160 lines
19 KiB
XML
160 lines
19 KiB
XML
<document id="522384F60F954B8EBC369E599555F6A5" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos & Cerdeño, Esperanza" docDate="2019" docId="B0798797FFAF287D72CBCD39FC77F93B" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Icochilus ulter Ameghino 1899" docType="treatment" docVersion="8" lastPageNumber="216" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="216" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="6DC214182DCD9621EF7E9459FD89CB8D">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="7259807A9D33DF931094F3FDC565BB5A">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="D6DAAE675F3FBABE0887B71BC0DDC624">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="BA2EAA3CE912F876ADA2D293C823F5FD">Cerdeño, Esperanza</mods:namePart>
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<mods:title id="6DF3FDCE0C3FFDB7BEDA23606F95E75D">Zootaxa</mods:title>
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<mods:date id="57D4E8B0BAF399C0B213BABA99ED4CF8">2019</mods:date>
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<mods:number id="35769CD70F58326A5871701FDEEB2200">2019-01-07</mods:number>
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<treatment id="B0798797FFAF287D72CBCD39FC77F93B" ID-DOI="http://doi.org/10.5281/zenodo.5931315" ID-GBIF-Taxon="154529766" ID-Zenodo-Dep="5931315" LSID="urn:lsid:plazi:treatment:B0798797FFAF287D72CBCD39FC77F93B" httpUri="http://treatment.plazi.org/id/B0798797FFAF287D72CBCD39FC77F93B" lastPageNumber="216" pageId="21" pageNumber="216">
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<subSubSection id="70CA650AFFAF287D72CBCD39FD97FCC2" box="[151,537,838,864]" pageId="21" pageNumber="216" type="nomenclature">
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<paragraph id="386F3681FFAF287D72CBCD39FD97FCC2" blockId="21.[151,537,838,864]" box="[151,537,838,864]" pageId="21" pageNumber="216">
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<heading id="632781EDFFAF287D72CBCD39FD97FCC2" bold="true" box="[151,537,838,864]" fontSize="11" level="1" pageId="21" pageNumber="216" reason="1">
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<taxonomicName id="FFD04D02FFAF287D72CBCD39FD97FCC2" authority="Ameghino, 1899" authorityName="Ameghino" authorityYear="1899" box="[151,537,838,864]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="ulter">
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<emphasis id="0AA4EA93FFAF287D72CBCD39FD97FCC2" bold="true" box="[151,537,838,864]" pageId="21" pageNumber="216">
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<emphasis id="0AA4EA93FFAF287D72CBCD39FEC9FCC2" bold="true" box="[151,327,838,864]" italics="true" pageId="21" pageNumber="216">Icochilus ulter</emphasis>
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<bibRefCitation id="5C414B70FFAF287D7313CD39FD97FCC2" author="Ameghino, F." box="[335,537,838,864]" pageId="21" pageNumber="216" pagination="1 - 13" refId="ref16324" refString="Ameghino, F. (1899) Sinopsis geologico-paleontologica de la Argentina. Suplemento (Adiciones y correcciones). Imprenta La Libertad, La Plata, pp. 1 - 13." type="book chapter" year="1899">Ameghino, 1899</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFAF287D72CBCDF2FE1BFB00" pageId="21" pageNumber="216" type="materials_examined">
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<paragraph id="386F3681FFAF287D72CBCDF2FEE2FC68" blockId="21.[151,1437,909,1690]" pageId="21" pageNumber="216">
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<materialsCitation id="88B83CDCFFAF287D72CBCDF2FEE2FC68" ID-GBIF-Occurrence="2147525030" collectionCode="MACN-A" pageId="21" pageNumber="216" specimenCode="MACN-A 11601" specimenCount="1" typeStatus="holotype">
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<emphasis id="0AA4EA93FFAF287D72CBCDF2FE85FC04" bold="true" box="[151,267,909,934]" pageId="21" pageNumber="216">
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<typeStatus id="E76B8823FFAF287D72CBCDF2FE88FC04" box="[151,262,909,934]" pageId="21" pageNumber="216" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="68769EFAFFAF287D7348CDF1FE5AFC04" box="[276,468,909,934]" collectionCode="MACN-A" pageId="21" pageNumber="216">MACN-A 11601</specimenCode>
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(
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<figureCitation id="A0EB2A04FFAF287D73BACDF2FDB8FC04" box="[486,566,909,934]" captionStart="FIGURE 6" captionStartId="19.[151,249,1490,1512]" captionTargetBox="[160,1405,212,1447]" captionTargetId="figure@19.[151,1436,193,1469]" captionTargetPageId="19" captionText="FIGURE 6. Icochilus multidentatus, lectotype MACN-A 3637, right maxilla with alveolus of extra tooth and C and dP1–M3, in occlusal view (A); paralectotype MACN-A 3638, right mandibular fragment with dp1–4 and m1–3, in oclusal (B) and lingual (C) views. Icochilus hegetotheroides, holotype MACN-A 9855, incomplete skull, anterior view of the premaxilla with labial view of both I1 (D) and right lateral view (E); holotype MACN-A 9856, incomplete mandible, in occlusal (F) and left lateral (G) views. Icochilus ulter, holotype MACN-A 11601, right mandibular fragment with p3–m3, in oclusal (H) and labial (I) views. Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617769/files/figure.png" pageId="21" pageNumber="216">Figs. 6</figureCitation>
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H–I), right mandibular fragment with p3–m3 and left mandibular fragment with p4–m1 series.
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</materialsCitation>
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</paragraph>
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<paragraph id="386F3681FFAF287D729BCDAAFD33FBB0" blockId="21.[151,1437,909,1690]" pageId="21" pageNumber="216">
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<emphasis id="0AA4EA93FFAF287D729BCDAAFD43FC4C" bold="true" box="[199,717,981,1006]" pageId="21" pageNumber="216">
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Age and provenance of the
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<materialsCitation id="88B83CDCFFAF287D707ECDAAFD43FC4C" ID-GBIF-Occurrence="2147524934" box="[546,717,981,1006]" country="Argentina" county="Miocene" location="Astrapothericulan" municipality="Pinturas Formation" pageId="21" pageNumber="216" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
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<typeStatus id="E76B8823FFAF287D707ECDAAFDD8FC4C" box="[546,598,981,1006]" pageId="21" pageNumber="216">type</typeStatus>
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material.
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</materialsCitation>
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</emphasis>
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<location id="3D0F605AFFAF287D7084CDAAFC2FFC4C" LSID="urn:lsid:plazi:treatment:B0798797FFAF287D72CBCD39FC77F93B:3D0F605AFFAF287D7084CDAAFC2FFC4C" box="[728,929,981,1006]" country="Argentina" county="Miocene" municipality="Pinturas Formation" name="Astrapothericulan" pageId="21" pageNumber="216" stateProvince="Santa Cruz">Astrapothericulan</location>
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beds (presumably the
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<collectingMunicipality id="D80BACFBFFAF287D76ECCDAAFA1EFC4C" box="[1200,1424,981,1006]" pageId="21" pageNumber="216">Pinturas Formation</collectingMunicipality>
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), early
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<collectingCounty id="D10E4E0DFFAF287D728ACD86FEB5FBB0" box="[214,315,1017,1042]" pageId="21" pageNumber="216">Miocene</collectingCounty>
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,
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<collectingRegion id="FA14F863FFAF287D731ACD85FDA2FBB0" box="[326,556,1017,1042]" country="Argentina" name="Santa Cruz" pageId="21" pageNumber="216">Santa Cruz Province</collectingRegion>
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(
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<collectingCountry id="40C77611FFAF287D7067CD86FD3FFBB0" box="[571,689,1017,1042]" name="Argentina" pageId="21" pageNumber="216">Argentina</collectingCountry>
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).
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</paragraph>
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<paragraph id="386F3681FFAF287D729BCA62FD3BFBF8" blockId="21.[151,1437,909,1690]" pageId="21" pageNumber="216">
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<emphasis id="0AA4EA93FFAF287D729BCA62FE37FB94" bold="true" box="[199,441,1053,1078]" pageId="21" pageNumber="216">Original description</emphasis>
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(
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<bibRefCitation id="5C414B70FFAF287D7394CA62FD10FB94" author="Ameghino, F." box="[456,670,1053,1078]" pageId="21" pageNumber="216" pagination="1 - 13" refId="ref16324" refString="Ameghino, F. (1899) Sinopsis geologico-paleontologica de la Argentina. Suplemento (Adiciones y correcciones). Imprenta La Libertad, La Plata, pp. 1 - 13." type="book chapter" year="1899">Ameghino 1899: 5</bibRefCitation>
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). “
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<emphasis id="0AA4EA93FFAF287D709DCA61FCECFB94" box="[705,866,1053,1078]" italics="true" pageId="21" pageNumber="216">De la talla del</emphasis>
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<taxonomicName id="FFD04D02FFAF287D7136CA62FBB6FB97" authorityName="Ameghino" authorityYear="1889" box="[874,1080,1053,1078]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="extensus">Icochilus extensus</taxonomicName>
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,
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<emphasis id="0AA4EA93FFAF287D761BCA61FD2FFBF8" italics="true" pageId="21" pageNumber="216">pero de rama mandibular más alta (20 milímetros debajo de la quinta muela)</emphasis>
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”.
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</paragraph>
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<paragraph id="386F3681FFAF287D729BCA1AFE1BFB00" blockId="21.[151,1437,909,1690]" pageId="21" pageNumber="216">
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English translation. With the size of
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<taxonomicName id="FFD04D02FFAF287D7034CA1AFCB8FBDF" authorityName="Ameghino" authorityYear="1889" box="[616,822,1125,1150]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="extensus">
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<emphasis id="0AA4EA93FFAF287D7034CA1AFCB8FBDF" box="[616,822,1125,1150]" italics="true" pageId="21" pageNumber="216">Icochilus extensus</emphasis>
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</taxonomicName>
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, but with a highest mandibular ramus (20 millimetres below the fifth molar).
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFAF287D729BCAD2FC77F93B" pageId="21" pageNumber="216" type="discussion">
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<paragraph id="386F3681FFAF287D729BCAD2FC77F93B" blockId="21.[151,1437,909,1690]" pageId="21" pageNumber="216">
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<emphasis id="0AA4EA93FFAF287D729BCAD2FEC1FB64" bold="true" box="[199,335,1197,1222]" pageId="21" pageNumber="216">Comments.</emphasis>
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<bibRefCitation id="5C414B70FFAF287D730ACAD2FD90FB64" author="Ameghino, F." box="[342,542,1197,1222]" pageId="21" pageNumber="216" pagination="1 - 13" refId="ref16324" refString="Ameghino, F. (1899) Sinopsis geologico-paleontologica de la Argentina. Suplemento (Adiciones y correcciones). Imprenta La Libertad, La Plata, pp. 1 - 13." type="book chapter" year="1899">Ameghino (1899)</bibRefCitation>
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briefly described this species without illustrating it, but he did not mention it in his catalogue. There is no specimen indicated as type of
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<taxonomicName id="FFD04D02FFAF287D7148CAADFCE3FB48" authorityName="Ameghino" authorityYear="1899" box="[788,877,1233,1258]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="ulter">
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<emphasis id="0AA4EA93FFAF287D7148CAADFCE3FB48" box="[788,877,1233,1258]" italics="true" pageId="21" pageNumber="216">Ic. ulter</emphasis>
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</taxonomicName>
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in the Ameghino Collection. Despite this,
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<bibRefCitation id="5C414B70FFAF287D770CCAADFF6FFAAC" author="Mones A." pageId="21" pageNumber="216" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
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indicated, but with doubts, “MACN-A 10601?” as type specimen; however, this specimen is also indicated by the author as the type of the isotemnid
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<taxonomicName id="FFD04D02FFAF287D7037CB66FC3EFA90" authorityName="Ameghino" authorityYear="1901" box="[619,944,1305,1330]" class="Mammalia" family="Isotemnidae" genus="Eochalicotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="crassidens">
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<emphasis id="0AA4EA93FFAF287D7037CB66FC3EFA90" box="[619,944,1305,1330]" italics="true" pageId="21" pageNumber="216">Eochalicotherium crassidens</emphasis>
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</taxonomicName>
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. MACN-A 10601 is in fact a lot constituted by three mandibular fragments—with one tooth each—that belong to the latter species; consequently, it is discarded as the
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<typeStatus id="E76B8823FFAF287D7300CB1EFE30FAD8" box="[348,446,1377,1402]" pageId="21" pageNumber="216" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="FFD04D02FFAF287D73B7CB1DFDC8FAD8" authorityName="Ameghino" authorityYear="1899" box="[491,582,1377,1402]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="ulter">
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<emphasis id="0AA4EA93FFAF287D73B7CB1DFDC8FAD8" box="[491,582,1377,1402]" italics="true" pageId="21" pageNumber="216">Ic. ulter</emphasis>
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</taxonomicName>
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. On the other hand, the lot MACN-A 11601, which is composed of two mandibular fragments of the same individual, with right p3–m3 and left p4–m1, is the only material assigned by F. Ameghino to
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<taxonomicName id="FFD04D02FFAF287D736ECBD5FE05FA60" authorityName="Ameghino" authorityYear="1899" box="[306,395,1449,1474]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="ulter">
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<emphasis id="0AA4EA93FFAF287D736ECBD5FE05FA60" box="[306,395,1449,1474]" italics="true" pageId="21" pageNumber="216">Ic. ulter</emphasis>
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</taxonomicName>
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within the Ameghino Collection. The right fragment matches the only measurement (length of the mandibular ramus below m1 =
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<quantity id="FF289B64FFAF287D7061CBB1FD02FA44" box="[573,652,1486,1510]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="216" unit="mm" value="20.0">20 mm</quantity>
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) given by
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<bibRefCitation id="5C414B70FFAF287D715FCBB2FC40FA44" author="Ameghino, F." box="[771,974,1485,1510]" pageId="21" pageNumber="216" pagination="1 - 13" refId="ref16324" refString="Ameghino, F. (1899) Sinopsis geologico-paleontologica de la Argentina. Suplemento (Adiciones y correcciones). Imprenta La Libertad, La Plata, pp. 1 - 13." type="book chapter" year="1899">Ameghino (1899)</bibRefCitation>
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. It is worth mentioning that there are two pieces of paper inside the container with Ameghino’s handwriting: one that reproduces the original description of the species and another with the name “
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<taxonomicName id="FFD04D02FFAF287D7000C86AFC8EF98C" authorityName="Ameghino" authorityYear="1899" box="[604,768,1557,1582]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="ulter">
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<emphasis id="0AA4EA93FFAF287D7000C86AFC8EF98C" box="[604,768,1557,1582]" italics="true" pageId="21" pageNumber="216">Icochilus ulter</emphasis>
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</taxonomicName>
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”. In the latter, it is also written “10601” but with someone else’s handwriting (probably from M. Soria; A. Kramarz, pers. comm.), and the first ‘0’ was crossed out and replaced by ‘1’; it is very likely that this mistake led Mones to mention “MACN-A 10601?” as the type of the species. As a result, MACN-A 11601 is identified as the
|
||
<typeStatus id="E76B8823FFAF287D714FC8FEFCFBF938" box="[787,885,1665,1690]" pageId="21" pageNumber="216" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="FFD04D02FFAF287D71C7C8FDFC7FF938" authorityName="Ameghino" authorityYear="1899" box="[923,1009,1665,1690]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="ulter">
|
||
<emphasis id="0AA4EA93FFAF287D71C7C8FDFC7FF938" box="[923,1009,1665,1690]" italics="true" pageId="21" pageNumber="216">Ic. ulter</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |