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<document id="202D33E40F1E159FE462E8F077A97864" ID-DOI="10.5281/zenodo.189744" ID-GBIF-Dataset="6a2f3119-e07a-40c1-9c23-c05b3bfb163b" ID-ISSN="1175-5326" ID-Zenodo-Dep="189744" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461104746989" checkinUser="plazi" docAuthor="Reid, Chris A. M., Jurado-Rivera, José A. &amp; Beatson, Max" docDate="2009" docId="011387B4FF80AC0EFF4BFF23C6120FAB" docLanguage="en" docName="zt02207p066.pdf" docOrigin="Zootaxa 2207" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Canobolas Reid, Jurado-Rivera &amp; Beatson, gen. nov." docType="treatment" docVersion="8" lastPageNumber="56" masterDocId="FD2AFFCCFF81AC0DFFDCFFB4C70F0B34" masterDocTitle="A new genus of Chrysomelinae from Australia (Coleoptera: Chrysomelidae)" masterLastPageNumber="66" masterPageNumber="53" pageNumber="54" updateTime="1698592537263" updateUser="plazi">
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<mods:titleInfo id="FB92D51BF649E552B23A4D2080C5E014">
<mods:title id="E4FE3D3C173E9DD215AB91F6ADEF3D77">A new genus of Chrysomelinae from Australia (Coleoptera: Chrysomelidae)</mods:title>
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<mods:name id="947C1C330660370EA5AAF98D98310DB2" type="personal">
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<mods:namePart id="1F7FDA5AA6E3554BFF396F19315785A0">Reid, Chris A. M.</mods:namePart>
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<mods:namePart id="CD171F14AB5EAB81419ECF49A297A7B9">Jurado-Rivera, José A.</mods:namePart>
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<mods:namePart id="14CF74011879EBAE8005A1224E236B3B">Beatson, Max</mods:namePart>
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<mods:title id="80216A575817C1BC55B1C4338D592EE2">Zootaxa</mods:title>
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<mods:date id="07C39329CF0784993CB9A91FC918B2F7">2009</mods:date>
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<mods:identifier id="2D9D77FABFC3F297B82FB7D131B491B8" type="DOI">10.5281/zenodo.189744</mods:identifier>
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<treatment id="011387B4FF80AC0EFF4BFF23C6120FAB" ID-DOI="http://doi.org/10.5281/zenodo.6214685" ID-GBIF-Taxon="119622895" ID-Zenodo-Dep="6214685" LSID="urn:lsid:plazi:treatment:011387B4FF80AC0EFF4BFF23C6120FAB" httpUri="http://treatment.plazi.org/id/011387B4FF80AC0EFF4BFF23C6120FAB" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54">
<subSubSection id="C1A06529FF80AC0CFF4BFF23C4290B85" box="[151,806,151,177]" pageId="1" pageNumber="54" type="nomenclature">
<paragraph id="890536A2FF80AC0CFF4BFF23C4290B85" blockId="1.[151,806,151,177]" box="[151,806,151,177]" pageId="1" pageNumber="54">
<heading id="D24D81CEFF80AC0CFF4BFF23C4290B85" bold="true" box="[151,806,151,177]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
<emphasis id="BBCEEAB0FF80AC0CFF4BFF23C4290B85" bold="true" box="[151,806,151,177]" pageId="1" pageNumber="54">
<taxonomicName id="4EBA4D21FF80AC0CFF4BFF23C5A30B85" authority="Reid, Jurado-Rivera &amp; Beatson" authorityName="Reid, Jurado-Rivera &amp; Beatson" box="[151,684,151,177]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBCEEAB0FF80AC0CFF4BFF23C6170B85" bold="true" box="[151,280,151,177]" italics="true" pageId="1" pageNumber="54">Canobolas</emphasis>
Reid, Jurado-Rivera &amp; Beatson
</taxonomicName>
,
<taxonomicNameLabel id="A0FD57CBFF80AC0CFD65FF23C4290B85" box="[697,806,151,177]" pageId="1" pageNumber="54" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C1A06529FF80AC0CFF4BFF55C4040BCC" box="[151,779,225,248]" pageId="1" pageNumber="54" type="materials_examined">
<paragraph id="890536A2FF80AC0CFF4BFF55C4040BCC" blockId="1.[151,779,225,248]" box="[151,779,225,248]" pageId="1" pageNumber="54">
<typeStatus id="56018800FF80AC0CFF4BFF55C7C30BCC" box="[151,204,225,248]" pageId="1" pageNumber="54">Type</typeStatus>
species.
<taxonomicName id="4EBA4D21FF80AC0CFEF0FF55C55F0BCC" authority="Lea, 1917" authorityName="Lea" authorityYear="1917" box="[300,592,225,248]" class="Insecta" family="Chrysomelidae" genus="Geomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="BBCEEAB0FF80AC0CFEF0FF55C6D30BCC" box="[300,476,225,248]" italics="true" pageId="1" pageNumber="54">Geomela nobilis</emphasis>
<bibRefCitation id="ED2B4B53FF80AC0CFE3FFF55C55F0BCC" author="Lea" box="[483,592,225,248]" pageId="1" pageNumber="66" refString="Lea, A. M. (1917) Descriptions of new species of Australian Coleoptera. Part xiii. Proceedings of the Linnean Society of New South Wales, 42 (3), 545 - 582, plate 29." type="journal article" year="1917">Lea, 1917</bibRefCitation>
</taxonomicName>
, this designation.
