147 lines
17 KiB
XML
147 lines
17 KiB
XML
<document id="66C5E76AFF54F22E11AF9FB94CCA09EB" ID-CLB-Dataset="181704" ID-DOI="10.11646/zootaxa.5203.1.1" ID-GBIF-Dataset="b6f94c7b-7033-4952-96a7-3db5fb6ad879" ID-ISSN="1175-5326" ID-Zenodo-Dep="7284841" ID-ZooBank="85C20298-5DF9-44DF-8485-3C64CF40CD08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667554441660" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2022" docId="0C081779B907785FC5B846C4FDD424AF" docLanguage="en" docName="zootaxa.5203.1.1.pdf" docOrigin="Zootaxa 5203 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5203.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena millenaria Perkins 2022, new species" docType="treatment" docVersion="5" lastPageNumber="22" masterDocId="F0316F01B913784AC52F422FFFE92502" masterDocTitle="Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="21" updateTime="1699409684053" updateUser="ExternalLinkService">
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<mods:title id="D8B0D877D4F9925FBCFA13889905464A">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<mods:name id="179EC01A242BBD1EBE70BAFEC506B8B6" type="personal">
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<mods:namePart id="2CDF7C6189F60F1E118862E225150995">Perkins, Philip D.</mods:namePart>
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<mods:title id="6E4B658545D58DF8C98AD2D286614B47">Zootaxa</mods:title>
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<mods:part id="746CF3EDAF128763968B7365DED84586">
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<mods:date id="05BF17D3071E6D83E301B4B49A1DAF65">2022</mods:date>
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<mods:number id="163E119F33008812C3505BE0C41C67B9">2022-11-03</mods:number>
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<mods:url id="DB19D208C25460A784C3367213A5CFFC">http://dx.doi.org/10.11646/zootaxa.5203.1.1</mods:url>
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<mods:classification id="2F71EFC65822B7396D48CAE94F98F4CA">journal article</mods:classification>
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<mods:identifier id="5AE3646C6A7FA0ED0A6D67C38150A0E3" type="CLB-Dataset">181704</mods:identifier>
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<treatment id="0C081779B907785FC5B846C4FDD424AF" ID-DOI="http://doi.org/10.5281/zenodo.7300202" ID-GBIF-Taxon="204205222" ID-Zenodo-Dep="7300202" LSID="urn:lsid:plazi:treatment:0C081779B907785FC5B846C4FDD424AF" httpUri="http://treatment.plazi.org/id/0C081779B907785FC5B846C4FDD424AF" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="CCBBF5E4B907785EC5B846C4FDDA2007" box="[151,563,1259,1286]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="841EA66FB907785EC5B846C4FDDA2007" blockId="20.[151,662,1259,1322]" box="[151,563,1259,1286]" pageId="20" pageNumber="21">
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<heading id="DF561103B907785EC5B846C4FDDA2007" bold="true" box="[151,563,1259,1286]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
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<emphasis id="B6D57A7DB907785EC5B846C4FDDA2007" bold="true" box="[151,563,1259,1286]" pageId="20" pageNumber="21">
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<taxonomicName id="43A1DDECB907785EC5B846C4FE7D2004" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,404,1259,1286]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="millenaria" status="sp. nov.">
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<emphasis id="B6D57A7DB907785EC5B846C4FE7D2004" bold="true" box="[151,404,1259,1286]" italics="true" pageId="20" pageNumber="21">Hydraena millenaria</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="ADE6C706B907785EC48C46C4FDDA2007" box="[419,563,1259,1285]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B907785EC5B84720FD7F202B" box="[151,662,1295,1322]" pageId="20" pageNumber="21" type="description">
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<paragraph id="841EA66FB907785EC5B84720FD7F202B" blockId="20.[151,662,1259,1322]" box="[151,662,1295,1322]" pageId="20" pageNumber="21">
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<figureCitation id="1C9ABAEAB907785EC5B84720FF0C202B" box="[151,229,1295,1321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="20" pageNumber="21">Figs. 4</figureCitation>
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,
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<figureCitation id="1C9ABAEAB907785EC5DE4720FEE42028" box="[241,269,1295,1322]" captionStart="FIGURE 25" captionStartId="50.[151,250,1991,2015]" captionTargetBox="[286,1303,192,1946]" captionTargetId="figure-25@50.[258,1329,181,1967]" captionTargetPageId="50" captionText="FIGURE 25. Hydraena millenaria, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284919" httpUri="https://zenodo.org/record/7284919/files/figure.png" pageId="20" pageNumber="21">25</figureCitation>
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(habitus), 25 (aedeagus), 40 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B907785EC5B84778FCD22097" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="841EA66FB907785EC5B84778FCD22097" blockId="20.[151,1437,1367,2042]" pageId="20" pageNumber="21">
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<emphasis id="B6D57A7DB907785EC5B84778FEA12073" bold="true" box="[151,328,1367,1393]" pageId="20" pageNumber="21">Type Material.</emphasis>
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<materialsCitation id="34C9AC32B907785EC4614778FEB82097" ID-GBIF-Occurrence="3962178350" collectingDate="1982-02-01" collectionCode="MCZ" collectorName="N. W. Prov. & P. D. Perkins" country="Cameroon" location="Bamenda" municipality="Skyline Hotel" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5B1A18CDB907785EC4614778FE5F2073" box="[334,438,1367,1393]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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(male): “
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<collectingCountry id="FCB6E6FFB907785EC7314778FD702073" box="[542,665,1367,1393]" name="Cameroon" pageId="20" pageNumber="21">Cameroon</collectingCountry>
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:
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<collectorName id="2954C3B9B907785EC78B4777FCC12070" box="[676,808,1368,1394]" pageId="20" pageNumber="21">N. W. Prov.</collectorName>
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,
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<location id="817EF0B4B907785EC61D4778FC762073" LSID="urn:lsid:plazi:treatment:0C081779B907785FC5B846C4FDD424AF:817EF0B4B907785EC61D4778FC762073" box="[818,927,1367,1393]" country="Cameroon" municipality="Skyline Hotel" name="Bamenda" pageId="20" pageNumber="21">Bamenda</location>
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, stream nr.
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<collectingMunicipality id="647A3C15B907785EC1314778FB522073" box="[1054,1211,1367,1393]" pageId="20" pageNumber="21">Skyline Hotel</collectingMunicipality>
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,
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<date id="F01F80AFB907785EC1E94778FAC52073" box="[1222,1324,1367,1393]" pageId="20" pageNumber="21" value="1982-02-01">
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<collectingDate id="E05B7947B907785EC1E94778FAC52073" box="[1222,1324,1367,1393]" pageId="20" pageNumber="21" value="1982-02-01">1.ii.1982</collectingDate>
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</date>
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, leg.
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<collectorName id="2954C3B9B907785EC0484777FF072097" pageId="20" pageNumber="21">P. D. Perkins</collectorName>
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” (
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<collectionCode id="E2B03EAAB907785EC4284754FEA32097" box="[263,330,1403,1429]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="20" pageNumber="21" type="Museum">MCZ</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="34C9AC32B907785EC4734753FCDE2097" ID-GBIF-Occurrence="3962178302" box="[348,823,1403,1430]" collectingDate="1982-02-01" collectionCode="MCZ" collectorName="N. W. Prov. & P. D. Perkins" country="Cameroon" location="Bamenda" municipality="Skyline Hotel" pageId="20" pageNumber="21" specimenCount="1" typeStatus="paratype">
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<typeStatus id="5B1A18CDB907785EC4734753FE2B2094" box="[348,450,1404,1430]" pageId="20" pageNumber="21" type="paratype">Paratype</typeStatus>
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: Same data as holotype (1
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<collectionCode id="E2B03EAAB907785EC7C24754FCC62097" box="[749,815,1403,1429]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="20" pageNumber="21" type="Museum">MCZ</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B907785EC5E847B0FDA8234B" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="841EA66FB907785EC5E847B0FDA8234B" blockId="20.[151,1437,1367,2042]" pageId="20" pageNumber="21">
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<emphasis id="B6D57A7DB907785EC5E847B0FE3820BB" bold="true" box="[199,465,1439,1465]" pageId="20" pageNumber="21">Differential Diagnosis.</emphasis>
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Differentiated from other members of the genus in
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<collectingCountry id="FCB6E6FFB907785EC12747B0FB9420BB" box="[1032,1149,1439,1465]" name="Cameroon" pageId="20" pageNumber="21">Cameroon</collectingCountry>
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by the combination of the absence of metaventral plaques, the frons and pronotum finely sparsely punctulate, the punctures of the pronotum very consistent in size throughout, and the aedeagus (
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<figureCitation id="1C9ABAEAB907785EC7C347C8FCA92300" box="[748,832,1511,1538]" captionStart="FIGURE 25" captionStartId="50.[151,250,1991,2015]" captionTargetBox="[286,1303,192,1946]" captionTargetId="figure-25@50.[258,1329,181,1967]" captionTargetPageId="50" captionText="FIGURE 25. Hydraena millenaria, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284919" httpUri="https://zenodo.org/record/7284919/files/figure.png" pageId="20" pageNumber="21">Fig. 25</figureCitation>
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). The dorsal punctation is much finer and sparser than that of
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<taxonomicName id="43A1DDECB907785EC5CA4423FE702327" authorityName="Perkins" authorityYear="2022" box="[229,409,1547,1573]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cameropetila">
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<emphasis id="B6D57A7DB907785EC5CA4423FE702327" box="[229,409,1547,1573]" italics="true" pageId="20" pageNumber="21">H. cameropetila</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="43A1DDECB907785EC4E14423FD612327" authorityName="Perkins" authorityYear="2022" box="[462,648,1548,1573]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="camerosetosa">
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<emphasis id="B6D57A7DB907785EC4E14423FD612327" box="[462,648,1548,1573]" italics="true" pageId="20" pageNumber="21">H. camerosetosa</emphasis>
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</taxonomicName>
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, two species which also lack metaventral plaques. All three species are in different Aedeagal Species Groups.
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B907785FC5E8447CFBFC2443" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
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<paragraph id="841EA66FB907785EC5E8447CFAF3226B" blockId="20.[151,1437,1367,2042]" pageId="20" pageNumber="21">
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<emphasis id="B6D57A7DB907785EC5E8447CFEB8236F" bold="true" box="[199,337,1619,1645]" pageId="20" pageNumber="21">Description</emphasis>
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. Size:
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<typeStatus id="5B1A18CDB907785EC4B3447CFE17236F" box="[412,510,1619,1645]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.20/0.53; head width 0.33; pronotum 0.31/0.44, PA 0.34, PB 0.39; elytra 0.75/0.53. Habitus as illustrated (
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<figureCitation id="1C9ABAEAB907785EC6CC4458FBDE2390" box="[995,1079,1655,1682]" captionStart="FIGURE 25" captionStartId="50.[151,250,1991,2015]" captionTargetBox="[286,1303,192,1946]" captionTargetId="figure-25@50.[258,1329,181,1967]" captionTargetPageId="50" captionText="FIGURE 25. Hydraena millenaria, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284919" httpUri="https://zenodo.org/record/7284919/files/figure.png" pageId="20" pageNumber="21">Fig. 25</figureCitation>
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). Dorsum brown to dark brown, head, pronotal fascia and elytra dark brown, pronotum rather widely bordered anteriorly and posteriorly slightly lighter brown, maxillary palpi testaceous, tip of last palpomere not darker. Frons and pronotum finely sparsely punctulate, punctures of pronotum slightly larger than those of frons, very consistent in size throughout, interstices shining, ca. 1–3xpd. Clypeus medially very finely sparsely punctulate, shining; laterally densely micropunctulate, dull. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral foveae deep, posterolateral and posterosubmedian foveae shallow.
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</paragraph>
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<paragraph id="841EA66FB907785EC5E8455CFBA922FB" blockId="20.[151,1437,1367,2042]" pageId="20" pageNumber="21">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals shining, not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
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<paragraph id="841EA66FB906785FC5B842B8FBFC2443" blockId="21.[151,1437,151,429]" pageId="21" pageNumber="22">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, surrounding areas smooth, shining. Genae raised, shining, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques absent. AIS width at straight posterior margin ca. 2x P2. All tibiae slender, straight. Abdominal apex slightly asymmetrical, with deep and wide apicomedian notch.</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B906785FC5E84365FC212467" box="[199,968,330,357]" pageId="21" pageNumber="22" type="distribution">
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<paragraph id="841EA66FB906785FC5E84365FC212467" blockId="21.[151,1437,151,429]" box="[199,968,330,357]" pageId="21" pageNumber="22">
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<emphasis id="B6D57A7DB906785FC5E84365FE882466" bold="true" box="[199,353,330,356]" pageId="21" pageNumber="22">Distribution.</emphasis>
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Currently known only from the
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<typeStatus id="5B1A18CDB906785FC7E34364FD152467" box="[716,764,331,357]" pageId="21" pageNumber="22">type</typeStatus>
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locality (
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<figureCitation id="1C9ABAEAB906785FC6484364FC522467" box="[871,955,331,357]" captionStart="FIGURES 39–41" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 39–41. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="21" pageNumber="22">Fig. 40</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="CCBBF5E4B906785FC5E84340FDD424AF" pageId="21" pageNumber="22" type="etymology">
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<paragraph id="841EA66FB906785FC5E84340FDD424AF" blockId="21.[151,1437,151,429]" pageId="21" pageNumber="22">
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<emphasis id="B6D57A7DB906785FC5E84340FEA7248B" bold="true" box="[199,334,367,393]" pageId="21" pageNumber="22">Etymology.</emphasis>
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Named in reference to the fact that the author’s goal of describing more than 1000 new species of water beetles is now happily attained.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |