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<document id="C72B0BEAB2D426FEA8A46568213EEB9D" ID-CLB-Dataset="181704" ID-DOI="10.11646/zootaxa.5203.1.1" ID-GBIF-Dataset="b6f94c7b-7033-4952-96a7-3db5fb6ad879" ID-ISSN="1175-5326" ID-Zenodo-Dep="7284841" ID-ZooBank="85C20298-5DF9-44DF-8485-3C64CF40CD08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667554441660" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2022" docId="0C081779B906785FC5B84069FB1223BF" docLanguage="en" docName="zootaxa.5203.1.1.pdf" docOrigin="Zootaxa 5203 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5203.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena camerocarinata Perkins 2022, new species" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="F0316F01B913784AC52F422FFFE92502" masterDocTitle="Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="22" updateTime="1699409684053" updateUser="ExternalLinkService">
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<mods:title id="5C1AE4524F1A1907A6F394F3B4F8EC4B">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="900C9DE7F28591C65F0C4AAB0AFD2CB7">Perkins, Philip D.</mods:namePart>
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<mods:title id="68C3F25EC1349E0B52C50A672446F0A4">Zootaxa</mods:title>
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<mods:date id="602710303BB2E3100EED529A94C0192F">2022</mods:date>
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<mods:number id="CA11C6DE08DB4861C271BBE0B7900C36">2022-11-03</mods:number>
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<mods:classification id="5EC640EF6992EB698609CCF1C08640C7">journal article</mods:classification>
<mods:identifier id="2BB27B4FDA33BF68B8C43EF18952F495" type="CLB-Dataset">181704</mods:identifier>
<mods:identifier id="849C84522945B810924A73D7BDA631E3" type="DOI">10.11646/zootaxa.5203.1.1</mods:identifier>
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<treatment id="0C081779B906785FC5B84069FB1223BF" ID-DOI="http://doi.org/10.5281/zenodo.7300273" ID-GBIF-Taxon="204205224" ID-Zenodo-Dep="7300273" LSID="urn:lsid:plazi:treatment:0C081779B906785FC5B84069FB1223BF" httpUri="http://treatment.plazi.org/id/0C081779B906785FC5B84069FB1223BF" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="CCBBF5E4B906785FC5B84069FD9D2762" box="[151,628,582,609]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="841EA66FB906785FC5B84069FD9D2762" blockId="21.[151,662,582,645]" box="[151,628,582,609]" pageId="21" pageNumber="22">
<heading id="DF561103B906785FC5B84069FD9D2762" bold="true" box="[151,628,582,609]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<emphasis id="B6D57A7DB906785FC5B84069FD9D2762" bold="true" box="[151,628,582,609]" pageId="21" pageNumber="22">
<taxonomicName id="43A1DDECB906785FC5B84069FE3F2763" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,470,582,609]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="camerocarinata" status="sp. nov.">
<emphasis id="B6D57A7DB906785FC5B84069FE3F2763" bold="true" box="[151,470,582,609]" italics="true" pageId="21" pageNumber="22">Hydraena camerocarinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B906785FC4CB4068FD9D2762" box="[484,628,582,609]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CCBBF5E4B906785FC5B84044FD7F2787" box="[151,662,619,645]" pageId="21" pageNumber="22" type="description">
<paragraph id="841EA66FB906785FC5B84044FD7F2787" blockId="21.[151,662,582,645]" box="[151,662,619,645]" pageId="21" pageNumber="22">
<figureCitation id="1C9ABAEAB906785FC5B84044FF0C2787" box="[151,229,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1956,1980]" captionTargetBox="[233,1355,489,1933]" captionTargetId="figure-134@6.[233,1355,489,1933]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Groups 3 and 4." figureDoi="http://doi.org/10.5281/zenodo.7284851" httpUri="https://zenodo.org/record/7284851/files/figure.png" pageId="21" pageNumber="22">Figs. 4</figureCitation>
,
<figureCitation id="1C9ABAEAB906785FC5DE4044FEE42787" box="[241,269,619,645]" captionStart="FIGURE 26" captionStartId="51.[151,250,1996,2020]" captionTargetBox="[247,1291,199,1954]" captionTargetId="figure-15@51.[211,1377,181,1973]" captionTargetPageId="51" captionText="FIGURE 26. Hydraena camerocarinata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284921" httpUri="https://zenodo.org/record/7284921/files/figure.png" pageId="21" pageNumber="22">26</figureCitation>
(habitus), 26 (aedeagus), 42 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5B8409CFF0727F3" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="841EA66FB906785FC5B8409CFF0727F3" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5B8409CFEA327CE" bold="true" box="[151,330,690,717]" pageId="21" pageNumber="22">Type Material.</emphasis>
<materialsCitation id="34C9AC32B906785FC47C409CFF0727F3" ID-GBIF-Occurrence="3962178307" collectingDate="1982-05-01" collectionCode="MCZ" collectorName="N. Ngaoundere &amp; P. D. Perkins" country="Cameroon" location="Cameroon" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB906785FC47C409CFE5227CF" box="[339,443,691,717]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B6D57A7DB906785FC730409DFDC427CE" bold="true" box="[543,557,690,716]" pageId="21" pageNumber="22"></emphasis>
<collectingCountry id="FCB6E6FFB906785FC702409CFD4B27CF" box="[557,674,691,717]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
:
<quantity id="43590B8AB906785FC79B409DFD1427CF" box="[692,765,690,717]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="km" value="35.0">35 km</quantity>
<collectorName id="2954C3B9B906785FC629409CFC5E27CF" box="[774,951,691,717]" pageId="21" pageNumber="22">N. Ngaoundere</collectorName>
, stream,
<date id="F01F80AFB906785FC130409CFB6827CF" box="[1055,1153,691,717]" pageId="21" pageNumber="22" value="1982-05-01">
<collectingDate id="E05B7947B906785FC130409CFB6827CF" box="[1055,1153,691,717]" pageId="21" pageNumber="22" value="1982-05-01">1.v.1982</collectingDate>
</date>
,
<collectorName id="2954C3B9B906785FC1A0409CFACC27CF" box="[1167,1317,691,717]" pageId="21" pageNumber="22">P. D. Perkins</collectorName>
collector” (
<collectionCode id="E2B03EAAB906785FC5B040F8FF0B27F3" box="[159,226,727,753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="21" pageNumber="22" type="Museum">MCZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E840D5FCC3265F" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="841EA66FB906785FC5E840D5FCC3265F" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E840D5FE382616" bold="true" box="[199,465,762,788]" pageId="21" pageNumber="22">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB906785FC12740D4FB942617" box="[1032,1149,763,789]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
by the combination of the nearly semicordate pronotum, the widely separated and carinate mesoventral plaques, and the aedeagus (
<figureCitation id="1C9ABAEAB906785FC0154130FA79263B" box="[1338,1424,799,825]" captionStart="FIGURE 26" captionStartId="51.[151,250,1996,2020]" captionTargetBox="[247,1291,199,1954]" captionTargetId="figure-15@51.[211,1377,181,1973]" captionTargetPageId="51" captionText="FIGURE 26. Hydraena camerocarinata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284921" httpUri="https://zenodo.org/record/7284921/files/figure.png" pageId="21" pageNumber="22">Fig. 26</figureCitation>
). Refer also to the diagnosis of
<taxonomicName id="43A1DDECB906785FC4C9416CFD0D265F" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[486,740,835,861]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="camerosemicordata" status="sp. nov.">
<emphasis id="B6D57A7DB906785FC4C9416CFD0D265F" box="[486,740,835,861]" italics="true" pageId="21" pageNumber="22">H. camerosemicordata</emphasis>
</taxonomicName>
<emphasis id="B6D57A7DB906785FC7C4416CFCC3265F" bold="true" box="[747,810,835,861]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="ADE6C706B906785FC7C4416CFCC3265F" box="[747,810,835,861]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E84149FE452377" pageId="21" pageNumber="22" type="description">
<paragraph id="841EA66FB906785FC5E84149FDA221A3" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E84149FEB82682" bold="true" box="[199,337,870,896]" pageId="21" pageNumber="22">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB906785FC48B4148FDEF2683" box="[420,518,871,897]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.35/0.57; head width 0.33; pronotum 0.35/0.44, PA 0.37, PB 0.33; elytra 0.81/0.57. Habitus as illustrated (
<figureCitation id="1C9ABAEAB906785FC12341A4FB8926A7" box="[1036,1120,907,933]" captionStart="FIGURE 26" captionStartId="51.[151,250,1996,2020]" captionTargetBox="[247,1291,199,1954]" captionTargetId="figure-15@51.[211,1377,181,1973]" captionTargetPageId="51" captionText="FIGURE 26. Hydraena camerocarinata, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284921" httpUri="https://zenodo.org/record/7284921/files/figure.png" pageId="21" pageNumber="22">Fig. 26</figureCitation>
). Pronotum almost semicordiform, anterior margin much wider than posterior margin. Dorsum brown except head and pronotal fascia black, palpi light brown to testaceous, tip of last palpomere not darker. Dorsum of head and pronotum moderately punctulate, more sparsely punctulate on fascia than on surrounding areas, interstices shining; punctures of frons disc fine and sparse, less than 1xef; punctures of pronotal disc ca. 2xef, interstices ca. 13xpd or slightly larger; punctures of clypeus fine, moderately dense. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; antero- and posterolateral pronotal foveae moderately deep, posterosubmedial foveae very shallow.
</paragraph>
<paragraph id="841EA66FB906785FC5E84684FD3E2057" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">Elytra with summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures 23xpd and deeper than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, very narrow, width ca. 1/2xpd or less, as are interstices between punctures of a row, each puncture with very short and very fine seta. Apices in dorsal aspect conjointly rounded, in posterior aspect margins not forming angle with one another.</paragraph>
<paragraph id="841EA66FB906785FC5B84770FE452377" blockId="21.[151,1437,690,1725]" pageId="21" pageNumber="22">Venter: Mentum and postmentum densely micropunctulate, dull. Genae very slightly raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.15/0.15/2. P1 laminate; median carina weakly sinuate in profile. P2 l/w ca. 2.5/1, sides almost parallel, apex blunt. Plaques very small, very narrow, carinate lines, straight, convergent anteriorly, very weakly raised, located at sides of deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS weakly concave, width at straight posterior margin ca. 2x P2. Protibiae slender, very weakly arcuate. Mesotibiae slender, straight. Metatibiae slender, straight, very slightly wider at midlength than apically. Abdominal apex symmetrical, with apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E84451FC21239B" box="[199,968,1662,1689]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="841EA66FB906785FC5E84451FC21239B" blockId="21.[151,1437,690,1725]" box="[199,968,1662,1689]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E84451FE88239A" bold="true" box="[199,353,1662,1688]" pageId="21" pageNumber="22">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB906785FC7E34450FD15239B" box="[716,764,1663,1689]" pageId="21" pageNumber="22">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB906785FC6484450FC52239B" box="[871,955,1663,1689]" captionStart="FIGURES 4243" captionStartId="65.[151,264,1727,1751]" captionTargetBox="[156,1432,181,1701]" captionTargetId="figure-14@65.[156,1432,181,1704]" captionTargetPageId="65" captionText="FIGURES 4243. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284967" httpUri="https://zenodo.org/record/7284967/files/figure.png" pageId="21" pageNumber="22">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B906785FC5E8448CFB1223BF" box="[199,1275,1699,1725]" pageId="21" pageNumber="22" type="etymology">
<paragraph id="841EA66FB906785FC5E8448CFB1223BF" blockId="21.[151,1437,690,1725]" box="[199,1275,1699,1725]" pageId="21" pageNumber="22">
<emphasis id="B6D57A7DB906785FC5E8448CFEA723BF" bold="true" box="[199,334,1699,1725]" pageId="21" pageNumber="22">Etymology.</emphasis>
Named in reference to the known distribution and the carinate metaventral plaques.
</paragraph>
</subSubSection>
</treatment>
</document>