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<document id="7A57197C8AFF22B7D7781E91550BE77B" ID-CLB-Dataset="181704" ID-DOI="10.11646/zootaxa.5203.1.1" ID-GBIF-Dataset="b6f94c7b-7033-4952-96a7-3db5fb6ad879" ID-ISSN="1175-5326" ID-Zenodo-Dep="7284841" ID-ZooBank="85C20298-5DF9-44DF-8485-3C64CF40CD08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667554441660" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2022" docId="0C081779B905785DC5B84715FBD6271B" docLanguage="en" docName="zootaxa.5203.1.1.pdf" docOrigin="Zootaxa 5203 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5203.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena cameroarida Perkins 2022, new species" docType="treatment" docVersion="7" lastPageNumber="24" masterDocId="F0316F01B913784AC52F422FFFE92502" masterDocTitle="Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="23" updateTime="1699409684053" updateUser="ExternalLinkService">
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<mods:title id="F0B686ECFF32E6E83013941B77BE8EF0">Thirty-three new species of water beetles in the genus Hydraena Kugelann from Cameroon (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="670351676DE9DDDC303689EADF6FAEDC">Perkins, Philip D.</mods:namePart>
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<mods:date id="06588E7BDEF25DC91B98F7B80ABB4048">2022</mods:date>
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<mods:number id="0D7FF64524929C2843B7B9BBDA6FD8BF">2022-11-03</mods:number>
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<mods:identifier id="C66FA6684549408CC7E9C9A1962C26F9" type="DOI">10.11646/zootaxa.5203.1.1</mods:identifier>
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<treatment id="0C081779B905785DC5B84715FBD6271B" ID-DOI="http://doi.org/10.5281/zenodo.7300206" ID-GBIF-Taxon="204205226" ID-Zenodo-Dep="7300206" LSID="urn:lsid:plazi:treatment:0C081779B905785DC5B84715FBD6271B" httpUri="http://treatment.plazi.org/id/0C081779B905785DC5B84715FBD6271B" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection id="CCBBF5E4B905785CC5B84715FDA62056" box="[151,591,1338,1365]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="841EA66FB905785CC5B84715FDA62056" blockId="22.[151,662,1338,1401]" box="[151,591,1338,1365]" pageId="22" pageNumber="23">
<heading id="DF561103B905785CC5B84715FDA62056" bold="true" box="[151,591,1338,1365]" fontSize="11" level="2" pageId="22" pageNumber="23" reason="4">
<emphasis id="B6D57A7DB905785CC5B84715FDA62056" bold="true" box="[151,591,1338,1365]" pageId="22" pageNumber="23">
<taxonomicName id="43A1DDECB905785CC5B84715FE592057" authority="Perkins, 2022" authorityName="Perkins" authorityYear="2022" box="[151,432,1338,1365]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cameroarida" status="sp. nov.">
<emphasis id="B6D57A7DB905785CC5B84715FE592057" bold="true" box="[151,432,1338,1365]" italics="true" pageId="22" pageNumber="23">Hydraena cameroarida</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADE6C706B905785CC4904715FDA62056" box="[447,591,1338,1364]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CCBBF5E4B905785CC5B84771FD7F207A" box="[151,662,1374,1401]" pageId="22" pageNumber="23" type="description">
<paragraph id="841EA66FB905785CC5B84771FD7F207A" blockId="22.[151,662,1338,1401]" box="[151,662,1374,1401]" pageId="22" pageNumber="23">
<figureCitation id="1C9ABAEAB905785CC5B84771FF0C207B" box="[151,229,1374,1401]" captionStart="FIGURE 5" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[162,1409,281,1916]" captionTargetId="figure-52@7.[151,1435,273,1927]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal habitus of Cameroonian species of Hydraena in Aedeagal Species Group 5." figureDoi="http://doi.org/10.5281/zenodo.7284853" httpUri="https://zenodo.org/record/7284853/files/figure.png" pageId="22" pageNumber="23">Figs. 5</figureCitation>
,
<figureCitation id="1C9ABAEAB905785CC5DE4771FEE4207A" box="[241,269,1374,1400]" captionStart="FIGURE 28" captionStartId="53.[151,250,1970,1994]" captionTargetBox="[305,1282,181,1946]" captionTargetId="figure-15@53.[305,1282,181,1946]" captionTargetPageId="53" captionText="FIGURE 28. Hydraena cameroarida, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284927" httpUri="https://zenodo.org/record/7284927/files/figure.png" pageId="22" pageNumber="23">28</figureCitation>
(habitus), 28 (aedeagus), 39 (map)
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785CC5B84789FEBC20E6" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="841EA66FB905785CC5B84789FEBC20E6" blockId="22.[151,1437,1446,2013]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5B84789FEA220C2" bold="true" box="[151,331,1446,1472]" pageId="22" pageNumber="23">Type Material.</emphasis>
<materialsCitation id="34C9AC32B905785CC47A4789FEBC20E6" ID-GBIF-Occurrence="3962178313" collectingDate="1982-01-14" collectionCode="MCZ" collectorName="P. D. Perkins" country="Cameroon" location="Cameroon" pageId="22" pageNumber="23" specimenCount="1" typeStatus="holotype">
<typeStatus id="5B1A18CDB905785CC47A4789FE5420C2" box="[341,445,1446,1472]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="FCB6E6FFB905785CC7034789FD4E20C2" box="[556,679,1446,1472]" name="Cameroon" pageId="22" pageNumber="23">Cameroon</collectingCountry>
:
<collectorName id="2954C3B9B905785CC79A4788FCFE20C3" box="[693,791,1447,1473]" pageId="22" pageNumber="23">N. Prov.</collectorName>
,
<quantity id="43590B8AB905785CC60A4789FC9B20C2" box="[805,882,1446,1472]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="23" unit="km" value="22.0">22 km</quantity>
.
<collectorName id="2954C3B9B905785CC6AF4788FC1620C3" box="[896,1023,1447,1473]" pageId="22" pageNumber="23">N. Maroua</collectorName>
, desert stream,
<date id="F01F80AFB905785CC1994789FAC820C2" box="[1206,1313,1446,1472]" pageId="22" pageNumber="23" value="1982-01-14">
<collectingDate id="E05B7947B905785CC1994789FAC820C2" box="[1206,1313,1446,1472]" pageId="22" pageNumber="23" value="1982-01-14">14.i.1982</collectingDate>
</date>
, leg.
<collectorName id="2954C3B9B905785CC04B4788FF0720E6" pageId="22" pageNumber="23">P. D. Perkins</collectorName>
” (
<collectionCode id="E2B03EAAB905785CC42847E5FEA320E6" box="[263,330,1482,1508]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="22" pageNumber="23" type="Museum">MCZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785CC5E847C1FE402376" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="841EA66FB905785CC5E847C1FE402376" blockId="22.[151,1437,1446,2013]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5E847C1FE38230A" bold="true" box="[199,465,1518,1544]" pageId="22" pageNumber="23">Differential Diagnosis.</emphasis>
Differentiated from other members of the genus in
<collectingCountry id="FCB6E6FFB905785CC12747C1FB94230A" box="[1032,1149,1518,1544]" name="Cameroon" pageId="22" pageNumber="23">Cameroon</collectingCountry>
by the combination of the brown to dark brown or black dorsum, the deep anterolateral pronotal foveae, and the aedeagus (
<figureCitation id="1C9ABAEAB905785CC1CF443DFADC232E" box="[1248,1333,1554,1580]" captionStart="FIGURE 28" captionStartId="53.[151,250,1970,1994]" captionTargetBox="[305,1282,181,1946]" captionTargetId="figure-15@53.[305,1282,181,1946]" captionTargetPageId="53" captionText="FIGURE 28. Hydraena cameroarida, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284927" httpUri="https://zenodo.org/record/7284927/files/figure.png" pageId="22" pageNumber="23">Fig. 28</figureCitation>
). Similar in habitus and size to
<taxonomicName id="43A1DDECB905785CC4A34418FD8D2352" authorityName="Perkins" authorityYear="2022" box="[396,612,1591,1616]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="camerocontinua">
<emphasis id="B6D57A7DB905785CC4A34418FD8D2352" box="[396,612,1591,1616]" italics="true" pageId="22" pageNumber="23">H. camerocontinua</emphasis>
</taxonomicName>
, but differing therefrom in denser dorsal punctation, ventral morphology, and aedeagal characters.
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B905785DC5E84451FC1624D3" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="841EA66FB905785CC5E84451FB0F2296" blockId="22.[151,1437,1446,2013]" pageId="22" pageNumber="23">
<emphasis id="B6D57A7DB905785CC5E84451FEB8239A" bold="true" box="[199,337,1662,1688]" pageId="22" pageNumber="23">Description</emphasis>
. Size:
<typeStatus id="5B1A18CDB905785CC48B4451FDEF239A" box="[420,518,1662,1688]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.37/0.57; head width 0.34; pronotum 0.32/0.45, PA 0.38, PB 0.40; elytra 0.80/0.57. Habitus as illustrated (
<figureCitation id="1C9ABAEAB905785CC106448DFB6923BE" box="[1065,1152,1698,1724]" captionStart="FIGURE 28" captionStartId="53.[151,250,1970,1994]" captionTargetBox="[305,1282,181,1946]" captionTargetId="figure-15@53.[305,1282,181,1946]" captionTargetPageId="53" captionText="FIGURE 28. Hydraena cameroarida, new species, habitus and aedeagus of holotype." figureDoi="http://doi.org/10.5281/zenodo.7284927" httpUri="https://zenodo.org/record/7284927/files/figure.png" pageId="22" pageNumber="23">Fig. 28</figureCitation>
). Dorsum brown to dark brown or black, frons and pronotal disc darker, maxillary palpi light brown, tip of last palpomere not darker. Frons finely moderately densely punctulate, ca. 1xef, interstices ca. 12xpd, interstices shining. Pronotum moderately coarsely moderately densely punctate, ca. 2xpd of frons punctures; interstices ca. 1xpd or slightly greater, shining. Clypeus medially very finely sparsely punctulate, shining; laterally very finely densely punctulate, dull. Labrum apicomedially excised, free margins not upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal foveae deep, posterolateral shallow, posterosubmedial moderately deep.
</paragraph>
<paragraph id="841EA66FB905785DC5E845B1FEB325FB" blockId="22.[151,1437,1446,2013]" lastBlockId="23.[151,1437,151,537]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">Elytra with summit of posterior declivity at or very near posterior 1/3; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior, each puncture with very short and very fine seta. Intervals not raised, width ca. 12xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very weak angle with one another.</paragraph>
<paragraph id="841EA66FB904785DC5B8432CFC1624D3" blockId="23.[151,1437,151,537]" pageId="23" pageNumber="24">Venter: Mentum sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate, dull. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques narrow, straight or very slightly arcuate, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 1.5x P2. Protibia very slender, straight. All tibiae slender, straight. Abdominal apex symmetrical, with deep apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E843F5FC2124F7" box="[199,968,474,501]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="841EA66FB904785DC5E843F5FC2124F7" blockId="23.[151,1437,151,537]" box="[199,968,474,501]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E843F5FE8824F6" bold="true" box="[199,353,474,500]" pageId="23" pageNumber="24">Distribution.</emphasis>
Currently known only from the
<typeStatus id="5B1A18CDB904785DC7E343F4FD1524F7" box="[716,764,475,501]" pageId="23" pageNumber="24">type</typeStatus>
locality (
<figureCitation id="1C9ABAEAB904785DC64843F4FC5224F7" box="[871,955,475,501]" captionStart="FIGURES 3941" captionStartId="64.[151,264,1911,1935]" captionTargetBox="[167,1420,182,1885]" captionTargetId="figure-25@64.[167,1420,181,1888]" captionTargetPageId="64" captionText="FIGURES 3941. Geographical distributions of Cameroonian Hydraena." figureDoi="http://doi.org/10.5281/zenodo.7284965" httpUri="https://zenodo.org/record/7284965/files/figure.png" pageId="23" pageNumber="24">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCBBF5E4B904785DC5E843D0FBD6271B" box="[199,1087,511,537]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="841EA66FB904785DC5E843D0FBD6271B" blockId="23.[151,1437,151,537]" box="[199,1087,511,537]" pageId="23" pageNumber="24">
<emphasis id="B6D57A7DB904785DC5E843D0FEA7271B" bold="true" box="[199,334,511,537]" pageId="23" pageNumber="24">Etymology.</emphasis>
Named in reference to the known distribution, and to the arid area.
</paragraph>
</subSubSection>
</treatment>
</document>