</paragraph>
</subSubSection>
<subSubSection id="C1A06529FF80AC0FFF4BFE93C4D30AF4" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54" type="description">
<paragraph id="890536A2FF80AC0CFF4BFE93C6740A82" blockId="1.[151,1437,295,2026]" pageId="1" pageNumber="54">
<emphasis id="BBCEEAB0FF80AC0CFF4BFE93C6820A75" bold="true" box="[151,397,295,321]" pageId="1" pageNumber="54">Description (adult).</emphasis>
Length 3.56.0mm; body (
<figureCitation id="11812A27FF80AC0CFD0FFE9CC4330A76" box="[723,828,296,322]" captionStart-0="FIGURES 1 4" captionStart-1="FIGURES 5 8" captionStartId-0="2.[151,269,2010,2034]" captionStartId-1="5.[151,269,1818,1842]" captionTargetBox-0="[306,1230,705,1953]" captionTargetBox-1="[185,1397,202,1767]" captionTargetId-0="figure@2.[280,1307,661,1984]" captionTargetId-1="figure@5.[151,1436,169,1794]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionText-0="FIGURES 1 4. Canobolas nobilis (Lea), male: 1, dorsal; 2, lateral; 3, ventral; 4, face." captionText-1="FIGURES 5 8. Canobolas species, dorsal: 5, C. jarrah sp. nov.; 6, C. minang sp. nov.; 7, C. tubrabucca sp. nov.; face: 8, C. jarrah sp. nov." httpUri-0="https://zenodo.org/record/189745/files/figure.png" httpUri-1="https://zenodo.org/record/189746/files/figure.png" pageId="1" pageNumber="54">Figs 18</figureCitation>
) almost circular to elongate-ovate, length 1.11.7x width, moderately to strongly convex in profile, length 1.72.4x height. Dorsum dark reddish-brown to black, with or without slight metallic green or blue reflections, glabrous except a few recumbent setae above antennal insertions.
</paragraph>
<paragraph id="890536A2FF80AC0CFF19FE76C38408E7" blockId="1.[151,1437,295,2026]" pageId="1" pageNumber="54">
Head (
<figureCitation id="11812A27FF80AC0CFECAFE76C6670AE8" box="[278,360,450,476]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,2010,2034]" captionTargetBox="[306,1230,705,1953]" captionTargetId="figure@2.[280,1307,661,1984]" captionTargetPageId="2" captionText="FIGURES 1 4. Canobolas nobilis (Lea), male: 1, dorsal; 2, lateral; 3, ventral; 4, face." httpUri="https://zenodo.org/record/189745/files/figure.png" pageId="1" pageNumber="54">Figs 4</figureCitation>
,
<figureCitation id="11812A27FF80AC0CFEAAFE76C6880AE8" box="[374,391,450,476]" captionStart="FIGURES 5 8" captionStartId="5.[151,269,1818,1842]" captionTargetBox="[185,1397,202,1767]" captionTargetId="figure@5.[151,1436,169,1794]" captionTargetPageId="5" captionText="FIGURES 5 8. Canobolas species, dorsal: 5, C. jarrah sp. nov.; 6, C. minang sp. nov.; 7, C. tubrabucca sp. nov.; face: 8, C. jarrah sp. nov." httpUri="https://zenodo.org/record/189746/files/figure.png" pageId="1" pageNumber="54">8</figureCitation>
,
<figureCitation id="11812A27FF80AC0CFE4AFE76C6A80AE8" box="[406,423,450,476]" captionStart="FIGURES 9 10" captionStartId="6.[151,269,1958,1982]" captionTargetBox="[188,1402,1267,1957]" captionTargetId="figure@6.[151,1436,1225,1964]" captionTargetPageId="6" captionText="FIGURES 9 10. Canobolas nobilis (Lea): 9, venter of head and pronotum; 10, venter of ovipositor." httpUri="https://zenodo.org/record/189747/files/figure.png" pageId="1" pageNumber="54">9</figureCitation>
,
<figureCitation id="11812A27FF80AC0CFE69FE76C50A0AE8" box="[437,517,450,476]" captionStart="FIGURES 11 14" captionStartId="7.[151,269,1948,1972]" captionTargetBox="[271,1265,691,1902]" captionTargetId="figure@7.[203,1384,671,1942]" captionTargetPageId="7" captionText="FIGURES 11 14. Canobolas species, male antenna (drawn to scale): 11, C. jarrah sp. nov.; 12, C. minang sp. nov.; 13, C. nobilis (Lea); 14, C. tubrabucca sp. nov." httpUri="https://zenodo.org/record/189748/files/figure.png" pageId="1" pageNumber="54">1114</figureCitation>
): fronto-clypeus smooth, without antero-lateral ridges and with an arcuate groove or shallow depression between (but not reaching) bases of antennae, which are adjacent to eyes; vertex obliquely grooved posteriorly from inner margin of each eye (groove often hidden by overlapping prothorax); eyes strongly transverse (narrow in dorsal view), height c.
<date id="FD041062FF80AC0CFC96FD82C4670964" box="[842,872,566,592]" pageId="1" pageNumber="54">3x</date>
width, partially ventral, inner margin shallowly concave; eye laterally projecting, anteriorly truncate and posteriorly abruptly constricted to parallel-sided head capsule, temples absent or almost so; antennae situated on anterior margin of head; venter of head with deep and smooth elongate subantennal groove, length c.
<date id="FD041062FF80AC0CFCF1FD1EC44509F0" box="[813,842,682,708]" pageId="1" pageNumber="54">3x</date>
width, bounded by ridges, for retention of antenna between eye and mouth; antennae c. 0.30.6x body length, antennomere 1 enlarged, ovate, 2 short (half length of 1), 3 relatively narrow and elongated (length 2.53.5x width), 6 reduced, shorter than either 5 or 7, 710 expanded from base to apex, length 0.91.7x width; labrum not densely setose, with 35 pairs of prominent setae; mandible with two apical teeth and thick membranous pad (pulvillus) near middle of inner margin; apical maxillary palpomere elongate, cylindrical or fusiform, preapical palpomere triangular, shorter than apical and of similar width; first maxillary palpomere without groove or ridge to receive second palpomere; mentum transverse, width c.
<date id="FD041062FF80AC0CFE35FC0DC50808E7" box="[489,519,953,979]" pageId="1" pageNumber="54">3x</date>
median length, but anterior angles narrowly produced.
</paragraph>
<paragraph id="890536A2FF80AC0CFF19FC54C3290C7B" blockId="1.[151,1437,295,2026]" pageId="1" pageNumber="54">
Thorax
<emphasis id="BBCEEAB0FF80AC0CFEFEFC54C62308CD" box="[290,300,992,1017]" italics="true" pageId="1" pageNumber="54">(</emphasis>
<figureCitation id="11812A27FF80AC0CFEF0FC54C69B08CE" box="[300,404,992,1018]" captionStart-0="FIGURES 1 4" captionStart-1="FIGURES 5 8" captionStart-2="FIGURES 9 10" captionStartId-0="2.[151,269,2010,2034]" captionStartId-1="5.[151,269,1818,1842]" captionStartId-2="6.[151,269,1958,1982]" captionTargetBox-0="[306,1230,705,1953]" captionTargetBox-1="[185,1397,202,1767]" captionTargetBox-2="[188,1402,1267,1957]" captionTargetId-0="figure@2.[280,1307,661,1984]" captionTargetId-1="figure@5.[151,1436,169,1794]" captionTargetId-2="figure@6.[151,1436,1225,1964]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 1 4. Canobolas nobilis (Lea), male: 1, dorsal; 2, lateral; 3, ventral; 4, face." captionText-1="FIGURES 5 8. Canobolas species, dorsal: 5, C. jarrah sp. nov.; 6, C. minang sp. nov.; 7, C. tubrabucca sp. nov.; face: 8, C. jarrah sp. nov." captionText-2="FIGURES 9 10. Canobolas nobilis (Lea): 9, venter of head and pronotum; 10, venter of ovipositor." httpUri-0="https://zenodo.org/record/189745/files/figure.png" httpUri-1="https://zenodo.org/record/189746/files/figure.png" httpUri-2="https://zenodo.org/record/189747/files/figure.png" pageId="1" pageNumber="54">Figs 19</figureCitation>
,
<figureCitation id="11812A27FF80AC0CFE7EFC54C6FF08CE" box="[418,496,992,1018]" captionStart="FIGURES 15 18" captionStartId="8.[151,270,1723,1747]" captionTargetBox="[255,1290,196,1679]" captionTargetId="figure@8.[235,1351,176,1717]" captionTargetPageId="8" captionText="FIGURES 15 18. Canobolas species, partial right side of venter, showing metaventrite, metacoxa and first abdominal ventrite, femoral plates shaded (drawn to scale): 15, C. jarrah sp. nov.; 16, C. minang sp. nov.; 17, C. nobilis (Lea); 18, C. tubrabucca sp. nov." httpUri="https://zenodo.org/record/189749/files/figure.png" pageId="1" pageNumber="54">1518</figureCitation>
): pronotum transverse, broadest at base, evenly convex in cross section or with feeble lateral depressions; anterior edge concave, posterior convex and lateral edges slightly convex; anterior and sides distinctly margined, base without margination; anterior angles rounded, posterior angles approximately 90°; pronotum without trichobothria, angles non-setose; hypomeron impunctate, without groove; prosternum with strong oblique ridge anterior to each coxa, for retention of antennae; prosternal process about as broad as long, or slightly broader, truncate, evenly expanded to apex, with lateral ridges converging anteriorly at or just before anterior margin of prosternum; procoxal cavities open, gap at least half width of procoxa; scutellum with rounded apex; elytra evenly convex, without basal or lateral depressions, with scutellary striole and 8 complete regular striae, 9th present at least in basal third, 10th present as series of large punctures above epipleural margin; elytral epipleuron narrow, width &lt;0.2x elytral width, entirely visible laterally, slightly sinuate, gradually contracted from base to apex, and without setae; anterior elytral wingbinding patch on oval swelling one third from base; mesoventrite visible between prosternal process and metaventrite, medially reduced to short strongly transverse (width&gt;
<date id="FD041062FF80AC0CFC73FA06C4C30EF8" box="[943,972,1458,1484]" pageId="1" pageNumber="54">3x</date>
length) median process, with shallowly concave apex, and laterally reduced to a thin arcuate ridge between each mesocoxa and prosternal process; anterior of mesoventrite process abruptly declivous to accommodate prosternal process; wings entirely absent; metaventrite evenly microreticulate, strongly transverse, width 3
<date id="FD041062FF80AC0CFC45F993C4A70D75" box="[921,936,1575,1601]" pageId="1" pageNumber="54">4</date>
x length, with deep median depression and anterolateral margins flattened and expanded as smooth areas (femoral plates); metepisternum smooth, not longitudinally grooved; metendosternite with short basal stalk and thin recurved lobes at apices of lateral arms; all femora flattened, slightly fusiform to almost parallel-sided, ventral surfaces defined by paired longitudinal ridges; all tibiae flattened, with one dorsal and two ventral keels, expanded from narrow base to beyond middle, then dorsally excavate before apex to accommodate first tarsomere; all first tarsomeres with ovate patch of spatulate setae in males, with dense non-spatulate setae diverging from midline in females; apex of third tarsomere shallowly concave; claws simple, without basal tooth.
</paragraph>
<paragraph id="890536A2FF80AC0FFF19F8E8C4D30AF4" blockId="1.[151,1437,295,2026]" lastBlockId="2.[151,1437,152,642]" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
Abdomen (
<figureCitation id="11812A27FF80AC0CFE8DF8E8C6AC0C42" box="[337,419,1884,1910]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,2010,2034]" captionTargetBox="[306,1230,705,1953]" captionTargetId="figure@2.[280,1307,661,1984]" captionTargetPageId="2" captionText="FIGURES 1 4. Canobolas nobilis (Lea), male: 1, dorsal; 2, lateral; 3, ventral; 4, face." httpUri="https://zenodo.org/record/189745/files/figure.png" pageId="1" pageNumber="54">Figs 3</figureCitation>
,
<figureCitation id="11812A27FF80AC0CFE6EF8E8C6DB0C42" box="[434,468,1884,1910]" captionStart="FIGURES 9 10" captionStartId="6.[151,269,1958,1982]" captionTargetBox="[188,1402,1267,1957]" captionTargetId="figure@6.[151,1436,1225,1964]" captionTargetPageId="6" captionText="FIGURES 9 10. Canobolas nobilis (Lea): 9, venter of head and pronotum; 10, venter of ovipositor." httpUri="https://zenodo.org/record/189747/files/figure.png" pageId="1" pageNumber="54">10</figureCitation>
,
<figureCitation id="11812A27FF80AC0CFE3FF8E8C5380C42" box="[483,567,1884,1910]" captionStart-0="FIGURES 15 18" captionStart-1="FIGURES 19 22" captionStart-2="FIGURES 23 24" captionStartId-0="8.[151,270,1723,1747]" captionStartId-1="9.[151,269,1911,1935]" captionStartId-2="10.[151,269,1305,1329]" captionTargetBox-0="[255,1290,196,1679]" captionTargetBox-1="[193,1389,633,1870]" captionTargetBox-2="[152,1386,204,1304]" captionTargetId-0="figure@8.[235,1351,176,1717]" captionTargetId-1="figure@9.[151,1436,578,1947]" captionTargetId-2="figure@10.[151,1436,168,1311]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 15 18. Canobolas species, partial right side of venter, showing metaventrite, metacoxa and first abdominal ventrite, femoral plates shaded (drawn to scale): 15, C. jarrah sp. nov.; 16, C. minang sp. nov.; 17, C. nobilis (Lea); 18, C. tubrabucca sp. nov." captionText-1="FIGURES 19 22. Canobolas species, male genitalia (drawn to scale): 19, C. jarrah sp. nov.; 20, C. minang sp. nov.; 21," captionText-2="FIGURES 23 24. Canobolas species, female genitalia (drawn to scale): 23, C. jarrah sp. nov.; 24, C. nobilis (Lea)." httpUri-0="https://zenodo.org/record/189749/files/figure.png" httpUri-1="https://zenodo.org/record/189750/files/figure.png" httpUri-2="https://zenodo.org/record/189751/files/figure.png" pageId="1" pageNumber="54">1524</figureCitation>
): pygidium deeply grooved from base to apex along midline, groove rectangular in cross-section, gradually narrowed from base to apex; tergite VIII with distinct spiracles; ventrites free, not fused, ventral surfaces of I and V laterally bound by elevated ridges; ventrite I with intercoxal process truncate at apex, anterior border flattened and expanded behind coxae (femoral plates); ventrite V smooth, not medially depressed, and truncate at apex in both sexes, but male with transverse rectangular internal flange; sternite IX of male reduced to two separate struts; male tegmen present, V-shaped; penis simple, flattened tubular, with small basal foramen, &lt;0.3x length of penis; base of penis with longitudinal split; female tergite 8 well-developed, sternite 8 with long, thin, basal apodeme (
<taxonomicName id="4EBA4D21FF83AC0FFB20FEB8C27E0A11" box="[1276,1393,268,293]" class="Insecta" family="Chrysomelidae" genus="Geomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="BBCEEAB0FF83AC0FFB20FEB8C27E0A11" box="[1276,1393,268,293]" italics="true" pageId="2" pageNumber="55">C. nobilis</emphasis>
</taxonomicName>
) or with short bifurcate apodeme (
<taxonomicName id="4EBA4D21FF83AC0FFDDDFE86C5600A7F" box="[513,623,306,331]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="jarrah">
<emphasis id="BBCEEAB0FF83AC0FFDDDFE86C5600A7F" box="[513,623,306,331]" italics="true" pageId="2" pageNumber="55">C. jarrah</emphasis>
</taxonomicName>
); ovipositor with pair of well-developed paraprocts, partly enclosing basal half of palpi, and pair of weakly sclerotised and poorly defined elongate proctigers dorsal to these; vaginal palpi 2-segmented, second segment large and elongate; spermatheca falciform, with long coiled spermathecal duct which is expanded at attachment to bursa copulatrix.
</paragraph>
</subSubSection>
<subSubSection id="C1A06529FF83AC0EFF1AFE79C6F00BCB" lastPageId="3" lastPageNumber="56" pageId="2" pageNumber="55" type="distribution">
<paragraph id="890536A2FF83AC0FFF1AFE79C63309B6" blockId="2.[151,1437,152,642]" pageId="2" pageNumber="55">
<emphasis id="BBCEEAB0FF83AC0FFF1AFE79C6F20AD3" bold="true" box="[198,509,461,487]" pageId="2" pageNumber="55">Distribution and biology.</emphasis>
<taxonomicName id="4EBA4D21FF83AC0FFDDFFE79C58D0AD2" box="[515,642,461,486]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBCEEAB0FF83AC0FFDDFFE79C58D0AD2" box="[515,642,461,486]" italics="true" pageId="2" pageNumber="55">Canobolas</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="F1AD7632FF83AC0FFCF7FE79C4980AD3" box="[811,919,461,487]" name="Australia" pageId="2" pageNumber="55">Australia</collectingCountry>
(
<figureCitation id="11812A27FF83AC0FFC79FE79C4F40AD3" box="[933,1019,461,487]" captionStart="FIGURE 25" captionStartId="12.[151,255,1880,1904]" captionText="FIGURE 25. Maps of south-western and south-eastern Australia, showing localities for: Canobolas jarrah sp. nov. (˔); C. minang sp. nov. (●); C. nobilis (Lea) (Ο); C. tubrabucca sp. nov. ()." pageId="2" pageNumber="55">Fig. 25</figureCitation>
), where it occurs in the South-west, Southeastern New South
<collectingCountry id="F1AD7632FF83AC0FFE63FE40C509093A" box="[447,518,500,526]" name="United Kingdom" pageId="2" pageNumber="55">Wales</collectingCountry>
and McPherson-Macleay bioregions (Crisp
<emphasis id="BBCEEAB0FF83AC0FFBD0FE40C3480939" box="[1036,1095,500,525]" italics="true" pageId="2" pageNumber="55">et al.</emphasis>
1999). Western localities are low altitude (&lt;
<quantity id="4E429B47FF83AC0FFE95FDAEC69B0900" box="[329,404,538,564]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="55" unit="m" value="500.0">500m</quantity>
), dominated by closed eucalypt forest, whereas eastern localities are high altitude (at least&gt;
<quantity id="4E429B47FF83AC0FFF3AFDF5C621096F" box="[230,302,577,603]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="55" unit="m" value="800.0">800m</quantity>
, probably&gt;
<quantity id="4E429B47FF83AC0FFE61FDF5C51C096F" box="[445,531,577,603]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="55" unit="m" value="1100.0">1100m</quantity>
), in subalpine woodland (
<figureCitation id="11812A27FF83AC0FFC95FDF5C4AC096F" box="[841,931,577,603]" captionStart="FIGURE 26" captionStartId="12.[151,257,1950,1974]" captionTargetBox="[266,1327,443,1840]" captionTargetId="figure@12.[252,1335,433,1854]" captionTargetPageId="12" captionText="FIGURE 26. Snowgum (Eucalyptus pauciflora) woodland, 1380 m, Mount Canobolas, New South Wales; habitat of Canobolas nobilis (Lea). Photographed 2 November 2006, José Jurado-Rivera." httpUri="https://zenodo.org/record/189752/files/figure.png" pageId="2" pageNumber="55">Fig. 26</figureCitation>
). Each species is known from only one or two localities.
</paragraph>
<caption id="DDC5662AFF83AC0FFF4BF86EC33D0CC5" box="[151,1074,2010,2034]" httpUri="https://zenodo.org/record/189745/files/figure.png" pageId="2" pageNumber="55" targetBox="[306,1230,705,1953]" targetPageId="2">
<paragraph id="890536A2FF83AC0FFF4BF86EC33D0CC5" blockId="2.[151,1074,2010,2034]" box="[151,1074,2010,2034]" pageId="2" pageNumber="55">
<emphasis id="BBCEEAB0FF83AC0FFF4BF86EC64D0CC6" bold="true" box="[151,322,2010,2034]" pageId="2" pageNumber="55">FIGURES 14.</emphasis>
<taxonomicName id="4EBA4D21FF83AC0FFE95F86EC5400CC5" authority="Lea" authorityName="Lea" box="[329,591,2010,2033]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="BBCEEAB0FF83AC0FFE95F86EC5030CC5" box="[329,524,2010,2033]" italics="true" pageId="2" pageNumber="55">Canobolas nobilis</emphasis>
(Lea)
</taxonomicName>
, male: 1, dorsal; 2, lateral; 3, ventral; 4, face.
</paragraph>
</caption>
<paragraph id="890536A2FF82AC0EFF19FF2CC6F00BCB" blockId="3.[151,1437,152,1183]" pageId="3" pageNumber="56">
No beetles have been observed feeding on plants. DNA data from plant fragments in a specimen of
<taxonomicName id="4EBA4D21FF82AC0EFF4BFF0BC6620BEC" box="[151,365,191,216]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis id="BBCEEAB0FF82AC0EFF4BFF0BC6620BEC" box="[151,365,191,216]" italics="true" pageId="3" pageNumber="56">Canobolas nobilis</emphasis>
</taxonomicName>
suggest that the species is polyphagous (see under species description below). Immature stages are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C1A06529FF82AC0EFF1AFEBFC29F09C2" pageId="3" pageNumber="56" type="etymology">
<paragraph id="890536A2FF82AC0EFF1AFEBFC5180A47" blockId="3.[151,1437,152,1183]" pageId="3" pageNumber="56">
<emphasis id="BBCEEAB0FF82AC0EFF1AFEBFC65C0A11" bold="true" box="[198,339,267,293]" pageId="3" pageNumber="56">Etymology.</emphasis>
The genus is named from the massif of Mount
<taxonomicName id="4EBA4D21FF82AC0EFC64FEB8C3380A12" box="[952,1079,268,294]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. nov.">Canobolas</taxonomicName>
, where the
<typeStatus id="56018800FF82AC0EFB12FEB8C20F0A12" box="[1230,1280,268,294]" pageId="3" pageNumber="56">type</typeStatus>
species was collected.
<taxonomicName id="4EBA4D21FF82AC0EFECCFE86C6830A78" box="[272,396,306,332]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. nov.">Canobolas</taxonomicName>
means 'two shoulders' in the local Wiradjuri language (
<bibRefCitation id="ED2B4B53FF82AC0EFBC1FE86C3F80A78" author="Anonymous" box="[1053,1271,306,332]" pageId="3" pageNumber="66" refString="Anonymous (2003) Mount Canobolas State Conservation Area plan of management. New South Wales National Parks and Wildlife Service, Hurstville, New South Wales. 30 pp. [sighted online, October 2008: http: // www. environment. nsw. gov. au / resources / parks / pomfinalcanobolas. pdf]" type="book" year="2003">Anonymous 2003</bibRefCitation>
). The name is arbitrarily designated masculine.
</paragraph>
<paragraph id="890536A2FF82AC0EFF1AFECBC29F09C2" blockId="3.[151,1437,152,1183]" pageId="3" pageNumber="56">
<emphasis id="BBCEEAB0FF82AC0EFF1AFECBC66F0AAD" bold="true" box="[198,352,383,409]" pageId="3" pageNumber="56">Systematics.</emphasis>
The new genus is most similar to the endemic Australian genera
<taxonomicName id="4EBA4D21FF82AC0EFBBDFE34C2100AAE" authority="Baly" authorityName="Baly" box="[1121,1311,384,410]" class="Insecta" family="Chrysomelidae" genus="Cyclonoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFBBDFE34C3EF0AAD" box="[1121,1248,384,409]" italics="true" pageId="3" pageNumber="56">Cyclonoda</emphasis>
Baly
</taxonomicName>
,
<taxonomicName id="4EBA4D21FF82AC0EFAF0FE34C7C70AF4" authority="Lea" authorityName="Lea" class="Insecta" family="Chrysomelidae" genus="Ethomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFAF0FE34C2930AAD" box="[1324,1436,384,409]" italics="true" pageId="3" pageNumber="56">Ethomela</emphasis>
Lea
</taxonomicName>
,
<taxonomicName id="4EBA4D21FF82AC0EFF09FE13C6B60AF4" authority="Lea" authorityName="Lea" box="[213,441,422,448]" class="Insecta" family="Chrysomelidae" genus="Eugastromela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFF09FE13C68F0AF4" box="[213,384,423,448]" italics="true" pageId="3" pageNumber="56">Eugastromela</emphasis>
Lea
</taxonomicName>
,
<taxonomicName id="4EBA4D21FF82AC0EFE1BFE13C5610AF4" authority="Lea" authorityName="Lea" box="[455,622,422,448]" class="Insecta" family="Chrysomelidae" genus="Geomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFE1BFE13C53A0AF4" box="[455,565,423,448]" italics="true" pageId="3" pageNumber="56">Geomela</emphasis>
Lea
</taxonomicName>
,
<taxonomicName id="4EBA4D21FF82AC0EFDA0FE13C48A0AF4" authority="Daccordi" authorityName="Daccordi" box="[636,901,422,448]" class="Insecta" family="Chrysomelidae" genus="Gibbiomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFDA0FE13C4020AF4" box="[636,781,423,448]" italics="true" pageId="3" pageNumber="56">Gibbiomela</emphasis>
Daccordi
</taxonomicName>
and
<taxonomicName id="4EBA4D21FF82AC0EFC1FFE13C2720AF4" authority="Daccordi" authorityName="Daccordi" box="[963,1405,422,448]" class="Insecta" family="Chrysomelidae" genus="Palaeomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFC1FFE13C35A0AF4" box="[963,1109,423,448]" italics="true" pageId="3" pageNumber="56">Palaeomela</emphasis>
Daccordi (
<bibRefCitation id="ED2B4B53FF82AC0EFB03FE12C27B0AF4" author="Reid" box="[1247,1396,422,448]" pageId="3" pageNumber="66" refString="Reid, C. A. M. (2006 a) A taxonomic revision of the Australian Chrysomelinae, with a key to the genera (Coleoptera: Chrysomelidae), Zootaxa, 1292, 1 - 119." type="journal article" year="2006" yearSuffix="a">Reid 2006a</bibRefCitation>
)
</taxonomicName>
. It shares with them: body relatively small and ovoid; colour black or almost so, often with slightly metallic blue or green reflection; eyes laterally produced; trichobothria absent from each corner of pronotum.
<taxonomicName id="4EBA4D21FF82AC0EFACDFE40C2930939" box="[1297,1436,500,525]" class="Insecta" family="Chrysomelidae" genus="Gibbiomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFACDFE40C2930939" box="[1297,1436,500,525]" italics="true" pageId="3" pageNumber="56">Gibbiomela</emphasis>
</taxonomicName>
and
<taxonomicName id="4EBA4D21FF82AC0EFF15FDAEC65A0907" box="[201,341,538,563]" class="Insecta" family="Chrysomelidae" genus="Palaeomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFF15FDAEC65A0907" box="[201,341,538,563]" italics="true" pageId="3" pageNumber="56">Palaeomela</emphasis>
</taxonomicName>
are easily distinguished from
<taxonomicName id="4EBA4D21FF82AC0EFD66FDAEC4360907" box="[698,825,538,563]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBCEEAB0FF82AC0EFD66FDAEC4360907" box="[698,825,538,563]" italics="true" pageId="3" pageNumber="56">Canobolas</emphasis>
</taxonomicName>
: the former by the extremely narrow prosternal and mesoventrite processes and absence of subantennal grooves; the latter by closed procoxal cavities, absence of femoral plates, bilobed third tarsomeres and pygidium without median groove.
<taxonomicName id="4EBA4D21FF82AC0EFB8FFDDCC3F609B5" box="[1107,1273,616,641]" class="Insecta" family="Chrysomelidae" genus="Eugastromela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFB8FFDDCC3F609B5" box="[1107,1273,616,641]" italics="true" pageId="3" pageNumber="56">Eugastromela</emphasis>
</taxonomicName>
is also easily distinguished, by its deeply grooved facial sutures, lack of antenna-retaining ridges and grooves on head and prosternum, tuberculate elytra, absent femoral plates, bilobed third tarsomeres and penis without basal split. The remaining genera,
<taxonomicName id="4EBA4D21FF82AC0EFE7AFD68C52A09C1" box="[422,549,732,757]" class="Insecta" family="Chrysomelidae" genus="Cyclonoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFE7AFD68C52A09C1" box="[422,549,732,757]" italics="true" pageId="3" pageNumber="56">Cyclonoda</emphasis>
</taxonomicName>
,
<taxonomicName id="4EBA4D21FF82AC0EFDE8FD68C5AB09C1" box="[564,676,732,757]" class="Insecta" family="Chrysomelidae" genus="Ethomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFDE8FD68C5AB09C1" box="[564,676,732,757]" italics="true" pageId="3" pageNumber="56">Ethomela</emphasis>
</taxonomicName>
and
<taxonomicName id="4EBA4D21FF82AC0EFD01FD68C44809C1" box="[733,839,732,757]" class="Insecta" family="Chrysomelidae" genus="Geomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFD01FD68C44809C1" box="[733,839,732,757]" italics="true" pageId="3" pageNumber="56">Geomela</emphasis>
</taxonomicName>
, are more similar in size and shape to
<taxonomicName id="4EBA4D21FF82AC0EFAD6FD68C28609C1" box="[1290,1417,732,757]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBCEEAB0FF82AC0EFAD6FD68C28609C1" box="[1290,1417,732,757]" italics="true" pageId="3" pageNumber="56">Canobolas</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C1A06529FF82AC0EFF1AFCB6C6120FAB" pageId="3" pageNumber="56" type="description">
<paragraph id="890536A2FF82AC0EFF1AFCB6C7E40883" blockId="3.[151,1437,152,1183]" pageId="3" pageNumber="56">
<taxonomicName id="4EBA4D21FF82AC0EFF1AFCB6C63F082F" box="[198,304,770,795]" class="Insecta" family="Chrysomelidae" genus="Geomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFF1AFCB6C63F082F" box="[198,304,770,795]" italics="true" pageId="3" pageNumber="56">Geomela</emphasis>
</taxonomicName>
and
<taxonomicName id="4EBA4D21FF82AC0EFEB1FCB6C6D2082F" box="[365,477,770,795]" class="Insecta" family="Chrysomelidae" genus="Ethomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFEB1FCB6C6D2082F" box="[365,477,770,795]" italics="true" pageId="3" pageNumber="56">Ethomela</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="4EBA4D21FF82AC0EFD20FCB6C474082F" box="[764,891,770,795]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBCEEAB0FF82AC0EFD20FCB6C474082F" box="[764,891,770,795]" italics="true" pageId="3" pageNumber="56">Canobolas</emphasis>
</taxonomicName>
by: mesoventrite process not abruptly raised, less transverse (width &lt;
<date id="FD041062FF82AC0EFE69FC9DC6DD0877" box="[437,466,809,835]" pageId="3" pageNumber="56">2x</date>
length), with anterior margin not excavated to retain truncate apex of prosternal process; apex third tarsomere bilobed; apex last ventrite in male without rectangular internal flange; tegmen absent. Furthermore
<taxonomicName id="4EBA4D21FF82AC0EFE44FCC3C50208A4" box="[408,525,887,912]" class="Insecta" family="Chrysomelidae" genus="Ethomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFE44FCC3C50208A4" box="[408,525,887,912]" italics="true" pageId="3" pageNumber="56">Ethomela</emphasis>
</taxonomicName>
has closed procoxal cavities and the tibiae of
<taxonomicName id="4EBA4D21FF82AC0EFB90FCC3C3B508A4" box="[1100,1210,887,912]" class="Insecta" family="Chrysomelidae" genus="Geomela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFB90FCC3C3B508A4" box="[1100,1210,887,912]" italics="true" pageId="3" pageNumber="56">Geomela</emphasis>
</taxonomicName>
are not externally keeled.
</paragraph>
<paragraph id="890536A2FF82AC0EFF19FC70C59D0F66" blockId="3.[151,1437,152,1183]" pageId="3" pageNumber="56">
<taxonomicName id="4EBA4D21FF82AC0EFF19FC70C64B08E9" box="[197,324,964,989]" class="Insecta" family="Chrysomelidae" genus="Cyclonoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFF19FC70C64B08E9" box="[197,324,964,989]" italics="true" pageId="3" pageNumber="56">Cyclonoda</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4EBA4D21FF82AC0EFD96FC70C5C608E9" box="[586,713,964,989]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBCEEAB0FF82AC0EFD96FC70C5C608E9" box="[586,713,964,989]" italics="true" pageId="3" pageNumber="56">Canobolas</emphasis>
</taxonomicName>
by: male clypeus asymmetrically lobed; groove absent above eye; last maxillary palpomere elongate-cylindrical and truncate; elytral epipleura strongly sinuate, abruptly attenuate in apical third; metaventrite and first abdominal ventrite without femoral plates; pygidium without median groove; vaginal palpi 1-segmented.
</paragraph>
<paragraph id="890536A2FF82AC0EFF19FBEAC6120FAB" blockId="3.[151,1437,152,1183]" pageId="3" pageNumber="56">
On these criteria
<taxonomicName id="4EBA4D21FF82AC0EFE40FBEBC52E0F4C" box="[412,545,1119,1144]" class="Insecta" family="Chrysomelidae" genus="Canobolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBCEEAB0FF82AC0EFE40FBEBC52E0F4C" box="[412,545,1119,1144]" italics="true" pageId="3" pageNumber="56">Canobolas</emphasis>
</taxonomicName>
is a distinct monophyletic group of species, possibly the sistergroup of
<taxonomicName id="4EBA4D21FF82AC0EFF4BFB31C6190FAA" box="[151,278,1157,1182]" class="Insecta" family="Chrysomelidae" genus="Cyclonoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="BBCEEAB0FF82AC0EFF4BFB31C6190FAA" box="[151,278,1157,1182]" italics="true" pageId="3" pageNumber="56">Cyclonoda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>