetPageId="62" captionText="FIGURE 37. Hydraena cameroradiata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284959" httpUri="https://zenodo.org/record/7284959/files/figure.png" pageId="28" pageNumber="29">Fig. 37</figureCitation>
). Dorsum brown to light brown, frons and diffusely margined pronotal macula slightly darker, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum sparcely finely punctulate, pronotum more finely sparsely punctulate on macula than on surrounding areas, interstices shining; punctures of pronotal disc ca. 1xef, interstices ca. 35xpd or slightly larger; punctures of frons disc slightly larger and more closely spaced than those of pronotal disc; punctures of clypeus very fine, nearly obsolete. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae moderately deep, posterolateral and posterosubmedial foveae absent.
</paragraph>
<paragraph id="841EA66FB90F7856C5E84714FDD420E7" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins narrow; punctures fine, on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 24xpd, as are interstices between punctures of a row. Apices in dorsal aspect separately rounded, in posterior aspect margins forming moderately strong angle with one another.</paragraph>
<paragraph id="841EA66FB90F7856C5B847C0FDE3239B" blockId="28.[151,1437,798,1761]" pageId="28" pageNumber="29">Venter: Mentum and postmentum sparsely finely punctulate, shining. Genae raised, shining, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques absent, median depression deep. AIS width at straight posterior margin ca. 2x P2. All tibiae slender. Pro- and mesotibiae very slightly arcuate, metatibia straight. Abdominal apex symmetrical, with shallow apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E8448DFC2123BF" box="[199,968,1698,1725]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="841EA66FB90F7856C5E8448DFC2123BF" blockId="28.[151,1437,798,1761]" box="[199,968,1698,1725]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E8448DFE8823BE" bold="true" box="[199,353,1698,1724]" pageId="28" pageNumber="29">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB90F7856C7E3448CFD1523BF" box="[716,764,1699,1725]" pageId="28" pageNumber="29">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB90F7856C648448CFC5323BF" box="[871,954,1699,1725]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="28" pageNumber="29">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90F7856C5E844E8FB1823E3" box="[199,1265,1735,1761]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="841EA66FB90F7856C5E844E8FB1823E3" blockId="28.[151,1437,798,1761]" box="[199,1265,1735,1761]" pageId="28" pageNumber="29">
<emphasis id="B6D57A7DB90F7856C5E844E8FEA723E3" bold="true" box="[199,334,1735,1761]" pageId="28" pageNumber="29">Etymology.</emphasis>
Named in reference to the known distribution, and to the highly reflective dorsum.
</paragraph>
</subSubSection>
</treatment>
</document>