140 lines
16 KiB
XML
140 lines
16 KiB
XML
<document id="D715637EBEC62D455051DF85ADE2C729" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos & Cerdeño, Esperanza" docDate="2019" docId="B0798797FFB4286672CBCFD2FE5CFBEE" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Icochilus robustus Ameghino 1891" docType="treatment" docVersion="8" lastPageNumber="209" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="209" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="5ED721ED436AAAFE3F029C547EA90883">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="F61BBFFEA4B286086F2A9296A0106840">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="E107F291D9683B66E9830E429FB50C96">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="363B679F339A9F2387E62C048040C76A">Cerdeño, Esperanza</mods:namePart>
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<mods:title id="1762380073DB83E925061CFBB234EB31">Zootaxa</mods:title>
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<mods:date id="3690D90D66CFB9BDEB49178078C6AF25">2019</mods:date>
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<mods:number id="C8EC1D9BC88EE97E35CD4E0F7DD39F93">2019-01-07</mods:number>
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<treatment id="B0798797FFB4286672CBCFD2FE5CFBEE" ID-DOI="http://doi.org/10.5281/zenodo.5931293" ID-GBIF-Taxon="154529762" ID-Zenodo-Dep="5931293" LSID="urn:lsid:plazi:treatment:B0798797FFB4286672CBCFD2FE5CFBEE" httpUri="http://treatment.plazi.org/id/B0798797FFB4286672CBCFD2FE5CFBEE" lastPageNumber="209" pageId="14" pageNumber="209">
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<subSubSection id="70CA650AFFB4286672CBCFD2FDC6FE65" box="[151,584,429,455]" pageId="14" pageNumber="209" type="nomenclature">
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<paragraph id="386F3681FFB4286672CBCFD2FDC6FE65" blockId="14.[151,584,429,455]" box="[151,584,429,455]" pageId="14" pageNumber="209">
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<heading id="632781EDFFB4286672CBCFD2FDC6FE65" bold="true" box="[151,584,429,455]" fontSize="11" level="1" pageId="14" pageNumber="209" reason="1">
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<taxonomicName id="FFD04D02FFB4286672CBCFD2FDC6FE65" authority="Ameghino, 1891" authorityName="Ameghino" authorityYear="1891" box="[151,584,429,455]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="robustus">
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<emphasis id="0AA4EA93FFB4286672CBCFD2FDC6FE65" bold="true" box="[151,584,429,455]" pageId="14" pageNumber="209">
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<emphasis id="0AA4EA93FFB4286672CBCFD2FEF8FE65" bold="true" box="[151,374,429,455]" italics="true" pageId="14" pageNumber="209">Icochilus robustus</emphasis>
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<bibRefCitation id="5C414B70FFB428667322CFD2FDC6FE65" author="Ameghino, F." box="[382,584,429,455]" pageId="14" pageNumber="209" pagination="289 - 328" refId="ref16209" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article" year="1891">Ameghino, 1891</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFB4286672CBCF8BFCFBFCAA" pageId="14" pageNumber="209" type="materials_examined">
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<paragraph id="386F3681FFB4286672CBCF8BFE70FD92" blockId="14.[151,1437,500,1101]" pageId="14" pageNumber="209">
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<materialsCitation id="88B83CDCFFB4286672CBCF8BFE70FD92" ID-GBIF-Occurrence="2147525207" collectionCode="MACN-A" pageId="14" pageNumber="209" specimenCode="MACN-A 3454, MACN-A 3505" specimenCount="1" typeStatus="holotype">
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<emphasis id="0AA4EA93FFB4286672CBCF8BFE85FDAF" bold="true" box="[151,267,500,525]" pageId="14" pageNumber="209">
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<typeStatus id="E76B8823FFB4286672CBCF8BFE88FDAF" box="[151,262,500,525]" pageId="14" pageNumber="209" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="68769EFAFFB42866734ECF8BFE4DFDAF" box="[274,451,500,525]" collectionCode="MACN-A" pageId="14" pageNumber="209">MACN-A 3454</specimenCode>
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(
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<figureCitation id="A0EB2A04FFB42866738FCF8BFDA4FDAE" box="[467,554,500,525]" captionStart="FIGURE 4" captionStartId="15.[151,249,1189,1211]" captionTargetBox="[151,1431,217,1130]" captionTargetId="figure@15.[151,1436,193,1167]" captionTargetPageId="15" captionText="FIGURE 4. Icochilus robustus, holotype MACN-A 3454, skull, in dorsal view (A); reproduction of figure 97 (B) from Ameghino (1891). Icochilus senilis, holotype MACN-A 3596, incomplete skull, in ventral view (C); holotype MACN-A 3597, incomplete mandible, in dorsal view (D). Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617765/files/figure.png" pageId="14" pageNumber="209">Fig. 4A</figureCitation>
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) to
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<specimenCode id="68769EFAFFB42866700BCF8BFC85FDAF" box="[599,779,500,525]" collectionCode="MACN-A" pageId="14" pageNumber="209">MACN-A 3505</specimenCode>
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, complete skull and mandible of the same individual, with associated postcranial elements.
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</materialsCitation>
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</paragraph>
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<paragraph id="386F3681FFB42866729BCC43FEAFFDDA" blockId="14.[151,1437,500,1101]" pageId="14" pageNumber="209">
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<emphasis id="0AA4EA93FFB42866729BCC43FD5DFDF7" bold="true" box="[199,723,572,597]" pageId="14" pageNumber="209">
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Age and provenance of the
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<materialsCitation id="88B83CDCFFB42866707BCC43FD5DFDF7" ID-GBIF-Occurrence="2147525208" box="[551,723,572,597]" collectionCode="SALMA" country="Argentina" location="Miocene" pageId="14" pageNumber="209" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
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<typeStatus id="E76B8823FFB42866707BCC43FDD4FDF7" box="[551,602,572,597]" pageId="14" pageNumber="209">type</typeStatus>
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material.
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</materialsCitation>
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</emphasis>
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Santacrucian SALMA, early
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<location id="3D0F605AFFB42866766ACC43FB15FDF7" LSID="urn:lsid:plazi:treatment:B0798797FFB4286672CBCFD2FE5CFBEE:3D0F605AFFB42866766ACC43FB15FDF7" box="[1078,1179,572,597]" country="Argentina" name="Miocene" pageId="14" pageNumber="209" stateProvince="Santa Cruz">Miocene</location>
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,
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<collectingRegion id="FA14F863FFB4286676F7CC43FA12FDF7" box="[1195,1436,572,597]" country="Argentina" name="Santa Cruz" pageId="14" pageNumber="209">Santa Cruz Province</collectingRegion>
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(
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<collectingCountry id="40C77611FFB4286672C2CC20FE9AFDDA" box="[158,276,607,632]" name="Argentina" pageId="14" pageNumber="209">Argentina</collectingCountry>
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).
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</paragraph>
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<paragraph id="386F3681FFB42866729BCCFBFBB2FD62" blockId="14.[151,1437,500,1101]" pageId="14" pageNumber="209">
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<emphasis id="0AA4EA93FFB42866729BCCFBFE37FD3F" bold="true" box="[199,441,644,669]" pageId="14" pageNumber="209">Original description</emphasis>
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(
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<bibRefCitation id="5C414B70FFB428667396CCFBFD4EFD3E" author="Ameghino, F." box="[458,704,644,669]" pageId="14" pageNumber="209" pagination="289 - 328" refId="ref16209" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article" year="1891">Ameghino 1891: 393</bibRefCitation>
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). “
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<emphasis id="0AA4EA93FFB4286670BDCCFBFC07FD3F" box="[737,905,644,669]" italics="true" pageId="14" pageNumber="209">Del tamaño de</emphasis>
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<taxonomicName id="FFD04D02FFB4286671CCCCFBFBD1FD3E" authorityName="Ameghino" authorityYear="1889" box="[912,1119,644,669]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="extensus">Icochilus extensus</taxonomicName>
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,
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<emphasis id="0AA4EA93FFB428667633CCFBFBA7FD62" italics="true" pageId="14" pageNumber="209">del que se distingue por el cráneo más ancho y por los frontales que se prolongan adelante entre los nasales</emphasis>
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”.
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</paragraph>
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<paragraph id="386F3681FFB42866729BCCB3FCFBFCAA" blockId="14.[151,1437,500,1101]" pageId="14" pageNumber="209">
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English translation. Of the same size as
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<taxonomicName id="FFD04D02FFB4286670FCCCB3FCFEFD46" authorityName="Ameghino" authorityYear="1889" box="[672,880,716,741]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="extensus">
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<emphasis id="0AA4EA93FFB4286670FCCCB3FCFEFD46" box="[672,880,716,741]" italics="true" pageId="14" pageNumber="209">Icochilus extensus</emphasis>
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</taxonomicName>
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, from which it is distinguished by presenting a wider skull and the frontals extended forward between the nasals.
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFB42866729BCD6BFE5CFBEE" pageId="14" pageNumber="209" type="discussion">
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<paragraph id="386F3681FFB42866729BCD6BFAA9FCF2" blockId="14.[151,1437,500,1101]" pageId="14" pageNumber="209">
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<emphasis id="0AA4EA93FFB42866729BCD6BFEC1FC8F" bold="true" box="[199,335,788,813]" pageId="14" pageNumber="209">Comments.</emphasis>
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<bibRefCitation id="5C414B70FFB42866730ACD6BFD6CFC8F" author="Ameghino, F." box="[342,738,788,813]" pageId="14" pageNumber="209" pagination="289 - 328" refId="ref16209" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article" year="1891">Ameghino (1891: 393, figs. 95–97)</bibRefCitation>
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briefly described this species in a footnote, stating that he had the entire skeleton of an individual. He also illustrated a foot, a right hand and an almost complete skull.
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</paragraph>
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<paragraph id="386F3681FFB42866729BCD23FE5CFBEE" blockId="14.[151,1437,500,1101]" pageId="14" pageNumber="209">
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According to Ameghino’s catalogue and
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<bibRefCitation id="5C414B70FFB4286670EBCD22FCD0FCD7" author="Mones A." box="[695,862,860,885]" pageId="14" pageNumber="209" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
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, MACN-A 3454 to MACN-A 3505, an almost complete individual, constitute the type specimen of the species. MACN-A 3454 is indeed the illustrated complete skull (
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<bibRefCitation id="5C414B70FFB428667280CDDBFE61FC1E" author="Ameghino, F." box="[220,495,932,957]" pageId="14" pageNumber="209" pagination="289 - 328" refId="ref16209" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article" year="1891">Ameghino 1891: fig. 97</bibRefCitation>
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), herein reproduced (
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<figureCitation id="A0EB2A04FFB428667082CDDBFCB3FC1E" box="[734,829,932,957]" captionStart="FIGURE 4" captionStartId="15.[151,249,1189,1211]" captionTargetBox="[151,1431,217,1130]" captionTargetId="figure@15.[151,1436,193,1167]" captionTargetPageId="15" captionText="FIGURE 4. Icochilus robustus, holotype MACN-A 3454, skull, in dorsal view (A); reproduction of figure 97 (B) from Ameghino (1891). Icochilus senilis, holotype MACN-A 3596, incomplete skull, in ventral view (C); holotype MACN-A 3597, incomplete mandible, in dorsal view (D). Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617765/files/figure.png" pageId="14" pageNumber="209">Fig. 4 B</figureCitation>
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), although it has suffered many breaks. Regarding the right manus and foot, only some elements (basipodia, metapodia and phalanges) are present. At the beginning of the current study, all of these materials were in different uncovered containers, so it is assumed that different elements have been missed over the years. Therefore, MACN 3454 to MACN-A 3505 are confirmed as elements of the
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<typeStatus id="E76B8823FFB42866729CCA4BFEACFBEF" box="[192,290,1076,1101]" pageId="14" pageNumber="209" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="FFD04D02FFB42866731BCA4AFE44FBEF" authorityName="Ameghino" authorityYear="1891" box="[327,458,1076,1101]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="robustus">
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<emphasis id="0AA4EA93FFB42866731BCA4AFE44FBEF" box="[327,458,1076,1101]" italics="true" pageId="14" pageNumber="209">Ic. robustus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |