516 lines
61 KiB
XML
516 lines
61 KiB
XML
<document id="0AB971CE755460F921E67E3355A4A620" ID-CLB-Dataset="28356" ID-DOI="10.11646/zootaxa.4567.2.10" ID-GBIF-Dataset="74d0dd04-67a2-427c-8638-16a0d306a232" ID-ISSN="1175-5326" ID-Zenodo-Dep="2594903" ID-ZooBank="24C5A2A2-9AB0-48DE-920D-A9BA260C1136" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552655962961" checkinUser="plazi" docAuthor="Sabella, Giorgio, Amore, Elena & Nicolosi, Giuseppe" docDate="2019" docId="B53A8785FFD2FF99E5B5BFD3FE5EA84D" docLanguage="en" docName="zootaxa.4567.2.10.pdf" docOrigin="Zootaxa 4567 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tychobythinus villasmundi Sabella & Amore & Nicolosi 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="385" masterDocId="4903FFFDFFD1FF9FE522BF0EFFD2A90D" masterDocTitle="A new troglobitic Tychobythinus from Sicily (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="386" masterPageNumber="379" pageNumber="382" updateTime="1698701799277" updateUser="ExternalLinkService">
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<mods:titleInfo id="9636C6C542F4A0A37E1E170A1B9336CC">
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<mods:title id="15F04BD1B6A8755E6F5B8C8628298186">A new troglobitic Tychobythinus from Sicily (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:name id="A0543E33D714668992EC012B3DA6F47A" type="personal">
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<mods:namePart id="D780E4AF8B888C51E238ADFDAFC61A9B">Sabella, Giorgio</mods:namePart>
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<mods:namePart id="80ED6E2F7322B1BB4B903D4310DB1DC0">Amore, Elena</mods:namePart>
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<mods:name id="C312CB9FF4506984EB0E958EF30F5017" type="personal">
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<mods:namePart id="4BEE63ACBA27616F9AAEC6BDBC918750">Nicolosi, Giuseppe</mods:namePart>
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<mods:title id="2BAEC0DCB328ACD160C04207D812ECDF">Zootaxa</mods:title>
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<mods:part id="FDE1404101CBE45040DABD2D8F5BBEF5">
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<mods:date id="EC1171BA5A66E47B27C9A3B58EC59709">2019</mods:date>
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<mods:number id="95F364AEDCD19D83C29C80F44C631909">2019-03-15</mods:number>
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<mods:number id="81D8350A8B99C56383118DA7C89409F7">4567</mods:number>
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<mods:identifier id="759429942B8276A23C0F39F5A504743F" type="DOI">10.11646/zootaxa.4567.2.10</mods:identifier>
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<treatment id="B53A8785FFD2FF99E5B5BFD3FE5EA84D" ID-DOI="http://doi.org/10.5281/zenodo.5933740" ID-GBIF-Taxon="154402288" ID-Zenodo-Dep="5933740" LSID="urn:lsid:plazi:treatment:B53A8785FFD2FF99E5B5BFD3FE5EA84D" httpUri="http://treatment.plazi.org/id/B53A8785FFD2FF99E5B5BFD3FE5EA84D" lastPageId="6" lastPageNumber="385" pageId="3" pageNumber="382">
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<subSubSection id="75896518FFD2FF9CE5B5BFD3FC79A9FA" box="[151,939,221,247]" pageId="3" pageNumber="382" type="nomenclature">
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<paragraph id="3D2C3693FFD2FF9CE5B5BFD3FC79A9FA" blockId="3.[151,939,221,280]" box="[151,939,221,247]" pageId="3" pageNumber="382">
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<heading id="666481FFFFD2FF9CE5B5BFD3FC79A9FA" bold="true" box="[151,939,221,247]" fontSize="11" level="1" pageId="3" pageNumber="382" reason="1">
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<emphasis id="0FE7EA81FFD2FF9CE5B5BFD3FC79A9FA" bold="true" box="[151,939,221,247]" pageId="3" pageNumber="382">
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<taxonomicName id="FA934D10FFD2FF9CE5B5BFD3FC94A9FA" ID-CoL="8YC4R" authority="Sabella, Amore and Nicolosi" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[151,838,221,247]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="382" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
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<emphasis id="0FE7EA81FFD2FF9CE5B5BFD3FE0CA9FA" bold="true" box="[151,478,221,247]" italics="true" pageId="3" pageNumber="382">Tychobythinus villasmundi</emphasis>
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||
Sabella, Amore and Nicolosi
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||
</taxonomicName>
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||
<taxonomicNameLabel id="14D457FAFFD2FF9CE66FBFD3FC79A9FA" box="[845,939,221,247]" pageId="3" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="75896518FFD2FF9CE5B5BFF1FECDA815" box="[151,287,255,280]" pageId="3" pageNumber="382" type="description">
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<paragraph id="3D2C3693FFD2FF9CE5B5BFF1FECDA815" blockId="3.[151,939,221,280]" box="[151,287,255,280]" pageId="3" pageNumber="382">
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(
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||
<figureCitation id="A5A82A16FFD2FF9CE5BDBFF1FEC5A815" box="[159,279,255,280]" captionStart-0="FIGURE 5" captionStart-1="FIGURES 6–9" captionStart-2="FIGURES 10–13" captionStartId-0="4.[151,249,1697,1719]" captionStartId-1="5.[151,263,1814,1837]" captionStartId-2="6.[151,263,1734,1757]" captionTargetBox-0="[397,1129,167,1633]" captionTargetBox-1="[244,1346,569,1757]" captionTargetBox-2="[357,1202,438,1646]" captionTargetId-0="figure@4.[334,1252,163,1676]" captionTargetId-1="figure@5.[174,1413,517,1794]" captionTargetId-2="figure@6.[300,1286,375,1713]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 5. Tychobythinus villasmundi sp. nov. Paratype female, habitus, dorsal view (Ph. A. Marletta)." captionText-1="FIGURES 6–9. Tychobythinus villasmundi sp. nov. 6. Head in dorsal view, schematic drawning. 7. Holotype male, head, dorsal view (Ph. A. Marletta). 8. Holotype male, head, lateral view (Ph. A. Marletta). 9. Holotype male, head, ventral view (Ph. A. Marletta). Scale: 0.1 mm." captionText-2="FIGURES 10–13. Tychobythinus villasmundi sp. nov. 10. Holotype male, left antenna. 11. Paratype female, rigth maxillary palpus. 12. Holotype male, aedeagus, dorsal view. 13. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." httpUri-0="https://zenodo.org/record/2594911/files/figure.png" httpUri-1="https://zenodo.org/record/2594914/files/figure.png" httpUri-2="https://zenodo.org/record/2594916/files/figure.png" pageId="3" pageNumber="382">Figs. 5–13</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="75896518FFD2FF9CE5B5BE49FE47AB19" pageId="3" pageNumber="382" type="materials_examined">
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<paragraph id="3D2C3693FFD2FF9CE5B5BE49FE47AB19" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="382">
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<emphasis id="0FE7EA81FFD2FF9CE5B5BE49FEEDA86D" bold="true" box="[151,319,327,352]" pageId="3" pageNumber="382">Type material</emphasis>
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.
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||
<materialsCitation id="8DFB3CCEFFD2FF9CE46CBE49FEE2A889" ID-GBIF-Occurrence="2013156507" collectingDate="2017-09-16" collectionCode="DBUC" collectorName="G. Nicolosi" country="Italy" county="Region" location="Villasmundo Cave" municipality="Melilli" pageId="3" pageNumber="382" specimenCount="1" specimenCount-male="1" stateProvince="Sicilia" typeStatus="holotype">
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<emphasis id="0FE7EA81FFD2FF9CE46CBE49FE69A86D" bold="true" box="[334,443,327,352]" pageId="3" pageNumber="382">
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<typeStatus id="E2288831FFD2FF9CE46CBE49FE69A86D" box="[334,443,327,352]" pageId="3" pageNumber="382" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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||
<emphasis id="0FE7EA81FFD2FF9CE4EEBE49FDCFA86D" bold="true" box="[460,541,327,352]" pageId="3" pageNumber="382">
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<collectingCountry id="45847603FFD2FF9CE4EEBE49FDCFA86D" box="[460,541,327,352]" name="Italy" pageId="3" pageNumber="382">ITALY</collectingCountry>
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||
</emphasis>
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||
:
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||
<collectingRegion id="FF57F871FFD2FF9CE70FBE49FDBCA86D" box="[557,622,327,352]" country="Italy" name="Sicilia" pageId="3" pageNumber="382">Sicily</collectingRegion>
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<collectingCounty id="D44D4E1FFFD2FF9CE755BE49FD1EA86D" box="[631,716,327,352]" pageId="3" pageNumber="382">Region</collectingCounty>
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:
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<collectingMunicipality id="DD48ACE9FFD2FF9CE7FBBE49FCF7A86D" box="[729,805,327,352]" pageId="3" pageNumber="382">Melilli</collectingMunicipality>
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(SR),
|
||
<location id="384C6048FFD2FF9CE650BE49FB97A86D" LSID="urn:lsid:plazi:treatment:B53A8785FFD2FF99E5B5BFD3FE5EA84D:384C6048FFD2FF9CE650BE49FB97A86D" box="[882,1093,327,352]" country="Italy" county="Region" municipality="Melilli" name="Villasmundo Cave" pageId="3" pageNumber="382" stateProvince="Sicilia">Villasmundo Cave</location>
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||
,
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||
<date id="492D1053FFD2FF9CE171BE49FB07A86D" box="[1107,1237,327,352]" pageId="3" pageNumber="382" value="2017-09-16">
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||
<collectingDate id="5969E9BBFFD2FF9CE171BE49FB07A86D" box="[1107,1237,327,352]" pageId="3" pageNumber="382" value="2017-09-16">16.IX.2017</collectingDate>
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||
</date>
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||
,
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||
<specimenCount id="2B95FD1AFFD2FF9CE1C0BE46FADDA852" box="[1250,1295,328,352]" pageId="3" pageNumber="382" type="male">1 ♂</specimenCount>
|
||
,
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||
<collectorName id="90665345FFD2FF9CE03FBE46FA4EA86D" box="[1309,1436,327,352]" pageId="3" pageNumber="382">G. Nicolosi</collectorName>
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||
leg. (
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<collectionCode id="5B82AE56FFD2FF9CE5F6BE62FEFAA889" box="[212,296,364,388]" pageId="3" pageNumber="382">DBUC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
|
||
<materialsCitation id="8DFB3CCEFFD2FF9CE41DBE62FD03A889" ID-GBIF-Occurrence="2013156514" box="[319,721,364,389]" country="Italy" location="Region" pageId="3" pageNumber="382" specimenCount="1" stateProvince="Sicilia" typeStatus="paratype">
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<emphasis id="0FE7EA81FFD2FF9CE41DBE62FE65A888" bold="true" box="[319,439,364,389]" pageId="3" pageNumber="382">
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||
<typeStatus id="E2288831FFD2FF9CE41DBE62FE65A888" box="[319,439,364,389]" pageId="3" pageNumber="382" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="0FE7EA81FFD2FF9CE4E8BE62FDC9A888" bold="true" box="[458,539,364,389]" pageId="3" pageNumber="382">
|
||
<collectingCountry id="45847603FFD2FF9CE4E8BE62FDC9A888" box="[458,539,364,389]" name="Italy" pageId="3" pageNumber="382">ITALY</collectingCountry>
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||
</emphasis>
|
||
:
|
||
<collectingRegion id="FF57F871FFD2FF9CE70DBE62FDA2A889" box="[559,624,364,388]" country="Italy" name="Sicilia" pageId="3" pageNumber="382">Sicily</collectingRegion>
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<location id="384C6048FFD2FF9CE75EBE62FD03A889" LSID="urn:lsid:plazi:treatment:B53A8785FFD2FF99E5B5BFD3FE5EA84D:384C6048FFD2FF9CE75EBE62FD03A889" box="[636,721,364,388]" country="Italy" name="Region" pageId="3" pageNumber="382" stateProvince="Sicilia">Region</location>
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</materialsCitation>
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:
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<materialsCitation id="8DFB3CCEFFD2FF9CE7C2BE62FB7EA889" ID-GBIF-Occurrence="2013156512" box="[736,1196,364,388]" collectionCode="DBUC" country="Italy" location="Region" pageId="3" pageNumber="382" specimenCount="2" specimenCount-female="2" stateProvince="Sicily" typeStatus="holotype">
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||
<specimenCount id="2B95FD1AFFD2FF9CE7C2BE62FCF7A88E" box="[736,805,364,388]" pageId="3" pageNumber="382" type="female">2 ♀♀</specimenCount>
|
||
, same data of
|
||
<typeStatus id="E2288831FFD2FF9CE6F8BE62FBEEA889" box="[986,1084,364,388]" pageId="3" pageNumber="382" type="holotype">holotype</typeStatus>
|
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(
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<collectionCode id="5B82AE56FFD2FF9CE172BE62FB76A889" box="[1104,1188,364,388]" pageId="3" pageNumber="382">DBUC</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="8DFB3CCEFFD2FF9CE19EBE62FDEFA8A5" ID-GBIF-Occurrence="2013156510" collectingDate="2018-05-31" collectionCode="DBUC" collectorName="G. Nicolosi" country="Italy" location="Region" pageId="3" pageNumber="382" specimenCount="1" specimenCount-female="1" stateProvince="Sicily" typeStatus="paratype">
|
||
<specimenCount id="2B95FD1AFFD2FF9CE19EBE62FB39A88E" box="[1212,1259,364,388]" pageId="3" pageNumber="382" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="492D1053FFD2FF9CE5B5BE9EFEDEA8A5" box="[151,268,400,424]" pageId="3" pageNumber="382" value="2018-05-31">
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||
<collectingDate id="5969E9BBFFD2FF9CE5B5BE9EFEDEA8A5" box="[151,268,400,424]" pageId="3" pageNumber="382" value="2018-05-31">31.V.2018</collectingDate>
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||
</date>
|
||
,
|
||
<collectorName id="90665345FFD2FF9CE439BE9EFE4EA8A5" box="[283,412,400,424]" pageId="3" pageNumber="382">G. Nicolosi</collectorName>
|
||
leg. (
|
||
<collectionCode id="5B82AE56FFD2FF9CE4C3BE9EFDE7A8A5" box="[481,565,400,424]" pageId="3" pageNumber="382">DBUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8DFB3CCEFFD2FF9CE769BE9EFAD3A8A5" ID-GBIF-Occurrence="2013156509" box="[587,1281,400,424]" collectingDate="2018-07-05" collectionCode="DBUC" collectorName="G. Nicolosi" country="Italy" location="Region" pageId="3" pageNumber="382" specimenCount="2" specimenCount-female="2" stateProvince="Sicily" typeStatus="paratype">
|
||
<specimenCount id="2B95FD1AFFD2FF9CE769BE9EFD5CA8AA" box="[587,654,400,424]" pageId="3" pageNumber="382" type="female">2 ♀♀</specimenCount>
|
||
, same locality,
|
||
<date id="492D1053FFD2FF9CE667BE9EFC03A8A5" box="[837,977,400,424]" pageId="3" pageNumber="382" value="2018-07-05">
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||
<collectingDate id="5969E9BBFFD2FF9CE667BE9EFC03A8A5" box="[837,977,400,424]" pageId="3" pageNumber="382" value="2018-07-05">05.VII.2018</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="90665345FFD2FF9CE6FDBE9EFBB2A8A5" box="[991,1120,400,424]" pageId="3" pageNumber="382">G. Nicolosi</collectorName>
|
||
leg. (
|
||
<collectionCode id="5B82AE56FFD2FF9CE187BE9EFB2BA8A5" box="[1189,1273,400,424]" pageId="3" pageNumber="382">DBUC</collectionCode>
|
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)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8DFB3CCEFFD2FF9CE032BE9EFD76A8C1" ID-GBIF-Occurrence="2013156508" collectingDate="2018-07-29" collectionCode="DBUC" collectorName="G. Nicolosi" country="Italy" location="Region" pageId="3" pageNumber="382" specimenCount="3" specimenCount-female="3" stateProvince="Sicily" typeStatus="paratype">
|
||
<specimenCount id="2B95FD1AFFD2FF9CE032BE9EFA81A8AA" box="[1296,1363,400,424]" pageId="3" pageNumber="382" type="female">3 ♀♀</specimenCount>
|
||
, same locality,
|
||
<date id="492D1053FFD2FF9CE5D4BEBAFE50A8C1" box="[246,386,436,460]" pageId="3" pageNumber="382" value="2018-07-29">
|
||
<collectingDate id="5969E9BBFFD2FF9CE5D4BEBAFE50A8C1" box="[246,386,436,460]" pageId="3" pageNumber="382" value="2018-07-29">29.VII.2018</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="90665345FFD2FF9CE4AFBEBAFDD8A8C1" box="[397,522,436,460]" pageId="3" pageNumber="382">G. Nicolosi</collectorName>
|
||
leg. (
|
||
<collectionCode id="5B82AE56FFD2FF9CE76BBEBAFD4FA8C1" box="[585,669,436,460]" pageId="3" pageNumber="382">DBUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8DFB3CCEFFD2FF9CE78DBEBAFA51A8C1" ID-GBIF-Occurrence="2013156513" box="[687,1411,436,460]" collectingDate="2018-10-11" collectionCode="DBUC" collectorName="G. Nicolosi" country="Italy" location="Region" pageId="3" pageNumber="382" specimenCount="2" specimenCount-female="2" stateProvince="Sicily" typeStatus="paratype">
|
||
<specimenCount id="2B95FD1AFFD2FF9CE78DBEBAFD3DA8C6" box="[687,751,436,460]" pageId="3" pageNumber="382" type="female">2 ♀♀</specimenCount>
|
||
, same locality,
|
||
<date id="492D1053FFD2FF9CE6B9BEBAFBC1A8C1" box="[923,1043,436,460]" pageId="3" pageNumber="382" value="2018-10-11">
|
||
<collectingDate id="5969E9BBFFD2FF9CE6B9BEBAFBC1A8C1" box="[923,1043,436,460]" pageId="3" pageNumber="382" value="2018-10-11">11.X.2018</collectingDate>
|
||
</date>
|
||
, trap 7,
|
||
<collectorName id="90665345FFD2FF9CE14EBEBAFB3BA8C1" box="[1132,1257,436,460]" pageId="3" pageNumber="382">G. Nicolosi</collectorName>
|
||
leg. (
|
||
<collectionCode id="5B82AE56FFD2FF9CE005BEBAFAA9A8C1" box="[1319,1403,436,460]" pageId="3" pageNumber="382">DBUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8DFB3CCEFFD2FF9CE0ACBEBAFC13A8FD" ID-GBIF-Occurrence="2013156511" collectingDate="2018-10-26" collectionCode="DBUC" collectorName="G. Nicolosi" country="Italy" location="Region" pageId="3" pageNumber="382" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Sicily" typeStatus="paratype">
|
||
<specimenCount id="2B95FD1AFFD2FF9CE0ACBEBAFF7EA8E2" pageId="3" pageNumber="382" type="male">1 ♂</specimenCount>
|
||
and
|
||
<specimenCount id="2B95FD1AFFD2FF9CE5C1BED6FEF6A8E2" box="[227,292,472,496]" pageId="3" pageNumber="382" type="female">2 ♀♀</specimenCount>
|
||
, same locality,
|
||
<date id="492D1053FFD2FF9CE4F1BED6FD99A8FD" box="[467,587,472,496]" pageId="3" pageNumber="382" value="2018-10-26">
|
||
<collectingDate id="5969E9BBFFD2FF9CE4F1BED6FD99A8FD" box="[467,587,472,496]" pageId="3" pageNumber="382" value="2018-10-26">26.X.2018</collectingDate>
|
||
</date>
|
||
, trap 7,
|
||
<collectorName id="90665345FFD2FF9CE785BED6FCF7A8FD" box="[679,805,472,496]" pageId="3" pageNumber="382">G. Nicolosi</collectorName>
|
||
leg. (
|
||
<collectionCode id="5B82AE56FFD2FF9CE647BED6FC6BA8FD" box="[869,953,472,496]" pageId="3" pageNumber="382">DBUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8DFB3CCEFFD2FF9CE6EEBED6FE43AB19" ID-GBIF-Occurrence="2013156515" collectingDate="2018-11-27" collectionCode="DBUC" collectorName="G. Nicolosi" country="Italy" location="Region" pageId="3" pageNumber="382" specimenCount="1" specimenCount-female="1" stateProvince="Sicily" typeStatus="paratype">
|
||
<specimenCount id="2B95FD1AFFD2FF9CE6EEBED6FC25A8E2" box="[972,1015,472,496]" pageId="3" pageNumber="382" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="492D1053FFD2FF9CE185BED6FAFBA8FD" box="[1191,1321,472,496]" pageId="3" pageNumber="382" value="2018-11-27">
|
||
<collectingDate id="5969E9BBFFD2FF9CE185BED6FAFBA8FD" box="[1191,1321,472,496]" pageId="3" pageNumber="382" value="2018-11-27">27.XI.2018</collectingDate>
|
||
</date>
|
||
, trap 7,
|
||
<collectorName id="90665345FFD2FF9CE0A7BED6FF24AB18" pageId="3" pageNumber="382">G. Nicolosi</collectorName>
|
||
leg. (
|
||
<collectionCode id="5B82AE56FFD2FF9CE417BEF2FE5BAB19" box="[309,393,508,532]" pageId="3" pageNumber="382">DBUC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75896518FFD2FF9BE5E5BD11FD6AAE6E" lastPageId="4" lastPageNumber="383" pageId="3" pageNumber="382" type="description">
|
||
<paragraph id="3D2C3693FFD2FF9CE5E5BD11FDA3ABA8" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="382">
|
||
<emphasis id="0FE7EA81FFD2FF9CE5E5BD11FE83AB35" bold="true" box="[199,337,543,568]" pageId="3" pageNumber="382">Description</emphasis>
|
||
.
|
||
<emphasis id="0FE7EA81FFD2FF9CE446BD11FE4FAB35" box="[356,413,543,568]" italics="true" pageId="3" pageNumber="382">Male</emphasis>
|
||
: Length
|
||
<quantity id="FA6B9B76FFD2FF9CE72EBD11FD67AB3A" box="[524,693,543,568]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="3" pageNumber="382" unit="mm" value="1.275" valueMax="1.3" valueMin="1.25">1.25–1.30 mm</quantity>
|
||
, apterous and anophthalmous. Entirely pale brown with lighter antennae, palpi, and legs. Pubescence consists of long and flattened setae (length:
|
||
<quantity id="FA6B9B76FFD2FF9CE15ABD4AFAF1AB51" box="[1144,1315,580,604]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7" pageId="3" pageNumber="382" unit="mm" value="0.075" valueMax="0.08" valueMin="0.07">0.07–0.08 mm</quantity>
|
||
) on head, pronotum, elytra and abdomen, other suberect shorter setae (length:
|
||
<quantity id="FA6B9B76FFD2FF9CE6B3BD66FBE4AB8D" box="[913,1078,616,640]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="382" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.03–0.04 mm</quantity>
|
||
) on antennae, and legs, and thin fluff on apical segment of maxillary palpi.
|
||
</paragraph>
|
||
<paragraph id="3D2C3693FFD2FF9CE5E5BDA1FBC1AC78" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="382">
|
||
Head (
|
||
<figureCitation id="A5A82A16FFD2FF9CE433BDA1FEAFABC5" box="[273,381,687,712]" captionStart="FIGURES 6–9" captionStartId="5.[151,263,1814,1837]" captionTargetBox="[244,1346,569,1757]" captionTargetId="figure@5.[174,1413,517,1794]" captionTargetPageId="5" captionText="FIGURES 6–9. Tychobythinus villasmundi sp. nov. 6. Head in dorsal view, schematic drawning. 7. Holotype male, head, dorsal view (Ph. A. Marletta). 8. Holotype male, head, lateral view (Ph. A. Marletta). 9. Holotype male, head, ventral view (Ph. A. Marletta). Scale: 0.1 mm." httpUri="https://zenodo.org/record/2594914/files/figure.png" pageId="3" pageNumber="382">Figs. 6–7</figureCitation>
|
||
) wider (
|
||
<quantity id="FA6B9B76FFD2FF9CE4F8BDBEFD93ABC5" box="[474,577,688,712]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="382" unit="mm" value="0.3">0.30 mm</quantity>
|
||
) than long (
|
||
<quantity id="FA6B9B76FFD2FF9CE7EABDA1FC59ABC5" box="[712,907,687,712]" metricMagnitude="-4" metricUnit="m" metricValue="2.375" metricValueMax="2.4" metricValueMin="2.3499999999999996" pageId="3" pageNumber="382" unit="mm" value="0.2375" valueMax="0.24" valueMin="0.235">0.235–0.240 mm</quantity>
|
||
) slightly narrower than pronotum. Frontal lobe wider (
|
||
<quantity id="FA6B9B76FFD2FF9CE5C4BDDAFE9FABE1" box="[230,333,724,749]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="382" unit="mm" value="0.15">0.15 mm</quantity>
|
||
) than long (
|
||
<quantity id="FA6B9B76FFD2FF9CE4F7BDDAFD99ABE1" box="[469,587,724,749]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="382" unit="mm" value="0.075">0.075 mm</quantity>
|
||
) with subparallel, protruding and sharp sides; antennal tubercle protruding. Frons between antennal tubercles with large median sulcus reaching the anterior edge of vertexal foveae. Sides of frontal region with some punctures. Clypeal carina well-defined, equally visible in dorsal as well in lateral view, extends to ocular region. Tempora rounded, convex occipital region traversed by median longitudinal sulcus reaching over posterior edge of vertexal foveae, late wide and well-impressed. Gular region (
|
||
<figureCitation id="A5A82A16FFD2FF9CE1F4BC6AFAEFAA71" box="[1238,1341,868,893]" captionStart="FIGURES 6–9" captionStartId="5.[151,263,1814,1837]" captionTargetBox="[244,1346,569,1757]" captionTargetId="figure@5.[174,1413,517,1794]" captionTargetPageId="5" captionText="FIGURES 6–9. Tychobythinus villasmundi sp. nov. 6. Head in dorsal view, schematic drawning. 7. Holotype male, head, dorsal view (Ph. A. Marletta). 8. Holotype male, head, lateral view (Ph. A. Marletta). 9. Holotype male, head, ventral view (Ph. A. Marletta). Scale: 0.1 mm." httpUri="https://zenodo.org/record/2594914/files/figure.png" pageId="3" pageNumber="382">Figs 8–9</figureCitation>
|
||
) behind labium with deep and broad impression margined posteriorly by transverse ridge projecting on each side of head. This ridge with long and acute median process, projecting ventrally, and bearing tuft of setae on the tip. Base of the gular region with two long and sturdy medians bristles. Antennae (
|
||
<figureCitation id="A5A82A16FFD2FF9CE6A3BCC1FC00AAE5" box="[897,978,975,1000]" captionStart="FIGURES 10–13" captionStartId="6.[151,263,1734,1757]" captionTargetBox="[357,1202,438,1646]" captionTargetId="figure@6.[300,1286,375,1713]" captionTargetPageId="6" captionText="FIGURES 10–13. Tychobythinus villasmundi sp. nov. 10. Holotype male, left antenna. 11. Paratype female, rigth maxillary palpus. 12. Holotype male, aedeagus, dorsal view. 13. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." httpUri="https://zenodo.org/record/2594916/files/figure.png" pageId="3" pageNumber="382">Fig. 10</figureCitation>
|
||
)
|
||
<quantity id="FA6B9B76FFD2FF9CE6C0BCC1FB72AAEA" box="[994,1184,975,1000]" metricMagnitude="-4" metricUnit="m" metricValue="6.375" metricValueMax="6.4" metricValueMin="6.35" pageId="3" pageNumber="382" unit="mm" value="0.6375" valueMax="0.64" valueMin="0.635">0.635–0.640 mm</quantity>
|
||
long, scape more than 3.5 times longer (
|
||
<quantity id="FA6B9B76FFD2FF9CE47BBCFAFDC9AD01" box="[345,539,1012,1037]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.85" metricValueMin="1.75" pageId="3" pageNumber="382" unit="mm" value="0.18" valueMax="0.185" valueMin="0.175">0.175–0.185 mm</quantity>
|
||
) than wide (
|
||
<quantity id="FA6B9B76FFD2FF9CE787BCFAFCDCAD01" box="[677,782,1012,1037]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="382" unit="mm" value="0.05">0.05 mm</quantity>
|
||
), narrowed and flattened with protruding and sharp medial margin in basal third, wider in middle. Pedicel ovoid, slightly asymmetric, about one and a half times longer than wide, slightly tighter than scape, and distinctly wider than funicular segments. Antennomere III longer than wide and slightly narrowed at base; antennomere IV as long as wide, antennomere V longer than wide, antennomeres VI–VII subequal and as long as wide, antennomere VIII wider than long. Antennal club consisting of last three antennomeres which are broaden progressively from IX to XI. Antennomere IX slightly wider than long, antennomere X distinctly wider than long, antennomere XI distinctly longer than wide and twice as long as combined length of antennomeres IX and X. Maxillary palpi with palpomere II elongated and gradually expanded from base to apex, its surface covered by 4–6 tubercles concentrated in the distal portion. Palpomere III slightly longer than wide, its surface with 2–3 tubercles; last palpomere about 4 times as long (
|
||
<quantity id="FA6B9B76FFD2FF9CE1A4BA36FB3DAC5D" box="[1158,1263,1336,1360]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="382" unit="mm" value="0.24">0.24 mm</quantity>
|
||
) as wide (
|
||
<quantity id="FA6B9B76FFD2FF9CE04BBA39FF15AC79" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="382" unit="mm" value="0.06">0.06 mm</quantity>
|
||
), widest in basal third, lateral margin slightly curved and sinuate in middle.
|
||
</paragraph>
|
||
<paragraph id="3D2C3693FFD2FF9CE5E5BA8FFD0AAF7D" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="382">
|
||
Pronotum slightly wider (
|
||
<quantity id="FA6B9B76FFD2FF9CE4CEBA8EFD87AC95" box="[492,597,1408,1432]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="382" unit="mm" value="0.31">0.31 mm</quantity>
|
||
) than long (
|
||
<quantity id="FA6B9B76FFD2FF9CE7C0BA8EFC9CAC95" box="[738,846,1408,1432]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="382" unit="mm" value="0.3">0.30 mm</quantity>
|
||
), widest near basal third, anteriorly slightly tapered with sinuates sides, posteriorly more tapered with straight sides. Its dorsal surface shiny with some rare and fain punctures. Pleural carina oblique, well-defined. Two well impressed antebasal lateral fovea linked by the antebasal sulcus. Tegument between pronotal posterior margin and antebasal sulcus rough, it makes difficult to recognize median antebasal fovea. Metaventrite with median egg-shaped impression beginning from its posterior margin and extending just before mesocoxal cavities, and bearing for its entire length a median longitudinal carina. Base of mesocoxal cavities with pubescent pit on each side.
|
||
</paragraph>
|
||
<paragraph id="3D2C3693FFD2FF9CE5E5B972FDDCAE0D" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="382">
|
||
Elytra distinctly wider (
|
||
<quantity id="FA6B9B76FFD2FF9CE4F1B972FDE9AF99" box="[467,571,1660,1685]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="382" unit="mm" value="0.55">0.55 mm</quantity>
|
||
) than long (
|
||
<quantity id="FA6B9B76FFD2FF9CE7E1B972FCFCAF99" box="[707,814,1660,1684]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="382" unit="mm" value="0.49">0.49 mm</quantity>
|
||
), convex, sides slightly rounded from base to the apex, widest near middle. Humeral calli strongly reduced. Dorsal surface shiny with only some superficial punctures. Each elytron with two basal foveae, subhumeral fovea well-defined. Both, marginal and sutural striae reaching elytral apex, discal striae lacking.
|
||
</paragraph>
|
||
<paragraph id="3D2C3693FFD2FF9CE5E5B802FC97AE28" blockId="3.[151,1437,327,2009]" box="[199,837,1804,1829]" pageId="3" pageNumber="382">Abdomen normally shaped without particular characters.</paragraph>
|
||
<paragraph id="3D2C3693FFD2FF9CE5E5B83FFAA4AE9D" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="382">
|
||
Legs relatively long and thin. Protrochanters with 1–2 tubercles on ventral surface, protibiae slightly flattened and sinuate in distal third, tarsomere II of protarsi slightly dilated. Meso and metatrochanters simple; femora simple, mesotibiae simple, length of metatibiae:
|
||
<quantity id="FA6B9B76FFD2FF9CE795B879FC8EAE9D" box="[695,860,1911,1936]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="3" pageNumber="382" unit="mm" value="0.455" valueMax="0.46" valueMin="0.45">0.45–0.46 mm</quantity>
|
||
, slightly enlarged and sinuate in the distal third.
|
||
</paragraph>
|
||
<paragraph id="3D2C3693FFD2FF9CE5E5B892FA92AED4" blockId="3.[151,1437,327,2009]" pageId="3" pageNumber="382">
|
||
Aedeagus (
|
||
<figureCitation id="A5A82A16FFD2FF9CE464B892FE48AEB9" box="[326,410,1948,1973]" captionStart="FIGURES 10–13" captionStartId="6.[151,263,1734,1757]" captionTargetBox="[357,1202,438,1646]" captionTargetId="figure@6.[300,1286,375,1713]" captionTargetPageId="6" captionText="FIGURES 10–13. Tychobythinus villasmundi sp. nov. 10. Holotype male, left antenna. 11. Paratype female, rigth maxillary palpus. 12. Holotype male, aedeagus, dorsal view. 13. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." httpUri="https://zenodo.org/record/2594916/files/figure.png" pageId="3" pageNumber="382">Fig. 12</figureCitation>
|
||
)
|
||
<quantity id="FA6B9B76FFD2FF9CE489B892FD83AEB9" box="[427,593,1948,1973]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="3" pageNumber="382" unit="mm" value="0.245" valueMax="0.25" valueMin="0.24">0.24–0.25 mm</quantity>
|
||
long, ovoid with relatively short parameres, that are convergent and with narrow apex, each bearing two subapical setae. Internal sac with 2 big and long teeth and 3 smaller spines.
|
||
</paragraph>
|
||
<caption id="69EC661BFFD5FF9BE5B5B9AFFB68AFBA" box="[151,1210,1697,1719]" httpUri="https://zenodo.org/record/2594911/files/figure.png" pageId="4" pageNumber="383" startId="4.[151,249,1697,1719]" targetBox="[397,1129,167,1633]" targetPageId="4">
|
||
<paragraph id="3D2C3693FFD5FF9BE5B5B9AFFB68AFBA" blockId="4.[151,1210,1697,1719]" box="[151,1210,1697,1719]" pageId="4" pageNumber="383">
|
||
<emphasis id="0FE7EA81FFD5FF9BE5B5B9AFFEDEAFBA" bold="true" box="[151,268,1697,1719]" pageId="4" pageNumber="383">FIGURE 5</emphasis>
|
||
.
|
||
<taxonomicName id="FA934D10FFD5FF9BE43AB9AFFDF5AFBA" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[280,551,1697,1719]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="383" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD5FF9BE43AB9AFFDF5AFBA" box="[280,551,1697,1719]" italics="true" pageId="4" pageNumber="383">Tychobythinus villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD5FF9BE70FB9ACFDAFAFBA" bold="true" box="[557,637,1698,1719]" pageId="4" pageNumber="383">
|
||
<taxonomicNameLabel id="14D457FAFFD5FF9BE70FB9ACFDAFAFBA" box="[557,637,1698,1719]" pageId="4" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Paratype female, habitus, dorsal view (Ph. A. Marletta).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3D2C3693FFD5FF9BE5E5B9D0FD6AAE6E" blockId="4.[151,1436,1758,2036]" pageId="4" pageNumber="383">
|
||
<emphasis id="0FE7EA81FFD5FF9BE5E5B9D0FEC9AFFA" box="[199,283,1758,1783]" italics="true" pageId="4" pageNumber="383">Female</emphasis>
|
||
: Similar to male (
|
||
<figureCitation id="A5A82A16FFD5FF9BE4CBB9D0FDFFAFFA" box="[489,557,1758,1783]" captionStart="FIGURE 5" captionStartId="4.[151,249,1697,1719]" captionTargetBox="[397,1129,167,1633]" captionTargetId="figure@4.[334,1252,163,1676]" captionTargetPageId="4" captionText="FIGURE 5. Tychobythinus villasmundi sp. nov. Paratype female, habitus, dorsal view (Ph. A. Marletta)." httpUri="https://zenodo.org/record/2594911/files/figure.png" pageId="4" pageNumber="383">Fig. 5</figureCitation>
|
||
), length
|
||
<quantity id="FA6B9B76FFD5FF9BE7B0B9D0FCEAAFFA" box="[658,824,1758,1783]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.45" metricValueMin="1.25" pageId="4" pageNumber="383" unit="mm" value="1.35" valueMax="1.45" valueMin="1.25">1.25–1.45 mm</quantity>
|
||
, head slightly wider (
|
||
<quantity id="FA6B9B76FFD5FF9BE10EB9D0FB01AFFA" box="[1068,1235,1758,1783]" metricMagnitude="-4" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="4" pageNumber="383" unit="mm" value="0.255" valueMax="0.26" valueMin="0.25">0.25–0.26 mm</quantity>
|
||
) than long (
|
||
<quantity id="FA6B9B76FFD5FF9BE079B9D1FF2DAE16" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="4" pageNumber="383" unit="mm" value="0.235" valueMax="0.24" valueMin="0.23">0.23– 0.24 mm</quantity>
|
||
), gular region unmodified, slightly convex. Maxillary palpi with surface of palpomere II covered by 12– 16 tubercles (
|
||
<figureCitation id="A5A82A16FFD5FF9BE410B828FE57AE32" box="[306,389,1830,1855]" captionStart="FIGURES 10–13" captionStartId="6.[151,263,1734,1757]" captionTargetBox="[357,1202,438,1646]" captionTargetId="figure@6.[300,1286,375,1713]" captionTargetPageId="6" captionText="FIGURES 10–13. Tychobythinus villasmundi sp. nov. 10. Holotype male, left antenna. 11. Paratype female, rigth maxillary palpus. 12. Holotype male, aedeagus, dorsal view. 13. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." httpUri="https://zenodo.org/record/2594916/files/figure.png" pageId="4" pageNumber="383">Fig. 11</figureCitation>
|
||
). Metaventrite lacking median impression; legs with protibiae, tarsomere II of protarsi, and metatibiae unmodified. Telisternite as in
|
||
<figureCitation id="A5A82A16FFD5FF9BE743B845FD66AE6E" box="[609,692,1867,1892]" captionStart="FIGURES 10–13" captionStartId="6.[151,263,1734,1757]" captionTargetBox="[357,1202,438,1646]" captionTargetId="figure@6.[300,1286,375,1713]" captionTargetPageId="6" captionText="FIGURES 10–13. Tychobythinus villasmundi sp. nov. 10. Holotype male, left antenna. 11. Paratype female, rigth maxillary palpus. 12. Holotype male, aedeagus, dorsal view. 13. Paratype female, telisternite, dorsal view. Scale: 0.1 mm." httpUri="https://zenodo.org/record/2594916/files/figure.png" pageId="4" pageNumber="383">Fig. 13</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75896518FFD5FF99E5E5B860FEBCA9D9" lastPageId="6" lastPageNumber="385" pageId="4" pageNumber="383" type="discussion">
|
||
<paragraph id="3D2C3693FFD5FF9BE5E5B860FC5DAEF9" blockId="4.[151,1436,1758,2036]" pageId="4" pageNumber="383">
|
||
<emphasis id="0FE7EA81FFD5FF9BE5E5B860FE9EAE8A" bold="true" box="[199,332,1902,1927]" pageId="4" pageNumber="383">Discussion.</emphasis>
|
||
<taxonomicName id="FA934D10FFD5FF9BE47EB87EFDD1AE8A" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[348,515,1902,1927]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="383" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD5FF9BE47EB87EFDD1AE8A" box="[348,515,1902,1927]" italics="true" pageId="4" pageNumber="383">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD5FF9BE731B860FDA7AE8A" bold="true" box="[531,629,1902,1927]" pageId="4" pageNumber="383">
|
||
<taxonomicNameLabel id="14D457FAFFD5FF9BE731B860FDA7AE8A" box="[531,629,1902,1927]" pageId="4" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from all other
|
||
<taxonomicName id="FA934D10FFD5FF9BE6BBB860FBE8AE8A" authorityName="Ganglbauer" authorityYear="1896" box="[921,1082,1902,1927]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="383" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0FE7EA81FFD5FF9BE6BBB860FBE8AE8A" box="[921,1082,1902,1927]" italics="true" pageId="4" pageNumber="383">Tychobythinus</emphasis>
|
||
</taxonomicName>
|
||
species by its aedeagal and exoskeleton features. However, it shows affinities with
|
||
<taxonomicName id="FA934D10FFD5FF9BE62EB89DFC7FAEA1" authorityName="Ganglbauer" authorityYear="1896" box="[780,941,1939,1964]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="383" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0FE7EA81FFD5FF9BE62EB89DFC7FAEA1" box="[780,941,1939,1964]" italics="true" pageId="4" pageNumber="383">Tychobythinus</emphasis>
|
||
</taxonomicName>
|
||
species from North Africa which belong to two distinct groups (
|
||
<bibRefCitation id="59024B62FFD5FF9BE459B8B8FD82AEC2" author="Sabella, G. & Viglianisi F. M. & D'Urso, V. & Nouira, S." box="[379,592,1974,1999]" pageId="4" pageNumber="383" pagination="579 - 586" refId="ref3321" refString="Sabella, G., Viglianisi F. M., D'Urso, V. & Nouira, S. (2014). A review of the Tychobythinus algiricus species group from North Africa (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 3893 (4), 579 - 586. https: // doi. org / 10.11646 / zootaxa. 3893.4.7" type="journal article" year="2014">
|
||
Sabella
|
||
<emphasis id="0FE7EA81FFD5FF9BE4F7B8B6FDDFAEC2" box="[469,525,1974,1999]" italics="true" pageId="4" pageNumber="383">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). Since their external morphology is closely related to their biology, it is not necessary they really represent two homogeneous phyletic lineages.
|
||
</paragraph>
|
||
<paragraph id="3D2C3693FFD4FF9AE5E5BF99FB9BA810" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="384">
|
||
The aedeagus of
|
||
<taxonomicName id="FA934D10FFD4FF9AE4B4BF97FDE9A9BD" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[406,571,151,176]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD4FF9AE4B4BF97FDE9A9BD" box="[406,571,151,176]" italics="true" pageId="5" pageNumber="384">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD4FF9AE765BF99FD74A9BD" bold="true" box="[583,678,151,176]" pageId="5" pageNumber="384">
|
||
<taxonomicNameLabel id="14D457FAFFD4FF9AE765BF99FD74A9BD" box="[583,678,151,176]" pageId="5" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is comparable with the species belonging to the
|
||
<taxonomicName id="FA934D10FFD4FF9AE1D9BF99FF29A9D8" baseAuthorityName="Raffray" baseAuthorityYear="1871" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="algiricus">
|
||
<emphasis id="0FE7EA81FFD4FF9AE1D9BF99FF29A9D8" italics="true" pageId="5" pageNumber="384">Tychobythinus algiricus</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<bibRefCitation id="59024B62FFD4FF9AE47BBFB2FDE9A9D9" author="Sabella, G. & Viglianisi F. M. & D'Urso, V. & Nouira, S." box="[345,571,188,213]" pageId="5" pageNumber="384" pagination="579 - 586" refId="ref3321" refString="Sabella, G., Viglianisi F. M., D'Urso, V. & Nouira, S. (2014). A review of the Tychobythinus algiricus species group from North Africa (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 3893 (4), 579 - 586. https: // doi. org / 10.11646 / zootaxa. 3893.4.7" type="journal article" year="2014">
|
||
Sabella
|
||
<emphasis id="0FE7EA81FFD4FF9AE49BBFB3FE27A9D8" box="[441,501,188,213]" italics="true" pageId="5" pageNumber="384">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
;
|
||
<taxonomicName id="FA934D10FFD4FF9AE769BFB2FC2EA9D8" authority="sensu Jeannel 1956" authorityName="Jeannel" authorityYear="1956" box="[587,1020,188,213]" class="Insecta" family="Staphylinidae" genus="Chiasmatobythus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0FE7EA81FFD4FF9AE769BFB2FCDFA9D8" box="[587,781,188,213]" italics="true" pageId="5" pageNumber="384">Chiasmatobythus</emphasis>
|
||
sensu
|
||
<bibRefCitation id="59024B62FFD4FF9AE643BFB2FC2EA9D8" author="Jeannel, R." box="[865,1020,188,213]" pageId="5" pageNumber="384" pagination="1 - 233" refId="ref2982" refString="Jeannel, R. (1956) Les Pselaphides de l'Afrique du Nord. Essai de biogeographie berbere. Memoires du Museum National d'Histoire Naturelle, Paris, n. s. (Serie A: Zoologie), 14, 1 - 233." type="journal article" year="1956">Jeannel 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
), especially to
|
||
<taxonomicName id="FA934D10FFD4FF9AE18DBFB3FF09A9F5" authority="(Raffray 1871)" authorityName="Raffray" authorityYear="1871" baseAuthorityName="Raffray" baseAuthorityYear="1871" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="algiricus">
|
||
<emphasis id="0FE7EA81FFD4FF9AE18DBFB3FAE3A9D8" box="[1199,1329,188,213]" italics="true" pageId="5" pageNumber="384">T. algiricus</emphasis>
|
||
(
|
||
<bibRefCitation id="59024B62FFD4FF9AE067BFB2FF00A9F5" author="Raffray, A." pageId="5" pageNumber="384" pagination="159 - 160" refId="ref3078" refString="Raffray, A. (1871) Petites nouvelles. Petites Nouvelles Entomologiques, 3, 159 - 160." type="journal article" year="1871">Raffray 1871</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which is widespread in the Maghreb (
|
||
<collectingCountry id="45847603FFD4FF9AE7B6BFD1FD39A9F5" box="[660,747,223,248]" name="Tunisia" pageId="5" pageNumber="384">Tunisia</collectingCountry>
|
||
, northern
|
||
<collectingCountry id="45847603FFD4FF9AE67DBFD1FC61A9F5" box="[863,947,223,248]" name="Algeria" pageId="5" pageNumber="384">Algeria</collectingCountry>
|
||
and
|
||
<collectingCountry id="45847603FFD4FF9AE6CEBFEEFB84A9F5" box="[1004,1110,224,248]" name="Morocco" pageId="5" pageNumber="384">Morocco</collectingCountry>
|
||
). However, all the species in this group are macro or microphthalmous, whereas
|
||
<taxonomicName id="FA934D10FFD4FF9AE7FABE0BFCA4A810" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[728,886,260,285]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="villasmundi">
|
||
<emphasis id="0FE7EA81FFD4FF9AE7FABE0BFCA4A810" box="[728,886,260,285]" italics="true" pageId="5" pageNumber="384">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
is anophthalmous.
|
||
</paragraph>
|
||
<paragraph id="3D2C3693FFD4FF9AE5E5BE26FE01A8DD" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="384">
|
||
Because of troglobitic adaptations,
|
||
<taxonomicName id="FA934D10FFD4FF9AE741BE27FCD7A84D" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[611,773,295,320]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="villasmundi">
|
||
<emphasis id="0FE7EA81FFD4FF9AE741BE27FCD7A84D" box="[611,773,295,320]" italics="true" pageId="5" pageNumber="384">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
can be also compared with the species belonging to the
|
||
<taxonomicName id="FA934D10FFD4FF9AE5B5BE42FE54A868" baseAuthorityName="Guillebeau" baseAuthorityYear="1894" box="[151,390,332,357]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="theryi">
|
||
<emphasis id="0FE7EA81FFD4FF9AE5B5BE42FE54A868" box="[151,390,332,357]" italics="true" pageId="5" pageNumber="384">Tychobythinus theryi</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<bibRefCitation id="59024B62FFD4FF9AE4C8BE42FD03A869" author="Sabella, G. & D'Urso, V. & Nouira, S. & Souissi, R. & Besuchet, C." box="[490,721,332,357]" pageId="5" pageNumber="384" pagination="63 - 75" refId="ref3248" refString="Sabella, G., D'Urso, V., Nouira, S., Souissi, R. & Besuchet, C. (2013) A review of Tychobythinus theryi species group with description of a new species from Algeria (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 3701 (1), 63 - 75. https: // doi. org / 10.11646 / zootaxa. 3701.1.5" type="journal article" year="2013">
|
||
Sabella
|
||
<emphasis id="0FE7EA81FFD4FF9AE769BE43FD50A868" box="[587,642,332,357]" italics="true" pageId="5" pageNumber="384">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
;
|
||
<taxonomicName id="FA934D10FFD4FF9AE7C1BE42FBBEA868" authority="sensu Jeannel 1956" authorityName="Jeannel" authorityYear="1956" box="[739,1132,332,357]" class="Insecta" family="Staphylinidae" genus="Anopsibythus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0FE7EA81FFD4FF9AE7C1BE42FC16A869" box="[739,964,332,357]" italics="true" pageId="5" pageNumber="384">Anopsibythus sensu</emphasis>
|
||
<bibRefCitation id="59024B62FFD4FF9AE6F0BE42FBBEA868" author="Jeannel, R." box="[978,1132,332,357]" pageId="5" pageNumber="384" pagination="1 - 233" refId="ref2982" refString="Jeannel, R. (1956) Les Pselaphides de l'Afrique du Nord. Essai de biogeographie berbere. Memoires du Museum National d'Histoire Naturelle, Paris, n. s. (Serie A: Zoologie), 14, 1 - 233." type="journal article" year="1956">Jeannel 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
) and, especially with the wingless and eyeless
|
||
<taxonomicName id="FA934D10FFD4FF9AE4B2BE7FFD1EA885" authority="(Guillebeau 1894)" authorityName="Guillebeau" authorityYear="1894" baseAuthorityName="Guillebeau" baseAuthorityYear="1894" box="[400,716,367,392]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="theryi">
|
||
<emphasis id="0FE7EA81FFD4FF9AE4B2BE7FFE22A885" box="[400,496,367,392]" italics="true" pageId="5" pageNumber="384">T. theryi</emphasis>
|
||
(
|
||
<bibRefCitation id="59024B62FFD4FF9AE723BE61FD16A885" author="Guillebeau, F." box="[513,708,367,392]" pageId="5" pageNumber="384" pagination="46 - 47" refId="ref2916" refString="Guillebeau, F. (1894) Description de nouveaux Pselaphides d'Algerie. L'Echange, Revue Linneenne, 10, 46 - 47." type="journal article" year="1894">Guillebeau 1894</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, known only from three females from Saint Charles (northern
|
||
<collectingCountry id="45847603FFD4FF9AE5B5BE9AFF22A8A0" box="[151,240,404,429]" name="Algeria" pageId="5" pageNumber="384">Algeria</collectingCountry>
|
||
), and
|
||
<taxonomicName id="FA934D10FFD4FF9AE413BE9BFDB5A8A0" authority="(Jeannel 1956)" authorityName="Jeannel" authorityYear="1956" baseAuthorityName="Jeannel" baseAuthorityYear="1956" box="[305,615,404,429]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="normandi">
|
||
<emphasis id="0FE7EA81FFD4FF9AE413BE9BFE6BA8A0" box="[305,441,404,429]" italics="true" pageId="5" pageNumber="384">T. normandi</emphasis>
|
||
(
|
||
<bibRefCitation id="59024B62FFD4FF9AE4E5BE9AFDB2A8A0" author="Jeannel, R." box="[455,608,404,429]" pageId="5" pageNumber="384" pagination="1 - 233" refId="ref2982" refString="Jeannel, R. (1956) Les Pselaphides de l'Afrique du Nord. Essai de biogeographie berbere. Memoires du Museum National d'Histoire Naturelle, Paris, n. s. (Serie A: Zoologie), 14, 1 - 233." type="journal article" year="1956">Jeannel 1956</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which is known only from a single male from Philippeville (old name of
|
||
<collectingRegion id="FF57F871FFD4FF9AE5B5BEB9FF35A8DD" box="[151,231,439,464]" country="Algeria" name="Skikda" pageId="5" pageNumber="384">Skikda</collectingRegion>
|
||
) in northern
|
||
<collectingCountry id="45847603FFD4FF9AE455BEB9FE1DA8DD" box="[375,463,439,464]" name="Algeria" pageId="5" pageNumber="384">Algeria</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption id="69EC661BFFD4FF9AE5B5B818FE6BAE67" httpUri="https://zenodo.org/record/2594914/files/figure.png" pageId="5" pageNumber="384" startId="5.[151,263,1814,1837]" targetBox="[244,1346,569,1757]" targetPageId="5">
|
||
<paragraph id="3D2C3693FFD4FF9AE5B5B818FE6BAE67" blockId="5.[151,1436,1814,1898]" pageId="5" pageNumber="384">
|
||
<emphasis id="0FE7EA81FFD4FF9AE5B5B818FEE4AE21" bold="true" box="[151,310,1814,1837]" pageId="5" pageNumber="384">FIGURES 6–9</emphasis>
|
||
.
|
||
<taxonomicName id="FA934D10FFD4FF9AE465B819FD8AAE20" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[327,600,1815,1837]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD4FF9AE465B819FD8AAE20" box="[327,600,1815,1837]" italics="true" pageId="5" pageNumber="384">Tychobythinus villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD4FF9AE740B819FD64AE21" bold="true" box="[610,694,1815,1836]" pageId="5" pageNumber="384">
|
||
<taxonomicNameLabel id="14D457FAFFD4FF9AE740B819FD64AE21" box="[610,694,1815,1836]" pageId="5" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
6. Head in dorsal view, schematic drawning. 7. Holotype male, head, dorsal view (Ph. A. Marletta). 8. Holotype male, head, lateral view (Ph. A. Marletta). 9. Holotype male, head, ventral view (Ph. A. Marletta). Scale: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3D2C3693FFD4FF99E5E5B89CFEBCA9D9" blockId="5.[151,1436,1938,2035]" lastBlockId="6.[151,1436,151,320]" lastPageId="6" lastPageNumber="385" pageId="5" pageNumber="384">
|
||
<taxonomicName id="FA934D10FFD4FF9AE5E5B89CFE21AEA6" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[199,499,1938,1963]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD4FF9AE5E5B89CFE21AEA6" box="[199,499,1938,1963]" italics="true" pageId="5" pageNumber="384">Tychobythinus villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD4FF9AE4D9B89CFD84AEA6" bold="true" box="[507,598,1938,1963]" pageId="5" pageNumber="384">
|
||
<taxonomicNameLabel id="14D457FAFFD4FF9AE4D9B89CFD84AEA6" box="[507,598,1938,1963]" pageId="5" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily distinguished from
|
||
<taxonomicName id="FA934D10FFD4FF9AE6F7B89DFBE1AEA6" baseAuthorityName="Guillebeau" baseAuthorityYear="1894" box="[981,1075,1938,1963]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="theryi">
|
||
<emphasis id="0FE7EA81FFD4FF9AE6F7B89DFBE1AEA6" box="[981,1075,1938,1963]" italics="true" pageId="5" pageNumber="384">T. theryi</emphasis>
|
||
</taxonomicName>
|
||
by its shiny tegument of dorsal surface (slightly but distinctly punctuate in
|
||
<taxonomicName id="FA934D10FFD4FF9AE7ABB8B9FD35AEC2" baseAuthorityName="Guillebeau" baseAuthorityYear="1894" box="[649,743,1974,1999]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="theryi">
|
||
<emphasis id="0FE7EA81FFD4FF9AE7ABB8B9FD35AEC2" box="[649,743,1974,1999]" italics="true" pageId="5" pageNumber="384">T. theryi</emphasis>
|
||
</taxonomicName>
|
||
), the wider frontal lobe (
|
||
<quantity id="FA6B9B76FFD4FF9AE12BB8B8FBA3AEC3" box="[1033,1137,1974,1999]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="384" unit="mm" value="0.15">0.15 mm</quantity>
|
||
in
|
||
<taxonomicName id="FA934D10FFD4FF9AE1B5B8B9FAE5AEC2" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[1175,1335,1974,1999]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD4FF9AE1B5B8B9FAE5AEC2" box="[1175,1335,1974,1999]" italics="true" pageId="5" pageNumber="384">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD4FF9AE062B8B8FA4EAEC2" bold="true" box="[1344,1436,1974,1999]" pageId="5" pageNumber="384">
|
||
<taxonomicNameLabel id="14D457FAFFD4FF9AE062B8B8FA4EAEC2" box="[1344,1436,1974,1999]" pageId="5" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
versus
|
||
<quantity id="FA6B9B76FFD4FF9AE5C5B8D4FE4BAEFF" box="[231,409,2010,2035]" metricMagnitude="-4" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="5" pageNumber="384" unit="mm" value="0.1275" valueMax="0.13" valueMin="0.125">0.125–0.13 mm</quantity>
|
||
in
|
||
<taxonomicName id="FA934D10FFD4FF9AE4E2B8D5FDCCAEFE" baseAuthorityName="Guillebeau" baseAuthorityYear="1894" box="[448,542,2010,2035]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="theryi">
|
||
<emphasis id="0FE7EA81FFD4FF9AE4E2B8D5FDCCAEFE" box="[448,542,2010,2035]" italics="true" pageId="5" pageNumber="384">T. theryi</emphasis>
|
||
</taxonomicName>
|
||
), longer antennae (
|
||
<quantity id="FA6B9B76FFD4FF9AE7D9B8D4FC70AEFF" box="[763,930,2010,2035]" metricMagnitude="-4" metricUnit="m" metricValue="6.35" metricValueMax="6.4" metricValueMin="6.3" pageId="5" pageNumber="384" unit="mm" value="0.635" valueMax="0.64" valueMin="0.63">0.63–0.64 mm</quantity>
|
||
in
|
||
<taxonomicName id="FA934D10FFD4FF9AE6EAB8D5FBBAAEFE" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[968,1128,2010,2035]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="384" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD4FF9AE6EAB8D5FBBAAEFE" box="[968,1128,2010,2035]" italics="true" pageId="5" pageNumber="384">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD4FF9AE153B8D4FB1EAEFE" bold="true" box="[1137,1228,2010,2035]" pageId="5" pageNumber="384">
|
||
<taxonomicNameLabel id="14D457FAFFD4FF9AE153B8D4FB1EAEFE" box="[1137,1228,2010,2035]" pageId="5" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, versus
|
||
<quantity id="FA6B9B76FFD4FF99E00EB8D4FF11A9A2" lastPageId="6" lastPageNumber="385" metricMagnitude="-4" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="5" pageNumber="384" unit="mm" value="0.555" valueMax="0.56" valueMin="0.55">0.55–0.56 mm</quantity>
|
||
in
|
||
<taxonomicName id="FA934D10FFD7FF99E5C7BF97FE93A9BD" baseAuthorityName="Guillebeau" baseAuthorityYear="1894" box="[229,321,151,176]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="theryi">
|
||
<emphasis id="0FE7EA81FFD7FF99E5C7BF97FE93A9BD" box="[229,321,151,176]" italics="true" pageId="6" pageNumber="385">T. theryi</emphasis>
|
||
</taxonomicName>
|
||
), and by the presence of less tubercles on the palpomere II (
|
||
<quantity id="FA6B9B76FFD7FF99E6FDBF99FB91A9BD" box="[991,1091,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.064" metricValueMin="3.048" pageId="6" pageNumber="385" unit="in" value="14.0" valueMax="16.0" valueMin="12.0">12–16 in</quantity>
|
||
<taxonomicName id="FA934D10FFD7FF99E16BBF97FB35A9BD" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[1097,1255,151,176]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD7FF99E16BBF97FB35A9BD" box="[1097,1255,151,176]" italics="true" pageId="6" pageNumber="385">T. villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD7FF99E1CFBF99FA94A9BD" bold="true" box="[1261,1350,151,176]" pageId="6" pageNumber="385">
|
||
<taxonomicNameLabel id="14D457FAFFD7FF99E1CFBF99FA94A9BD" box="[1261,1350,151,176]" pageId="6" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
against
|
||
<quantity id="FA6B9B76FFD7FF99E5B5BFB2FF28A9D8" box="[151,250,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" metricValueMax="7.62" metricValueMin="6.096" pageId="6" pageNumber="385" unit="in" value="27.0" valueMax="30.0" valueMin="24.0">24–30 in</quantity>
|
||
<taxonomicName id="FA934D10FFD7FF99E423BFB3FE8FA9D8" baseAuthorityName="Guillebeau" baseAuthorityYear="1894" box="[257,349,188,213]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="theryi">
|
||
<emphasis id="0FE7EA81FFD7FF99E423BFB3FE8FA9D8" box="[257,349,188,213]" italics="true" pageId="6" pageNumber="385">T. theryi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75896518FFD7FF99E5E5BFD1FE5EA84D" pageId="6" pageNumber="385" type="description">
|
||
<paragraph id="3D2C3693FFD7FF99E5E5BFD1FE5EA84D" blockId="6.[151,1436,151,320]" pageId="6" pageNumber="385">
|
||
<taxonomicName id="FA934D10FFD7FF99E5E5BFD1FE23A9F5" authority="Sabella & Amore & Nicolosi, 2019" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[199,497,223,248]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="villasmundi" status="sp. nov.">
|
||
<emphasis id="0FE7EA81FFD7FF99E5E5BFD1FE23A9F5" box="[199,497,223,248]" italics="true" pageId="6" pageNumber="385">Tychobythinus villasmundi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0FE7EA81FFD7FF99E4D5BFD1FD82A9F5" bold="true" box="[503,592,223,248]" pageId="6" pageNumber="385">
|
||
<taxonomicNameLabel id="14D457FAFFD7FF99E4D5BFD1FD82A9F5" box="[503,592,223,248]" pageId="6" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="FA934D10FFD7FF99E7C0BFEFFCB8A9F5" baseAuthorityName="Jeannel" baseAuthorityYear="1956" box="[738,874,223,248]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="normandi">
|
||
<emphasis id="0FE7EA81FFD7FF99E7C0BFEFFCB8A9F5" box="[738,874,223,248]" italics="true" pageId="6" pageNumber="385">T. normandi</emphasis>
|
||
</taxonomicName>
|
||
mainly by the features on the tempora (rounded in
|
||
<taxonomicName id="FA934D10FFD7FF99E5B5BE0AFE16A810" authorityName="Sabella & Amore & Nicolosi" authorityYear="2019" box="[151,452,260,285]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="villasmundi">
|
||
<emphasis id="0FE7EA81FFD7FF99E5B5BE0AFE16A810" box="[151,452,260,285]" italics="true" pageId="6" pageNumber="385">Tychobythinus villasmundi</emphasis>
|
||
</taxonomicName>
|
||
, angulated and dug in
|
||
<taxonomicName id="FA934D10FFD7FF99E7E9BE0BFC87A810" baseAuthorityName="Jeannel" baseAuthorityYear="1956" box="[715,853,260,285]" class="Insecta" family="Staphylinidae" genus="Tychobythinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="385" phylum="Arthropoda" rank="species" species="normandi">
|
||
<emphasis id="0FE7EA81FFD7FF99E7E9BE0BFC87A810" box="[715,853,260,285]" italics="true" pageId="6" pageNumber="385">T. normandi</emphasis>
|
||
</taxonomicName>
|
||
). From both species the new species differs by the structure of aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |