184 lines
20 KiB
XML
184 lines
20 KiB
XML
<document id="D3AEF3A0D8DF893D3DCEBE757CFC172A" ID-CLB-Dataset="27706" ID-DOI="10.11646/zootaxa.4543.2.2" ID-GBIF-Dataset="7438332f-a925-43aa-b17f-059781bdfc12" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617757" ID-ZooBank="11F546CD-109F-4CBC-B87D-EA8D1AD0B96F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928261775" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos & Cerdeño, Esperanza" docDate="2019" docId="B0798797FFAE287D72CBC8B9FD6BFD50" docLanguage="en" docName="zootaxa.4543.2.2.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Icochilus hegetotheroides Ameghino 1894" docType="treatment" docVersion="8" lastPageNumber="216" masterDocId="4C40FFEFFFBA2868725CCE7FFF8EFFA2" masterDocTitle="On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina" masterLastPageNumber="220" masterPageNumber="195" pageNumber="215" updateTime="1701997211094" updateUser="plazi">
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<mods:title id="81504D5A24CFAE797A3BB6CB99C81544">On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina</mods:title>
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<mods:namePart id="1158E20963335200194D37D183E62501">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="35A47071B8F4881A82EE82FC7AC2C711">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="2F99A838701F3F700F39874AAA3422FE">Cerdeño, Esperanza</mods:namePart>
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<mods:title id="CAB36A68DF9E2D785A014E4ABA945B23">Zootaxa</mods:title>
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<mods:date id="55A19B35DF923490238035404A7DAE81">2019</mods:date>
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<mods:number id="FEC0F82438FDE97C8EAE23EB712015CE">2019-01-07</mods:number>
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<treatment id="B0798797FFAE287D72CBC8B9FD6BFD50" ID-DOI="http://doi.org/10.5281/zenodo.5931313" ID-GBIF-Taxon="154529767" ID-Zenodo-Dep="5931313" LSID="urn:lsid:plazi:treatment:B0798797FFAE287D72CBC8B9FD6BFD50" httpUri="http://treatment.plazi.org/id/B0798797FFAE287D72CBC8B9FD6BFD50" lastPageId="21" lastPageNumber="216" pageId="20" pageNumber="215">
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<subSubSection id="70CA650AFFAE287C72CBC8B9FD23F942" box="[151,685,1734,1760]" pageId="20" pageNumber="215" type="nomenclature">
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<paragraph id="386F3681FFAE287C72CBC8B9FD23F942" blockId="20.[151,685,1734,1760]" box="[151,685,1734,1760]" pageId="20" pageNumber="215">
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<heading id="632781EDFFAE287C72CBC8B9FD23F942" bold="true" box="[151,685,1734,1760]" fontSize="11" level="1" pageId="20" pageNumber="215" reason="1">
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<taxonomicName id="FFD04D02FFAE287C72CBC8B9FD23F942" authority="Ameghino, 1894 a" authorityName="Ameghino" authorityYear="1894" box="[151,685,1734,1760]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="215" phylum="Chordata" rank="species" species="hegetotheroides">
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<emphasis id="0AA4EA93FFAE287C72CBC8B9FD23F942" bold="true" box="[151,685,1734,1760]" pageId="20" pageNumber="215">
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<emphasis id="0AA4EA93FFAE287C72CBC8B9FE42F942" bold="true" box="[151,460,1734,1760]" italics="true" pageId="20" pageNumber="215">Icochilus hegetotheroides</emphasis>
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<bibRefCitation id="5C414B70FFAE287C7388C8B9FD23F942" author="Ameghino, F." box="[468,685,1734,1760]" pageId="20" pageNumber="215" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino, 1894a</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFAE287C72CBC972FA12F87E" pageId="20" pageNumber="215" type="materials_examined">
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<paragraph id="386F3681FFAE287C72CBC972FD45F8E8" blockId="20.[151,1436,1805,2013]" pageId="20" pageNumber="215">
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<materialsCitation id="88B83CDCFFAE287C72CBC972FD45F8E8" ID-GBIF-Occurrence="2147525201" collectionCode="MACN-A" pageId="20" pageNumber="215" specimenCode="MACN-A 9855, MACN-A 9856" specimenCount="1" typeStatus="holotype">
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<emphasis id="0AA4EA93FFAE287C72CBC972FE85F884" bold="true" box="[151,267,1805,1830]" pageId="20" pageNumber="215">
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<typeStatus id="E76B8823FFAE287C72CBC972FE88F884" box="[151,262,1805,1830]" pageId="20" pageNumber="215" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="68769EFAFFAE287C7344C971FE41F884" box="[280,463,1805,1830]" collectionCode="MACN-A" pageId="20" pageNumber="215">MACN-A 9855</specimenCode>
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(
