219 lines
18 KiB
XML
219 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1160.98022" ID-Pensoft-Pub="1313-2970-1160-1" ID-Pensoft-UUID="B0708A590ABD58FEB4A1A474CA268AA4" ID-ZooBank="B7E449240D3D4529987F19A51FAA115E" ModsDocID="1313-2970-1160-1" checkinTime="1683131562219" checkinUser="pensoft" docAuthor="Man, Nem Sian, Lwin, Ngwe, Sutcharit, Chirasak & Panha, Somsak" docDate="2023" docId="B0C5CC33056352C8ABE877501B1AD032" docLanguage="en" docName="ZooKeys 1160: 1-59" docOrigin="ZooKeys 1160" docPubDate="2023-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1160.98022" docTitle="Oospira luneainopsis Man & Panha 2023, sp. nov." docType="treatment" docUuid="B8866312-31E1-4E9A-993D-6A3E025C8F17" docUuidSource="ZooBank" docVersion="1" id="B0708A590ABD58FEB4A1A474CA268AA4" lastPageNumber="1" masterDocId="B0708A590ABD58FEB4A1A474CA268AA4" masterDocTitle="Further notes on the taxonomy of the land snail family Clausiliidae Gray, 1855 (Stylommatophora, Helicina) from Myanmar with description of two new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="1" updateTime="1683131562219" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Further notes on the taxonomy of the land snail family Clausiliidae Gray, 1855 (Stylommatophora, Helicina) from Myanmar with description of two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Man, Nem Sian</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lwin, Ngwe</mods:namePart>
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<mods:affiliation>Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1160</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>59</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1160.98022</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1160.98022</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1160-1</mods:identifier>
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<mods:identifier type="ZooBank">B7E449240D3D4529987F19A51FAA115E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">B0708A590ABD58FEB4A1A474CA268AA4</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:B8866312-31E1-4E9A-993D-6A3E025C8F17" httpUri="http://treatment.plazi.org/id/B0C5CC33056352C8ABE877501B1AD032" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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12
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<taxonomicName LSID="https://zoobank.org/B8866312-31E1-4E9A-993D-6A3E025C8F17" authority="Man & Panha" authorityName="Man & Panha" authorityYear="2023" class="Gastropoda" family="Clausiliidae" genus="Oospira" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Oospira luneainopsis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="luneainopsis" status="sp. nov.">Oospira luneainopsis Man & Panha</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Shell and clausilial apparatus of A-C Oospira luneainopsis sp. nov. A shell of holotype CUMZ 13062 from Zwekabin Mountain, Kayin State B, C paratypes CUMZ 13063 from the type locality B clausilial apparatus C shell and D-F Oospira zediopsis sp. nov. D clausilial apparatus of paratype CUMZ 13065 from the type locality E shell of holotype CUMZ 13064 from Weibyan Cave, Kayin State and F shell of paratype CUMZ 13065 from the type locality." figureDoi="10.3897/zookeys.1160.98022.figure16" httpUri="https://binary.pensoft.net/fig/845478" pageId="0" pageNumber="1">Figs 16A-C</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Synoptic view of the clausiliid species recorded from Myanmar A Indonenia admirabilis, holotype NHMUK 20200185 B Oospira andersoniana, specimen CUMZ 13054 C Oospira arakana, lectotype NHMUK 1888.12.4.1037 D Cylindrophaedusa bacillum, specimen NHMUK 1906.2.2.345 E Oospira bulbus, possible syntype NHMUK 1903.7.1.1278 F Phaedusa burmanica, holotype NHMUK 1888.12.4.1058 G Oospira decollata, paratype ZISP H Indonenia excellens, specimen NHMUK 20200189 I Oospira fusiformis, holotype NHMUK 1906.2.2.380 J Oospira gouldiana, lectotype NHMUK 196548 K Oospira gracilior, lectotype NHMUK 1907.12.30.243 L Oospira insignis, lectotype USNM 117120 M Phaedusa kazueae, holotype NHMUK 20200187 N Phaedusa lemani (after Gude, 1914: fig. 107) O Oospira limborgi, holotype NHMUK 1903.7.1.1266, and P Oospira luneainopsis sp. nov., holotype CUMZ 13062." figureDoi="10.3897/zookeys.1160.98022.figure18" httpUri="https://binary.pensoft.net/fig/845480" pageId="0" pageNumber="1">, 18P</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 13062 (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Shell and clausilial apparatus of A-C Oospira luneainopsis sp. nov. A shell of holotype CUMZ 13062 from Zwekabin Mountain, Kayin State B, C paratypes CUMZ 13063 from the type locality B clausilial apparatus C shell and D-F Oospira zediopsis sp. nov. D clausilial apparatus of paratype CUMZ 13065 from the type locality E shell of holotype CUMZ 13064 from Weibyan Cave, Kayin State and F shell of paratype CUMZ 13065 from the type locality." figureDoi="10.3897/zookeys.1160.98022.figure16" httpUri="https://binary.pensoft.net/fig/845478" pageId="0" pageNumber="1">16A</figureCitation>
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). Measurements: shell height 20 mm, shell width 5.8 mm, and 7 whorls.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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CUMZ 13063 (8 shells; Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Shell and clausilial apparatus of A-C Oospira luneainopsis sp. nov. A shell of holotype CUMZ 13062 from Zwekabin Mountain, Kayin State B, C paratypes CUMZ 13063 from the type locality B clausilial apparatus C shell and D-F Oospira zediopsis sp. nov. D clausilial apparatus of paratype CUMZ 13065 from the type locality E shell of holotype CUMZ 13064 from Weibyan Cave, Kayin State and F shell of paratype CUMZ 13065 from the type locality." figureDoi="10.3897/zookeys.1160.98022.figure16" httpUri="https://binary.pensoft.net/fig/845478" pageId="0" pageNumber="1">16B, C</figureCitation>
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), NHMUK 20220504 (2 shells).
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</paragraph>
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<caption doi="10.3897/zookeys.1160.98022.figure16" httpUri="https://binary.pensoft.net/fig/845478" pageId="0" pageNumber="1" start="Figure 16" startId="F16">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 16.</emphasis>
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Shell and clausilial apparatus of
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<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
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<taxonomicName authorityName="Man & Panha" authorityYear="2023" class="Gastropoda" family="Clausiliidae" genus="Oospira" kingdom="Animalia" lsidName="Oospira luneainopsis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="luneainopsis">
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<emphasis italics="true" pageId="0" pageNumber="1">Oospira luneainopsis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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shell of holotype CUMZ 13062 from Zwekabin Mountain, Kayin State
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<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
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paratypes CUMZ 13063 from the type locality
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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clausilial apparatus
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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shell and
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<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
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<taxonomicName authorityName="Man & Panha" authorityYear="2023" class="Gastropoda" family="Clausiliidae" genus="Oospira" kingdom="Animalia" lsidName="Oospira zediopsis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="zediopsis">
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<emphasis italics="true" pageId="0" pageNumber="1">Oospira zediopsis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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clausilial apparatus of paratype CUMZ 13065 from the type locality
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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shell of holotype CUMZ 13064 from Weibyan Cave, Kayin State and
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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shell of paratype CUMZ 13065 from the type locality.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Limestone outcrop at Zwegabin Mountain, Hpa-an Township, Hpa-an District, Kayin State, Myanmar (
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<geoCoordinate degrees="16" direction="north" minutes="48" orientation="latitude" precision="1" seconds="44.5" value="16.81236">16°48'44.5"N</geoCoordinate>
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,
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<geoCoordinate degrees="97" direction="east" minutes="40" orientation="longitude" precision="1" seconds="25.5" value="97.673744">97°40'25.5"E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The specific name
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<emphasis italics="true" pageId="0" pageNumber="1">luneainopsis</emphasis>
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is a compound of
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<normalizedToken originalValue="‘lun-eain’">'lun-eain'</normalizedToken>
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, which in the Myanmar language refers to a bobbin used for traditional Myanmar weaving, and the Greek suffix -
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<emphasis italics="true" pageId="0" pageNumber="1">opsis</emphasis>
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meaning 'having the appearance of or
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<normalizedToken originalValue="like’">like'</normalizedToken>
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. It refers to the bobbin shape of the shell of this species.
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</paragraph>
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<caption doi="10.3897/zookeys.1160.98022.figure17" httpUri="https://binary.pensoft.net/fig/845479" pageId="0" pageNumber="1" start="Figure 17" startId="F17">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 17.</emphasis>
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Shell of
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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<taxonomicName baseAuthorityName="Likharev" baseAuthorityYear="1962" class="Gastropoda" family="Clausiliidae" genus="Oospira" kingdom="Animalia" lsidName="Oospira decollata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="decollata">
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<emphasis italics="true" pageId="0" pageNumber="1">Oospira decollata</emphasis>
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</taxonomicName>
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, paratype ZISP from West Burma, near Pakoku on Mount Victoria. 2200 m a.s.l.
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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<taxonomicName baseAuthorityName="Gould" baseAuthorityYear="1843" class="Gastropoda" family="Clausiliidae" genus="Oospira" kingdom="Animalia" lsidName="Oospira insignis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="insignis">
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<emphasis italics="true" pageId="0" pageNumber="1">Oospira insignis</emphasis>
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</taxonomicName>
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, lectotype USNM 117120 from Tavoy, Burmah
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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<taxonomicName authorityName="H. Nordsieck" authorityYear="1973" class="Gastropoda" family="Clausiliidae" genus="Oospira" kingdom="Animalia" lsidName="Oospira malaisei" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="malaisei">
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<emphasis italics="true" pageId="0" pageNumber="1">Oospira malaisei</emphasis>
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</taxonomicName>
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, holotype SMNH 3014 from NO-Birma: Pungkataung ca. 3000 ft. and
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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<taxonomicName baseAuthorityName="Gould" baseAuthorityYear="1856" class="Gastropoda" family="Clausiliidae" genus="Oospira" kingdom="Animalia" lsidName="Oospira vespa" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vespa">
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<emphasis italics="true" pageId="0" pageNumber="1">Oospira vespa</emphasis>
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</taxonomicName>
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, lectotype MCZ 169455 from Tavoy, British Burma. Photo: A Persson, SMNH (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell ovate-fusiform, decollated with smooth surface and dense striations near suture. Last whorl neck subquadrate; aperture nearly roundish; peristome detached. Inferior lamella straight ascending and reaching at peristome; palatal plicae four, distinct, equally, and parallelly arranged, and superior lamella developed.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell ovate-fusiform, decollated, opaque, pinkish brown; spire blunt. Shell surface smooth to with fine growth lines on last whorl, denser near suture and last whorl neck subquadrate; suture impressed and distinct. Whorls 7-8, convex, regularly growing, and bluntly attenuated to apex. Aperture nearly round; peristome detached, thickened, expanded and little reflexed. Superior lamella developed, continuous with spiralis, and slightly low at transition to spiralis. Inferior lamella steeply ascending, distant from superior lamella and ending at peristome. Subcolumellaris emerged and invisible in oblique view. Principalis running along lateral-dorsal side and anterior end visible through oblique apertural view. Palatal plicae lateral, four: first and fourth plicae strong and longest; remaining plicae parallel and equal length; sometimes tiny lowest plica present. Clausilium plate lateral side, narrow and slightly pointed at tip.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Man & Panha" authorityYear="2023" class="Gastropoda" family="Clausiliidae" genus="Oospira" kingdom="Animalia" lsidName="Oospira luneainopsis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="luneainopsis">
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<emphasis italics="true" pageId="0" pageNumber="1">Oospira luneainopsis</emphasis>
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</taxonomicName>
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sp. nov. is only known from the type locality.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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No ethanol preserved specimens were available for anatomical study. This species can be distinguished from
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<taxonomicName lsidName="O. gouldiana" pageId="0" pageNumber="1" rank="species" species="gouldiana">
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<emphasis italics="true" pageId="0" pageNumber="1">O. gouldiana</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="O. magna" pageId="0" pageNumber="1" rank="species" species="magna">
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<emphasis italics="true" pageId="0" pageNumber="1">O. magna</emphasis>
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</taxonomicName>
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, and
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<taxonomicName lsidName="O. zediopsis" pageId="0" pageNumber="1" rank="species" species="zediopsis">
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<emphasis italics="true" pageId="0" pageNumber="1">O. zediopsis</emphasis>
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</taxonomicName>
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sp. nov. by its small size, decollated, fewer whorls, roundish aperture, more spiral inferior lamella, and equal palatal plicae. This new species is clearly distinct from
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<taxonomicName lsidName="O. decollata" pageId="0" pageNumber="1" rank="species" species="decollata">
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<emphasis italics="true" pageId="0" pageNumber="1">O. decollata</emphasis>
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</taxonomicName>
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by its spiral inferior lamella visible from aperture, smoother shell, pink color, and more bulging whorls.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Compared to the Vietnamese dextral species,
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<taxonomicName lsidName="O. duci" pageId="0" pageNumber="1" rank="species" species="duci">
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<emphasis italics="true" pageId="0" pageNumber="1">O. duci</emphasis>
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</taxonomicName>
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Maassen & Gittenberger, 2007 exhibits thinner and oblique aperture, sinus higher, obviously emerged subcolumellaris, very fine striations and more ventrally unequal palatal plicae than this species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |