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<document id="A6DF314BCF14C2D6EAC2689D15C702AA" ID-CLB-Dataset="181704" ID-DOI="10.11646/zootaxa.5203.1.1" ID-GBIF-Dataset="b6f94c7b-7033-4952-96a7-3db5fb6ad879" ID-ISSN="1175-5326" ID-Zenodo-Dep="7284841" ID-ZooBank="85C20298-5DF9-44DF-8485-3C64CF40CD08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667554441660" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2022" docId="0C081779B90A7850C5B847C1FADD263B" docLanguage="en" docName="zootaxa.5203.1.1.pdf" docOrigin="Zootaxa 5203 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5203.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena camerocordata Perkins 2022, new species" docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="F0316F01B913784AC52F422FFFE92502" masterDocTitle="Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="26" updateTime="1699409684053" updateUser="ExternalLinkService">
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<mods:title id="FA6DFFC9C987DAAFBF16C0D1F0BC99CE">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="23AAAA8F0E1C8D3D3B3005C05DCEDB89">Perkins, Philip D.</mods:namePart>
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<mods:date id="2091D3451559B3245ABC30CCCFD4BDCF">2022</mods:date>
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<mods:number id="2ABE54CDE7757DA3C50FB9FAC3411356">2022-11-03</mods:number>
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<mods:classification id="F832D5828B965FAD6F83E47835AF7D6A">journal article</mods:classification>
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<mods:identifier id="8B5321827DC78BEB89D256741A71FD58" type="DOI">10.11646/zootaxa.5203.1.1</mods:identifier>
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<treatment id="0C081779B90A7850C5B847C1FADD263B" ID-DOI="http://doi.org/10.5281/zenodo.7300280" ID-GBIF-Taxon="204205218" ID-Zenodo-Dep="7300280" LSID="urn:lsid:plazi:treatment:0C081779B90A7850C5B847C1FADD263B" httpUri="http://treatment.plazi.org/id/0C081779B90A7850C5B847C1FADD263B" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<subSubSection id="CCBBF5E4B90A7853C5B847C1FD83230A" box="[151,618,1518,1545]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="841EA66FB90A7853C5B847C1FD83230A" blockId="25.[151,662,1518,1581]" box="[151,618,1518,1545]" pageId="25" pageNumber="26">
<heading id="DF561103B90A7853C5B847C1FD83230A" bold="true" box="[151,618,1518,1545]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="4">
<emphasis id="B6D57A7DB90A7853C5B847C1FD83230A" bold="true" box="[151,618,1518,1545]" pageId="25" pageNumber="26">
<taxonomicName id="43A1DDECB90A7853C5B847C1FE25230B" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,460,1518,1545]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="camerocordata" status="sp. nov.">
<emphasis id="B6D57A7DB90A7853C5B847C1FE25230B" bold="true" box="[151,460,1518,1545]" italics="true" pageId="25" pageNumber="26">Hydraena camerocordata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B90A7853C4F447C0FD83230A" box="[475,618,1518,1545]" pageId="25" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CCBBF5E4B90A7853C5B8443CFD7F232F" box="[151,662,1554,1581]" pageId="25" pageNumber="26" type="description">
<paragraph id="841EA66FB90A7853C5B8443CFD7F232F" blockId="25.[151,662,1518,1581]" box="[151,662,1554,1581]" pageId="25" pageNumber="26">
<figureCitation id="1C9ABAEAB90A7853C5B8443CFF0C232F" box="[151,229,1554,1581]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="25" pageNumber="26">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB90A7853C5DE443CFEE4232F" box="[241,269,1555,1581]" captionStart="FIGURE 33" captionStartId="58.[151,250,1989,2013]" captionTargetBox="[290,1325,191,1941]" captionTargetId="figure-25@58.[210,1377,181,1965]" captionTargetPageId="58" captionText="FIGURE 33. Hydraena camerocordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284945" httpUri="https://zenodo.org/record/7284945/files/figure.png" pageId="25" pageNumber="26">33</figureCitation>
(habitus), 33 (aedeagus), 41 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B90A7850C5B84474FA8427F3" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="841EA66FB90A7853C5B84474FC81239B" blockId="25.[151,1437,1626,2013]" pageId="25" pageNumber="26">
<emphasis id="B6D57A7DB90A7853C5B84474FEA02376" bold="true" box="[151,329,1626,1653]" pageId="25" pageNumber="26">Type Material.</emphasis>
<materialsCitation id="34C9AC32B90A7853C47F4474FE9D239B" ID-GBIF-Occurrence="3962178317" collectingDate="1982-04-22" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Etoug Ebe" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB90A7853C47F4474FE512377" box="[336,440,1627,1653]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB90A7853C70C4474FD772377" box="[547,670,1627,1653]" name="Cameroon" pageId="25" pageNumber="26">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B90A7853C7864474FCCF2377" box="[681,806,1627,1653]" pageId="25" pageNumber="26">C. S. Prov.</collectorName>
,
<location id="817EF0B4B90A7853C61D4474FC432377" LSID="urn:lsid:plazi:treatment:0C081779B90A7850C5B847C1FADD263B:817EF0B4B90A7853C61D4474FC432377" box="[818,938,1627,1653]" country="Cameroon" name="Etoug Ebe" pageId="25" pageNumber="26">Etoug Ebe</location>
, stream with dead leaves,
<date id="F01F80AFB90A7853C1FC4474FAA52377" box="[1235,1356,1627,1653]" pageId="25" pageNumber="26" value="1982-04-22">
<collectingDate id="E05B7947B90A7853C1FC4474FAA52377" box="[1235,1356,1627,1653]" pageId="25" pageNumber="26" value="1982-04-22">22.iv.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B90A7853C0A64474FEF9239B" pageId="25" pageNumber="26">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB90A7853C4064450FE85239B" box="[297,364,1663,1689]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="34C9AC32B90A7853C4514450FC8D239B" ID-GBIF-Occurrence="3962178318" box="[382,868,1663,1689]" collectingDate="1982-04-22" collectionCode="MCZ" collectorName="C. S. Prov. &amp; P. D. Perkins" country="Cameroon" location="Etoug Ebe" pageId="25" pageNumber="26" specimenCount="1" typeStatus="paratype">
<typeStatus id="5B1A18CDB90A7853C4514450FE06239B" box="[382,495,1663,1689]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
: Same data as holotype (2
<collectionCode id="E2B03EAAB90A7853C6354450FCB5239B" box="[794,860,1663,1689]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="25" pageNumber="26" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="841EA66FB90A7853C5E8448CFF2D2207" blockId="25.[151,1437,1626,2013]" pageId="25" pageNumber="26">
Differential Diagnosis. Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB90A7853C12B448CFB9023BF" box="[1028,1145,1699,1725]" name="Cameroon" pageId="25" pageNumber="26">Cameroon</collectingCountry>
by the combination of the large body size (ca.
<quantity id="43590B8AB90A7853C45744E9FE0923E3" box="[376,480,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="25" pageNumber="26" unit="mm" value="1.53">1.53 mm</quantity>
), the markedly cordiform pronotum, the very wide P1 and P2, and the aedeagus (
<figureCitation id="1C9ABAEAB90A7853C04044E8FF5F2207" captionStart="FIGURE 33" captionStartId="58.[151,250,1989,2013]" captionTargetBox="[290,1325,191,1941]" captionTargetId="figure-25@58.[210,1377,181,1965]" captionTargetPageId="58" captionText="FIGURE 33. Hydraena camerocordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284945" httpUri="https://zenodo.org/record/7284945/files/figure.png" pageId="25" pageNumber="26">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="841EA66FB90A7850C5E84520FE582443" blockId="25.[151,1437,1626,2013]" lastBlockId="26.[151,1437,151,825]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Description. Size:
<typeStatus id="5B1A18CDB90A7853C4B24520FE16222B" box="[413,511,1807,1833]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.53/0.69; head width 0.36; pronotum 0.38/0.52, PA 0.41, PB 0.38; elytra 0.90/0.69. Habitus as illustrated (
<figureCitation id="1C9ABAEAB90A7853C133451CFB98224F" box="[1052,1137,1843,1869]" captionStart="FIGURE 33" captionStartId="58.[151,250,1989,2013]" captionTargetBox="[290,1325,191,1941]" captionTargetId="figure-25@58.[210,1377,181,1965]" captionTargetPageId="58" captionText="FIGURE 33. Hydraena camerocordata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284945" httpUri="https://zenodo.org/record/7284945/files/figure.png" pageId="25" pageNumber="26">Fig. 33</figureCitation>
). Pronotum markedly cordiform, width at midlength much wider than anterior and posterior margins. Dorsum dark brown to black, except labrum, clypeus, and maxillary palpi light brown to testaceous, tip of last palpomere not darker. Frons moderately finely moderately sparsely punctate, punctures ca. 1xef. Pronotum coarsely densely punctate, especially anterior and posterior to moderately convex disc, interstices in densely punctate areas narrow walls; interstices on disc ca. 1xpd, shining; punctures laterally much smaller and sparser. Punctures of clypeus very fine, moderately dense on disc, very dense laterally. Labrum apicomedian excision deep and wide, free margins weakly upturned, angulate laterally and medially. Pronotum anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae deep, shallowly confluent; posterosubmedial foveae deep, area between antero- and posterolateral foveae distinctly convex.
</paragraph>
<paragraph id="841EA66FB9097850C5E84364FE74271B" blockId="26.[151,1437,151,825]" pageId="26" pageNumber="27">Elytra with posterior declivity rather abrupt, summit at or very near midlength; lateral explanate margins wide; on basal 1/3 punctures 1xpd or slightly larger and deeper than largest pronotal punctures, punctures becoming gradually slightly smaller laterally and toward posterior. Intervals not raised, width on disc ca. 1xpd, much wider laterally and posteriorly, interstices between punctures of a row ca. 16xpd, each puncture with very short and very fine seta. Apices in dorsal aspect conjointly rather sharply rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="841EA66FB9097850C5B8400CFA8427F3" blockId="26.[151,1437,151,825]" pageId="26" pageNumber="27">Venter: Mentum finely sparsely punctulate, shining. Postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.1/0.2/1.5. P1 and P2 very wide. P2 l/w ca. 1/1, sides parallel, apex blunt. Plaques very small, very short, widely separated carinate lines, located at sides of deep median depression. AIS very wide, weakly concave, width at straight posterior margin ca. 1.2x P2. Tibiae slender, straight. Protibia very slightly widened and bearing short spine at ca. distal 2/3. Mesotibia slightly widened at ca. distal 2/3. Abdominal apex symmetrical, with rather wide and deep apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5E840D5FC212617" box="[199,968,762,789]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="841EA66FB9097850C5E840D5FC212617" blockId="26.[151,1437,151,825]" box="[199,968,762,789]" pageId="26" pageNumber="27">
<emphasis id="B6D57A7DB9097850C5E840D5FEB02616" bold="true" box="[199,345,762,788]" pageId="26" pageNumber="27">Distribution</emphasis>
. Currently known only from the
<typeStatus id="5B1A18CDB9097850C7E340D4FD152617" box="[716,764,763,789]" pageId="26" pageNumber="27">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB9097850C64840D4FC532617" box="[871,954,763,789]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="26" pageNumber="27">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B9097850C5E84130FADD263B" box="[199,1332,799,825]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="841EA66FB9097850C5E84130FADD263B" blockId="26.[151,1437,151,825]" box="[199,1332,799,825]" pageId="26" pageNumber="27">
<emphasis id="B6D57A7DB9097850C5E84130FEA1263B" bold="true" box="[199,328,799,825]" pageId="26" pageNumber="27">Etymology</emphasis>
. Named in reference to the known distribution, and to the markedly cordiform pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>