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<figureCitation id="A0EB2A04FFAE287C73B9C972FDB6F884" box="[485,568,1805,1830]" captionStart="FIGURE 6" captionStartId="19.[151,249,1490,1512]" captionTargetBox="[160,1405,212,1447]" captionTargetId="figure@19.[151,1436,193,1469]" captionTargetPageId="19" captionText="FIGURE 6. Icochilus multidentatus, lectotype MACN-A 3637, right maxilla with alveolus of extra tooth and C and dP1–M3, in occlusal view (A); paralectotype MACN-A 3638, right mandibular fragment with dp1–4 and m1–3, in oclusal (B) and lingual (C) views. Icochilus hegetotheroides, holotype MACN-A 9855, incomplete skull, anterior view of the premaxilla with labial view of both I1 (D) and right lateral view (E); holotype MACN-A 9856, incomplete mandible, in occlusal (F) and left lateral (G) views. Icochilus ulter, holotype MACN-A 11601, right mandibular fragment with p3–m3, in oclusal (H) and labial (I) views. Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617769/files/figure.png" pageId="20" pageNumber="215">Figs. 6</figureCitation>
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D–E), anterior cranial fragment, and
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<specimenCode id="68769EFAFFAE287C7658C971FB35F884" box="[1028,1211,1805,1830]" collectionCode="MACN-A" pageId="20" pageNumber="215">MACN-A 9856</specimenCode>
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(
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<figureCitation id="A0EB2A04FFAE287C768DC972FAABF884" box="[1233,1317,1805,1830]" captionStart="FIGURE 6" captionStartId="19.[151,249,1490,1512]" captionTargetBox="[160,1405,212,1447]" captionTargetId="figure@19.[151,1436,193,1469]" captionTargetPageId="19" captionText="FIGURE 6. Icochilus multidentatus, lectotype MACN-A 3637, right maxilla with alveolus of extra tooth and C and dP1–M3, in occlusal view (A); paralectotype MACN-A 3638, right mandibular fragment with dp1–4 and m1–3, in oclusal (B) and lingual (C) views. Icochilus hegetotheroides, holotype MACN-A 9855, incomplete skull, anterior view of the premaxilla with labial view of both I1 (D) and right lateral view (E); holotype MACN-A 9856, incomplete mandible, in occlusal (F) and left lateral (G) views. Icochilus ulter, holotype MACN-A 11601, right mandibular fragment with p3–m3, in oclusal (H) and labial (I) views. Scale bar = 10 mm." httpUri="https://zenodo.org/record/2617769/files/figure.png" pageId="20" pageNumber="215">Figs. 6</figureCitation>
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F–G), an incomplete mandible, both of the same individual.
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</materialsCitation>
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</paragraph>
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<paragraph id="386F3681FFAE287C729BC92AFEAFF830" blockId="20.[151,1436,1805,2013]" pageId="20" pageNumber="215">
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<emphasis id="0AA4EA93FFAE287C729BC92AFD5DF8CC" bold="true" box="[199,723,1877,1902]" pageId="20" pageNumber="215">
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Age and provenance of the
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<materialsCitation id="88B83CDCFFAE287C707BC92AFD5DF8CC" ID-GBIF-Occurrence="2147525110" box="[551,723,1877,1902]" collectionCode="SALMA" country="Argentina" location="Miocene" pageId="20" pageNumber="215" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
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<typeStatus id="E76B8823FFAE287C707BC92AFDD4F8CC" box="[551,602,1877,1902]" pageId="20" pageNumber="215">type</typeStatus>
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material.
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</materialsCitation>
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</emphasis>
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Santacrucian SALMA, early
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<location id="3D0F605AFFAE287C766AC92AFB15F8CC" LSID="urn:lsid:plazi:treatment:B0798797FFAE287D72CBC8B9FD6BFD50:3D0F605AFFAE287C766AC92AFB15F8CC" box="[1078,1179,1877,1902]" country="Argentina" name="Miocene" pageId="20" pageNumber="215" stateProvince="Santa Cruz">Miocene</location>
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,
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<collectingRegion id="FA14F863FFAE287C76F7C929FA12F8CC" box="[1195,1436,1877,1902]" country="Argentina" name="Santa Cruz" pageId="20" pageNumber="215">Santa Cruz Province</collectingRegion>
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(
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<collectingCountry id="40C77611FFAE287C72C2C906FE9AF830" box="[158,276,1913,1938]" name="Argentina" pageId="20" pageNumber="215">Argentina</collectingCountry>
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).
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</paragraph>
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<paragraph id="386F3681FFAE287C729BC9E2FA12F87E" blockId="20.[151,1436,1805,2013]" pageId="20" pageNumber="215">
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<emphasis id="0AA4EA93FFAE287C729BC9E2FE32F814" bold="true" box="[199,444,1949,1974]" pageId="20" pageNumber="215">Original description</emphasis>
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(
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<bibRefCitation id="5C414B70FFAE287C7391C9E2FD4BF814" author="Ameghino, F." box="[461,709,1949,1974]" pageId="20" pageNumber="215" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino 1894a: 17</bibRefCitation>
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). “
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<emphasis id="0AA4EA93FFAE287C70B4C9E1FDEAF879" italics="true" pageId="20" pageNumber="215">
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Cette espèce se distingue par le grand développement de la paire d'incisives internes supérieures (
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<date id="4C6E1041FFAE287C7004C9BBFDEAF879" box="[600,612,1988,2011]" pageId="20" pageNumber="215">i.</date>
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</emphasis>
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<emphasis id="0AA4EA93FFAE287C7039C9BFFB20F87E" box="[613,1198,1984,2013]" italics="true" pageId="20" pageNumber="215">
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<superScript id="CFA59BC9FFAE287C7039C9BFFDE3F86C" attach="right" box="[613,621,1984,1998]" fontSize="6" pageId="20" pageNumber="215">1</superScript>
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), comparables sous ce rapport à celles du genre
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</emphasis>
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<taxonomicName id="FFD04D02FFAE287C76E4C9BBFAEEF87F" authorityName="Ameghino" authorityYear="1887" box="[1208,1376,1988,2013]" class="Mammalia" family="Hegetotheriidae" genus="Hegetotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="216" phylum="Chordata" rank="genus">Hegetotherium</taxonomicName>
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.
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<emphasis id="0AA4EA93FFAE287C772EC9BAFA12F87E" box="[1394,1436,1989,2012]" italics="true" pageId="20" pageNumber="215">Ces</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="70CA650AFFAF287D72CBCEE6FB27FDB8" pageId="21" pageNumber="216" type="description">
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<paragraph id="386F3681FFAF287D72CBCEE6FC33FEE1" blockId="21.[151,1437,151,755]" pageId="21" pageNumber="216">
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<emphasis id="0AA4EA93FFAF287D72CBCEE6FE1AFEE1" italics="true" pageId="21" pageNumber="216">
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incisives sont convexes en avant, concaves en arrière, et avec la surface de la couronne en arc de cercle. La deuxième incisive est petite, et la troisième incisive ainsi que la canine sont atrophiées. La partie antérieure du crâne est très raccourcie et proportionnellement très large. La couronne de chacune des incisives internes a
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<quantity id="FF289B64FFAF287D7704CE9FFA17FF5A" box="[1368,1433,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="216" unit="mm" value="5.0">5 mm</quantity>
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. 5 de diamètre transverse en ligne droite. Le palais au niveau des canines a
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<quantity id="FF289B64FFAF287D71A2CF7BFBC1FEBE" box="[1022,1103,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="21" pageNumber="216" unit="mm" value="13.0">13 mm</quantity>
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. de largeur, et la distance du bord antérieur de la p.
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</emphasis>
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||
<emphasis id="0AA4EA93FFAF287D73C8CF58FD1FFEE1" box="[404,657,295,323]" italics="true" pageId="21" pageNumber="216">
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||
<superScript id="CFA59BC9FFAF287D73C8CF58FE12FE97" attach="left" box="[404,412,295,309]" fontSize="6" pageId="21" pageNumber="216">1</superScript>
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au coin interne de l'i.
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</emphasis>
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||
<emphasis id="0AA4EA93FFAF287D70CDCF58FC24FEE1" box="[657,938,295,323]" italics="true" pageId="21" pageNumber="216">
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||
<superScript id="CFA59BC9FFAF287D70CDCF58FD17FE97" attach="left" box="[657,665,295,309]" fontSize="6" pageId="21" pageNumber="216">1</superScript>
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est de seulement
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<quantity id="FF289B64FFAF287D7102CF55FC24FEE1" box="[862,938,298,323]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="21" pageNumber="216" unit="mm" value="46.0">46 mm</quantity>
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</emphasis>
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||
”.
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</paragraph>
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<paragraph id="386F3681FFAF287D729BCF31FB27FDB8" blockId="21.[151,1437,151,755]" pageId="21" pageNumber="216">
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English translation. This species is distinguished by the great development of the internal upper incisors (
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<date id="4C6E1041FFAF287D7729CF30FA02FEC5" box="[1397,1420,335,359]" pageId="21" pageNumber="216">I1</date>
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), similar in this aspect to those of the genus
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<taxonomicName id="FFD04D02FFAF287D70DECF0DFCA4FE29" authorityName="Ameghino" authorityYear="1887" box="[642,810,370,395]" class="Mammalia" family="Hegetotheriidae" genus="Hegetotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="genus">
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<emphasis id="0AA4EA93FFAF287D70DECF0DFCA4FE29" box="[642,810,370,395]" italics="true" pageId="21" pageNumber="216">Hegetotherium</emphasis>
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</taxonomicName>
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. These incisors are labially convex, lingually concave, and with an arched surface of the crown. The second incisor is small, and the third incisor and the canine are atrophied. The anterior part of the skull is very short and proportionally very broad. The crown of each internal incisor measures
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<quantity id="FF289B64FFAF287D7301CFA1FE2AFE55" box="[349,420,478,503]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="216" unit="mm" value="5.0">5 mm</quantity>
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||
. 5 of transverse diameter in a straight line. The palate is
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||
<quantity id="FF289B64FFAF287D766BCFA0FB06FE55" box="[1079,1160,479,503]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="21" pageNumber="216" unit="mm" value="13.0">13 mm</quantity>
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||
wide at the level of the canines, and the distance from the anterior part of P1 to the internal part of
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<date id="4C6E1041FFAF287D71BECC7CFC77FDB9" box="[994,1017,515,539]" pageId="21" pageNumber="216">I1</date>
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||
is only
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||
<quantity id="FF289B64FFAF287D760ECC7DFB2BFDB8" box="[1106,1189,514,539]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="21" pageNumber="216" unit="mm" value="46.0">46 mm</quantity>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="70CA650AFFAF287D729BCC59FD6BFD50" pageId="21" pageNumber="216" type="discussion">
|
||
<paragraph id="386F3681FFAF287D729BCC59FD6BFD50" blockId="21.[151,1437,151,755]" pageId="21" pageNumber="216">
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<emphasis id="0AA4EA93FFAF287D729BCC59FEC1FD9D" bold="true" box="[199,335,550,575]" pageId="21" pageNumber="216">Comments.</emphasis>
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||
<bibRefCitation id="5C414B70FFAF287D7307CC59FDBAFD9D" author="Ameghino, F." box="[347,564,550,575]" pageId="21" pageNumber="216" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
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did not illustrate any specimen of this species. According to Ameghino’s catalogue and
|
||
<bibRefCitation id="5C414B70FFAF287D731ACC34FE62FDC1" author="Mones A." box="[326,492,586,611]" pageId="21" pageNumber="216" pagination="1 - 625" refId="ref17293" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article" year="1986">Mones (1986)</bibRefCitation>
|
||
, MACN-A 9855 (anterior portion of a skull) and MACN-A 9856 (incomplete mandible), which belong to the same individual, are the type specimens of the species. The measurements of MACN-A 9855 (MDL of
|
||
<date id="4C6E1041FFAF287D73E2CCECFE5BFD09" box="[446,469,659,683]" pageId="21" pageNumber="216">I1</date>
|
||
=
|
||
<quantity id="FF289B64FFAF287D73A8CCEDFDF1FD08" box="[500,639,658,683]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.7" metricValueMin="5.5" pageId="21" pageNumber="216" unit="mm" value="5.6" valueMax="5.7" valueMin="5.5">5.5–5.7 mm</quantity>
|
||
; width of the palate at the C level =
|
||
<quantity id="FF289B64FFAF287D7647CCECFB0DFD08" box="[1051,1155,659,683]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="21" pageNumber="216" unit="mm" value="13.2">13.2 mm</quantity>
|
||
; length of
|
||
<date id="4C6E1041FFAF287D76A7CCEDFA81FD09" box="[1275,1295,658,683]" pageId="21" pageNumber="216">I1</date>
|
||
–dP1 =
|
||
<quantity id="FF289B64FFAF287D7737CCEDFF4AFD6D" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="21" pageNumber="216" unit="mm" value="16.3">16.3 mm</quantity>
|
||
) match approximately those provided by
|
||
<bibRefCitation id="5C414B70FFAF287D70C2CCC9FCF8FD6D" author="Ameghino, F." box="[670,886,694,719]" pageId="21" pageNumber="216" refId="ref16256" refString="Ameghino, F. (1894 a) Enumeration synoptique des especes de mammiferes fossiles des formations eocenes de Patagonie. Coni e hijos, Buenos Aires, 196 pp." type="book" year="1894">Ameghino (1894a)</bibRefCitation>
|
||
. Therefore, MACN-A 9855 and MACN-A 9856 are confirmed as the
|
||
<typeStatus id="E76B8823FFAF287D73DECCA5FE6AFD51" box="[386,484,730,755]" pageId="21" pageNumber="216" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="FFD04D02FFAF287D7055CCA3FD50FD51" authorityName="Ameghino" authorityYear="1894" box="[521,734,730,755]" class="Mammalia" family="Interatheriidae" genus="Icochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="21" pageNumber="216" phylum="Chordata" rank="species" species="hegetotheroides">
|
||
<emphasis id="0AA4EA93FFAF287D7055CCA3FD50FD51" box="[521,734,730,755]" italics="true" pageId="21" pageNumber="216">Ic. hegetotheroides</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |