822 lines
92 KiB
XML
822 lines
92 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.99.107974" ID-Pensoft-Pub="1860-0743-2-557" ID-Pensoft-UUID="1F2637E73C8459FB8AC4165BA9D0DBAF" ID-ZooBank="1549ABF34A14448B89AF549F2088D947" ModsDocID="1860-0743-99-2-557" checkinTime="1701752218675" checkinUser="pensoft" docAuthor="Katnoum, Chanikan, Keetapithchayakul, Tosaphol Saetung, Rahim, Azman Abdul & Wongkamhaeng, Koraon" docDate="2023" docId="ADFF785E4AB25672B2F53B6336FBD91E" docLanguage="en" docName="ZoosystEvol 99(2): 557-574" docOrigin="Zoosystematics and Evolution 99 (2)" docPubDate="2023-12-04" docSource="http://dx.doi.org/10.3897/zse.99.107974" docTitle="Cerapus rivulus Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng 2023, sp. nov." docType="treatment" docUuid="3074048C-ECA2-499C-A3D4-C54C6DAECB7C" docUuidSource="ZooBank" docVersion="1" id="1F2637E73C8459FB8AC4165BA9D0DBAF" lastPageNumber="557" masterDocId="1F2637E73C8459FB8AC4165BA9D0DBAF" masterDocTitle="A new species of Cerapus (Amphipoda, Senticaudata, Ischyroceridae) from Mae Klong Estuary, with a discussion on their nesting and types of mating behaviour" masterLastPageNumber="574" masterPageNumber="557" pageNumber="557" updateTime="1701752218675" updateUser="pensoft">
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<mods:title id="EBA49949DD148C158AD66A29CDB2F976">A new species of Cerapus (Amphipoda, Senticaudata, Ischyroceridae) from Mae Klong Estuary, with a discussion on their nesting and types of mating behaviour</mods:title>
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<mods:name id="373912E9A3F139E7AA22E1C9B804FA82" type="personal">
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<mods:roleTerm id="C787E80A8DF2485D60469B342B5FBD39">Author</mods:roleTerm>
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<mods:namePart id="CB216C28688E5776C902501CBFDE8D41">Katnoum, Chanikan</mods:namePart>
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<mods:affiliation id="2DF64597751B8BCE82901C16DB3FEB7D">Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand & Biodiversity Center Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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<mods:name id="CBA639234461670A66A0194849D66F83" type="personal">
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<mods:roleTerm id="FB7C22E69AA55245EDE2546D84309379">Author</mods:roleTerm>
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<mods:namePart id="D226CE58330ADB54E3FB664998391F33">Keetapithchayakul, Tosaphol Saetung</mods:namePart>
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<mods:nameIdentifier id="66022B044B72D5EE4D9D54180554D9EF" type="ORCID">https://orcid.org/0000-0001-7565-4701</mods:nameIdentifier>
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<mods:affiliation id="8EFCDDAD65BA0B8E9F081FEFB16E8875">Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="2FB09372F227A9CF4AF5CC7565DA9001" type="personal">
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<mods:role id="4E26BD7B7591F897F166F169AA2AEF63">
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<mods:roleTerm id="53F7857FD4FEAFA4CC29CF7915B7A2D1">Author</mods:roleTerm>
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<mods:namePart id="168352E3A3BFD15A12E0ADCE83009BDC">Rahim, Azman Abdul</mods:namePart>
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<mods:nameIdentifier id="EC35A7842F5011F3B8AC6F08124B6C50" type="ORCID">https://orcid.org/0000-0002-8964-6638</mods:nameIdentifier>
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<mods:affiliation id="000332766A7ADB6855C3967F73191FE8">Marine Ecosystem Research Centre, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, 43600 UKM Bangi, Selangor, Malaysia</mods:affiliation>
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</mods:name>
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<mods:name id="489DB63EC775EFA5477E6A077E58861C" type="personal">
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<mods:roleTerm id="643176E90149717DD4C289655CD24F70">Author</mods:roleTerm>
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<mods:namePart id="0AECB56CF142C0E03A88FEBCD1BAFBF3">Wongkamhaeng, Koraon</mods:namePart>
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<mods:nameIdentifier id="7CB49704DE57666397A7921BEFA3132F" type="ORCID">https://orcid.org/0000-0001-7671-8869</mods:nameIdentifier>
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<mods:affiliation id="0BF8AC1850557FAB1B28D1A2B0F518C8">Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand & Biodiversity Center Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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<mods:nameIdentifier id="D7A94ECD9CA9C37DD187F11C7B40FAF6" type="email">koraon@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="89A41DBBBFDF1DFEEF3D28A89056984A">text</mods:typeOfResource>
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<mods:title id="F2B62605BAF8ADA3563632F73B882CFB">Zoosystematics and Evolution</mods:title>
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<mods:part id="E1F585821080E73DC212CA47ECAABA7C">
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<mods:date id="3B9F2BEC9BB0DED75499126A3F032B2E">2023</mods:date>
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<mods:number id="ADB1FE5783344B98F0447EE7B2658716">2023-12-04</mods:number>
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<mods:number id="5D756D2EC27F3DF84AE24AC16E75DA48">99</mods:number>
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<mods:start id="CF26B54BD1712F5D4DA9C2AAB88CF37B">557</mods:start>
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<mods:url id="D8130EA650ACFC9F08F6935CC2A85B77">http://dx.doi.org/10.3897/zse.99.107974</mods:url>
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<mods:classification id="96206D5BD66B221B7C4FC1660C077371">journal article</mods:classification>
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<mods:identifier id="2A86624B4CDA92CAD6E8E6B34042E7EA" type="DOI">http://dx.doi.org/10.3897/zse.99.107974</mods:identifier>
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<treatment id="ADFF785E4AB25672B2F53B6336FBD91E" LSID="urn:lsid:zoobank.org:act:3074048C-ECA2-499C-A3D4-C54C6DAECB7C" httpUri="http://treatment.plazi.org/id/ADFF785E4AB25672B2F53B6336FBD91E" lastPageNumber="557" pageId="0" pageNumber="557">
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<subSubSection id="E2BDED39F390D215D288D7393772AA3E" pageId="0" pageNumber="557" type="nomenclature">
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<paragraph id="D8761393566B88B351BF662EE5AB562C" pageId="0" pageNumber="557">
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<taxonomicName id="659CDA0E9311B4A953BDBF3C20709771" LSID="https://zoobank.org/3074048C-ECA2-499C-A3D4-C54C6DAECB7C" authority="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng, 2023" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus" status="sp. nov.">Cerapus rivulus</taxonomicName>
|
||
<taxonomicNameLabel id="0A15162A41671445DB622D67D8DF89B9" pageId="0" pageNumber="557">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="358E29E40174F85686359A60D900800A" pageId="0" pageNumber="557" type="description">
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<paragraph id="0E586ABF3C5C89BD92DF2AE136494A50" pageId="0" pageNumber="557">
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||
<figureCitation id="B14EEC01103892EC64EA8599B4F4D46F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerapus rivulus sp. nov. A. Holotype male lateral (THNHM - 19377); B. A specimen in its tube with protruding head and antennae; C. Paratype female front (THNHM- 19379)." figureDoi="10.3897/zse.99.107974.figure2" httpUri="https://binary.pensoft.net/fig/947231" pageId="0" pageNumber="557">Figs 2</figureCitation>
|
||
<figureCitation id="31264FD7F31882867138E09F89F58A26" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Habitus of the male holotype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure3" httpUri="https://binary.pensoft.net/fig/947232" pageId="0" pageNumber="557">, 3</figureCitation>
|
||
<figureCitation id="BE2E58E807C2CC4D88E757FB55AD0974" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pereopod 3; B. Pereopod 4; C. Pereopod 5; D. Pereopod 6; E. Pereopod 7. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure4" httpUri="https://binary.pensoft.net/fig/947233" pageId="0" pageNumber="557">, 4</figureCitation>
|
||
<figureCitation id="66F85F2AA71CA4372A913E54C3E0C2D6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">, 5</figureCitation>
|
||
<figureCitation id="9F70F5D1DA52EE879B702E5E610EE112" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Left mandible; B. Right mandible; C. Maxilliped; D. Maxilla 2; E. Maxilla 1; F. Upper lip; G. Lower lip. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure6" httpUri="https://binary.pensoft.net/fig/947235" pageId="0" pageNumber="557">, 6</figureCitation>
|
||
<figureCitation id="583675060B31CA2DE925C0A2CB419765" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Habitus of the female paratype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure7" httpUri="https://binary.pensoft.net/fig/947236" pageId="0" pageNumber="557">, 7</figureCitation>
|
||
<figureCitation id="3D800A315B2B28AEE6AA8968A0896211" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Pereopod 3; B. Pereopod 4; C. Pereopod 5; D. Pereopod 6; E. Pereopod 7. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure8" httpUri="https://binary.pensoft.net/fig/947237" pageId="0" pageNumber="557">, 8</figureCitation>
|
||
<figureCitation id="BDD83DCA96A3959B37E7125098F740EC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson. Scale bars: 0.1 mm (A-C); 0.2 mm (D)." figureDoi="10.3897/zse.99.107974.figure9" httpUri="https://binary.pensoft.net/fig/947238" pageId="0" pageNumber="557">, 9</figureCitation>
|
||
<figureCitation id="714C10C90C6AD4D8BB8DAA6070038CD0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Maxilliped; B. Maxilla 2; C. Maxilla 1; D. Upper lip; E. Lower lip; F. Left mandible; G. Right mandible. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure10" httpUri="https://binary.pensoft.net/fig/947239" pageId="0" pageNumber="557">, 10</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39DF2F31EB8CEC13273FB690FFB489F8" pageId="0" pageNumber="557" type="materials_examined">
|
||
<paragraph id="FE4B385C2BCB81447626A42A35FD42B6" pageId="0" pageNumber="557">Material examined.</paragraph>
|
||
<paragraph id="3069C8E16F703516DE71BD735B168707" pageId="0" pageNumber="557">
|
||
<materialsCitation id="A6A705A4F23892AB294FA1355C38C208" collectingDate="2021-04-10" collectorName="C. Katnoum" country="THAILAND" elevation="5" latitude="13.494722" location="Darunanukroh School" longLatPrecision="1" longitude="99.92369" municipality="Mae Klong River" specimenCount="♂" stateProvince="Samut Songkhram Province" typeStatus="Holotype">
|
||
<emphasis id="68D2C560BF55C5895E65E87C75DD26F5" bold="true" pageId="0" pageNumber="557">
|
||
<emphasis id="418260737C3B52339605C71679669ECF" italics="true" pageId="0" pageNumber="557">
|
||
<typeStatus id="B2570ECE9598D321970DD515CA91F4C5">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="262BA46534ADDA68EB7B3CE079AB9E1D" name="Thailand">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="D329CC2A305FAC3EED149899BCD6A240">♂</specimenCount>
|
||
,
|
||
<quantity id="16C880764AAB4C916E2C4A9068967F54" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
|
||
;
|
||
<collectingRegion id="C37AE3D76C0AA33DD9349533EC895C83" country="Thailand" name="Samut Songkhram">Samut Songkhram Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="8AECEE25CCCFBB4B211B2ACEB5313AAE">Mae Klong River</collectingMunicipality>
|
||
,
|
||
<location id="3E3EF0BD3F8930C7FB85477F58993027" LSID="urn:lsid:plazi:treatment:ADFF785E4AB25672B2F53B6336FBD91E:3E3EF0BD3F8930C7FB85477F58993027" country="THAILAND" latitude="13.494722" longLatPrecision="1" longitude="99.92369" municipality="Mae Klong River" name="Darunanukroh School" stateProvince="Samut Songkhram Province">Darunanukroh School</location>
|
||
;
|
||
<geoCoordinate id="4DB104E70843DB1F338707C24E2D82FD" degrees="13" direction="north" minutes="29" orientation="latitude" precision="1" seconds="41.0" value="13.494722">13°29'41.0"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E61768A4E5FF1FE8E3AEA09466135C05" degrees="99" direction="east" minutes="55" orientation="longitude" precision="1" seconds="25.3" value="99.92369">99°55'25.3"E</geoCoordinate>
|
||
;
|
||
<elevation id="038CF4AC21F23C032C8E969BE8F6823D" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">
|
||
<quantity id="A22892D7BF67D7B4616C5DA2BEB827B9" metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5 m</quantity>
|
||
depth
|
||
</elevation>
|
||
;
|
||
<collectingDate id="C8AB6F90975BB9D4678D8233F2F4BD0B" value="2021-04-10">10 Apr 2021</collectingDate>
|
||
;
|
||
<collectorName id="1F1A1FAB1DDFE628694A9C018D8C4429">C. Katnoum</collectorName>
|
||
leg.; THNHM-lv-19376.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="1CB93DEF0A9B1EB8443C6A88B0FA7F63" pageId="0" pageNumber="557">
|
||
<materialsCitation id="C4E9434D5C22FF4F30769BA01F335029" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="C. Katnoum" country="THAILAND" elevation="5" latitude="13.494722" location="THAILAND" longLatPrecision="1" longitude="99.92369" municipality="Mae Klong River" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Samut Songkhram" typeStatus="Paratype">
|
||
<emphasis id="BFCE66C2D7B5DE9A47458E376E390E0C" bold="true" pageId="0" pageNumber="557">
|
||
<emphasis id="A77B0DBBF6CF840084815554523BDD25" italics="true" pageId="0" pageNumber="557">
|
||
<typeStatus id="48AA544A33AC14DC514FB535A840B1B7">Paratype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="1D695DFB02B523FCF4D2BF96F2050D3C" name="Thailand">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="22C32AF62186F7E1E780ADFE6B3710C7" type="male">5 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="818AF8AB2DD48EB9A272ECEBFE3F5518" type="female">5 ♀</specimenCount>
|
||
; same data as for holotype • THNHM-lv-19379
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1C4D0F0968BFD5A5D7552BAE4AD2DA9" pageId="0" pageNumber="557" type="type locality">
|
||
<paragraph id="AF35ECFD40C86F4882DFB8D4E74E1892" pageId="0" pageNumber="557">Type locality.</paragraph>
|
||
<paragraph id="8EBE0637E61DD6D5A59677CB3C8C97F7" pageId="0" pageNumber="557">
|
||
Darunanukroh School (
|
||
<geoCoordinate id="A28CB0896A576D8C04250FE21ACB3FEE" degrees="13" direction="north" minutes="29" orientation="latitude" precision="1" seconds="41.0" value="13.494722">13°29'41.0"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="34C5498F4C1E6F21B7A1684F826413AB" degrees="99" direction="east" minutes="55" orientation="longitude" precision="1" seconds="25.3" value="99.92369">99°55'25.3"E</geoCoordinate>
|
||
) Mae Klong River, Amphawa District, Samut Songkhram Province, Thailand.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F1605005361BE4508AD5883FD52528FF" pageId="0" pageNumber="557" type="habitat">
|
||
<paragraph id="FA416DC74910367F6627D9E31BAF615A" pageId="0" pageNumber="557">Habitat.</paragraph>
|
||
<paragraph id="849F638EC034BD5037CD053075670F08" pageId="0" pageNumber="557">Freshwater and brackish water (0.05-2.03 ppt) in Mae Klong River of Amphawa District, Samut Songkhram Province, Thailand.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C2883390EB6BF3669BC0878FFEB03D3" pageId="0" pageNumber="557" type="etymology">
|
||
<paragraph id="4BF2F625B0036D60FAED2499712F28A6" pageId="0" pageNumber="557">Etymology.</paragraph>
|
||
<paragraph id="3EC7B1E930DE8E416BCD5C4A10AA998E" pageId="0" pageNumber="557">
|
||
The species is named after their habitat which is located in the river (
|
||
<taxonomicName id="39148072891971753A19DE7E37249346" authorityName="Poey in Poey" authorityYear="1860" class="Actinopteri" family="Rivulidae" genus="Rivulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rivulus" order="Cyprinodontiformes" pageId="0" pageNumber="557" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC15A8752D3FDE1CF61B426F0B1441D" italics="true" pageId="0" pageNumber="557">Rivulus</emphasis>
|
||
</taxonomicName>
|
||
: Latin for river/stream).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E5CA7407ECD6D6D7842623782AC2B7E7" pageId="0" pageNumber="557" type="description">
|
||
<paragraph id="97984D3B2D2D2A99B5B6873FA4F035B8" pageId="0" pageNumber="557">Description.</paragraph>
|
||
<paragraph id="03E9E206E9485984DECD2411AC3E4623" pageId="0" pageNumber="557">Based on holotype, male, size 10 mm, body elongated, cylindrical THNHM-lv-19376.</paragraph>
|
||
<paragraph id="F4CD3D34986BC473E10F34E2C49267E7" pageId="0" pageNumber="557">
|
||
<emphasis id="AAF583F782B1982B13D60C7D1671E4DD" bold="true" pageId="0" pageNumber="557">Head.</emphasis>
|
||
(Fig.
|
||
<figureCitation id="E7920B8A44EA0A407218B9E94B8B4C1A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Habitus of the male holotype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure3" httpUri="https://binary.pensoft.net/fig/947232" pageId="0" pageNumber="557">3A</figureCitation>
|
||
)
|
||
<emphasis id="2BF25305980DE220C695D5EBF5BEBF65" bold="true" italics="true" pageId="0" pageNumber="557">Eye</emphasis>
|
||
medium, 0.1 mm (0.14
|
||
<normalizedToken id="3CFCE6FF4A5E1A509B1E03BDD52E8681" originalValue="×">x</normalizedToken>
|
||
head length); rostrum short (0.16
|
||
<normalizedToken id="34A98B07287CC30D6A947D8D7EF4056A" originalValue="×">x</normalizedToken>
|
||
length of head), acute; Head lateral cephalic lobe well-developed and distally round.
|
||
<emphasis id="78336FEACC2387FDB683BFD397276A86" bold="true" italics="true" pageId="0" pageNumber="557">Antenna 1</emphasis>
|
||
(Fig.
|
||
<figureCitation id="06DD1C4E14FEDCDD28EAFDFBC0026F6F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Habitus of the male holotype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure3" httpUri="https://binary.pensoft.net/fig/947232" pageId="0" pageNumber="557">3B</figureCitation>
|
||
) anterior margin with short setae, posterior margin with long setae, about 0.4
|
||
<normalizedToken id="A8ACBA9EE99AF3FAD97AAA350A1DA5E1" originalValue="×">x</normalizedToken>
|
||
the body length, peduncle 3.1
|
||
<normalizedToken id="0B04F5A0CCBF8E721A148AA344AE1E51" originalValue="×">x</normalizedToken>
|
||
the length of flagellum peduncle article 1 with proximoventral swelling, peduncle article 3 subequal to peduncle article 2, both are slender, several aesthetascs, flagellum with 5 articles, article 1 being the longest, flagellum article 5 the shortest with 1 aesthetasc.
|
||
<emphasis id="632E4A538C79EB4016FFDA2209C5D8CA" bold="true" italics="true" pageId="0" pageNumber="557">Antenna 2</emphasis>
|
||
(Fig.
|
||
<figureCitation id="995E26C82B5D3EE20A439B2B7C4A1496" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Habitus of the male holotype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure3" httpUri="https://binary.pensoft.net/fig/947232" pageId="0" pageNumber="557">3C</figureCitation>
|
||
) anterior margin with short setae, posterior margin with long setae, slightly shorter than antenna 1, setation similar to antenna 1; peduncle article 5 subequal to article 4.
|
||
</paragraph>
|
||
<caption id="58EB1F44783B2B3739651709F3112BBF" doi="10.3897/zse.99.107974.figure2" httpUri="https://binary.pensoft.net/fig/947231" pageId="0" pageNumber="557" start="Figure 2" startId="F2">
|
||
<paragraph id="45CF893D8EE06FE25A2F78A749C46366" pageId="0" pageNumber="557">
|
||
<emphasis id="B6274E6DC4D2D7C48D964639D157225F" bold="true" pageId="0" pageNumber="557">Figure 2.</emphasis>
|
||
<taxonomicName id="3B78E9515504689D20E7271A499BC98B" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="F53E4106E8F471446D9267460BE5EAD8" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="7A00DB52D8F41155E8DD5107604CB967" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Holotype male lateral (THNHM -19377);
|
||
<emphasis id="9323AEDAA1AF73818BE30E90BC7EAA92" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
A specimen in its tube with protruding head and antennae;
|
||
<emphasis id="838FEA81F9D47F8712454A97095DF296" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Paratype female front (THNHM-19379).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5C3090973A603591150BC3BF1C1BDDC7" doi="10.3897/zse.99.107974.figure3" httpUri="https://binary.pensoft.net/fig/947232" pageId="0" pageNumber="557" start="Figure 3" startId="F3">
|
||
<paragraph id="B2565F8FC89C10D1980215E491913582" pageId="0" pageNumber="557">
|
||
<emphasis id="F8C82AA23D78B8C0C976E49A1D455803" bold="true" pageId="0" pageNumber="557">Figure 3.</emphasis>
|
||
<taxonomicName id="D6D0FA9AE89ABE12780EEEFA5B969191" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="D3C421FF3238F1F44B8A6E609028B604" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, male (THNHM -19377).
|
||
<emphasis id="55AE5E5232CEE91ACB1D0934069BB938" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Habitus of the male holotype;
|
||
<emphasis id="6FBB2B95296031E328F53F46A81BD229" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Antenna 1;
|
||
<emphasis id="DE57926FFE015CC881708AD1F6C425E6" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Antenna 2;
|
||
<emphasis id="28B391670FAA6D2B2562E92C3551535C" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Gnathopod 1;
|
||
<emphasis id="3A4CB71FF1F59834AFE29EE29ACB17FA" bold="true" pageId="0" pageNumber="557">E.</emphasis>
|
||
Palm of gnathopod 1;
|
||
<emphasis id="C2A2B9D0B532354E66D9F54BF396CC6F" bold="true" pageId="0" pageNumber="557">F.</emphasis>
|
||
Gnathopod 2;
|
||
<emphasis id="311F63C6C11C9E7A390C096A2FF60EDC" bold="true" pageId="0" pageNumber="557">G.</emphasis>
|
||
Palm of gnathopod 2. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="64EFA54E3B529FFCDCACD92D09407788" pageId="0" pageNumber="557">
|
||
<emphasis id="90EB7FB53503CC223D8A93490D3CFE8D" bold="true" italics="true" pageId="0" pageNumber="557">Upper lip</emphasis>
|
||
(Fig.
|
||
<figureCitation id="7D9DF57388B113B70D396A8B26C8B541" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Left mandible; B. Right mandible; C. Maxilliped; D. Maxilla 2; E. Maxilla 1; F. Upper lip; G. Lower lip. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure6" httpUri="https://binary.pensoft.net/fig/947235" pageId="0" pageNumber="557">6F</figureCitation>
|
||
) symmetrical, notched, with small setulae apically.
|
||
<emphasis id="7A37839AD65B90E74DD44E495267E456" bold="true" italics="true" pageId="0" pageNumber="557">Lower lip</emphasis>
|
||
(Fig.
|
||
<figureCitation id="EB1303C8261CC4ADB8A69803D8673591" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Left mandible; B. Right mandible; C. Maxilliped; D. Maxilla 2; E. Maxilla 1; F. Upper lip; G. Lower lip. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure6" httpUri="https://binary.pensoft.net/fig/947235" pageId="0" pageNumber="557">6G</figureCitation>
|
||
) with inner lobe, smooth, setulose on inner and outer lobes.
|
||
<emphasis id="E60304CCEA6E7A98531E7C60B7811864" bold="true" italics="true" pageId="0" pageNumber="557">Mandibles</emphasis>
|
||
(Fig.
|
||
<figureCitation id="63E8F642C9E54EFE97E9924FAEADF13E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Left mandible; B. Right mandible; C. Maxilliped; D. Maxilla 2; E. Maxilla 1; F. Upper lip; G. Lower lip. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure6" httpUri="https://binary.pensoft.net/fig/947235" pageId="0" pageNumber="557">6A, B</figureCitation>
|
||
) left incisor with 4 teeth; right lacinia mobilis 4-cuspidate (4 teeth); right mandible with 3 broad accessory spines; molar triturative; molar flake absent; palp elongated, robust; article 2 1.3
|
||
<normalizedToken id="826EA5D2743D9BF281206E4736465054" originalValue="×">x</normalizedToken>
|
||
as long as article 3, with 17 marginal setae; article 3 with 10 long setae.
|
||
<emphasis id="F8F6EE29B37636ED67708E8E32D85D34" bold="true" italics="true" pageId="0" pageNumber="557">Maxilla 1</emphasis>
|
||
(Fig.
|
||
<figureCitation id="D3A94148C2026162C890CD73DBC51F29" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Left mandible; B. Right mandible; C. Maxilliped; D. Maxilla 2; E. Maxilla 1; F. Upper lip; G. Lower lip. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure6" httpUri="https://binary.pensoft.net/fig/947235" pageId="0" pageNumber="557">6E</figureCitation>
|
||
) inner plate small without seta; outer plate with 7 apical robust setae; palp biarticulate, with 6 serrated apical robust setae and 4 subapical setae.
|
||
<emphasis id="AA543860ECFA8EA2140041DA3FB761BA" bold="true" italics="true" pageId="0" pageNumber="557">Maxilla 2</emphasis>
|
||
(Fig.
|
||
<figureCitation id="5B0B53D4E0E675E4A115474A5A94568C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Left mandible; B. Right mandible; C. Maxilliped; D. Maxilla 2; E. Maxilla 1; F. Upper lip; G. Lower lip. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure6" httpUri="https://binary.pensoft.net/fig/947235" pageId="0" pageNumber="557">6D</figureCitation>
|
||
) outer plate broader than inner plate, with 12 simple setae; inner plate with 12 setae.
|
||
</paragraph>
|
||
<paragraph id="7007D5EDCFC8159B84F5AD4D81FF5DFE" pageId="0" pageNumber="557">
|
||
<emphasis id="0485396B36BF697A5B9EDA0DB39A32F7" bold="true" pageId="0" pageNumber="557">
|
||
Pereon.
|
||
<emphasis id="F0690DCF8F6FCE0A0A52FEAD91F4E61A" italics="true" pageId="0" pageNumber="557">Gnathopod 1</emphasis>
|
||
</emphasis>
|
||
(Fig.
|
||
<figureCitation id="29471771B82F8B2942ADB650E20F31E6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Habitus of the male holotype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure3" httpUri="https://binary.pensoft.net/fig/947232" pageId="0" pageNumber="557">3D, E</figureCitation>
|
||
) length ratio from basis to dactylus 2.6:1:1:1.7:1.8; coxa 2
|
||
<normalizedToken id="A2AF9A1B2177C8AFFDD767016FBFD3EF" originalValue="×">x</normalizedToken>
|
||
as broad as deep, carpus subequal length to propodus, lobate posterior margin with setae and plumose setae, anterior margin less setae; propodus oval, length 1.7
|
||
<normalizedToken id="9CF9E8C02D909820989BB70114A1C1F6" originalValue="×">x</normalizedToken>
|
||
the width, posterior margin with setae and plumose setae, palm oblique, serrated and with setae; dactylus well developed, palmar margin serrated, crenulated posterior margin withone robust seta.
|
||
<emphasis id="04FA49D0C50455B01E393971FF7E3EC7" bold="true" italics="true" pageId="0" pageNumber="557">Gnathopod 2</emphasis>
|
||
(Fig.
|
||
<figureCitation id="E816AA32E716D3EE0F4A9C11AF174987" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Habitus of the male holotype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure3" httpUri="https://binary.pensoft.net/fig/947232" pageId="0" pageNumber="557">3F, G</figureCitation>
|
||
) carpochelate, length ratio from basis to dactylus 3:1:2:3.5:3:2.5; coxa broader than deep, subequal to coxa 1; basis longer than wide, with 2 marginal setae on anterior margin; carpus massive, length 1.6
|
||
<normalizedToken id="CC7D8058FE635C3CCB5F10E631E65803" originalValue="×">x</normalizedToken>
|
||
the width, palm margin transverse with long posterior defining tooth and well-developed anterodistal recurved spine adjacent to propodus articulation; propodus length 2
|
||
<normalizedToken id="33733A6381669563FB5981B523D74184" originalValue="×">x</normalizedToken>
|
||
the width, with two acute spines and 1 rounded projection on posterior margin, with long setae on anterior and posterior margins; dactylus length 3
|
||
<normalizedToken id="F53AF5A7085A662F17A1835D82FB6ACA" originalValue="×">x</normalizedToken>
|
||
the width, with long setae on posterior margin and one seta on anterior margin.
|
||
</paragraph>
|
||
<paragraph id="77435E6839FFF7BD4FF1C8FF6935F9DF" pageId="0" pageNumber="557">
|
||
<emphasis id="A4BD6DE08074C938D70BC48E2AE23129" bold="true" italics="true" pageId="0" pageNumber="557">Pereopod 3</emphasis>
|
||
(Fig.
|
||
<figureCitation id="84CBA507862E61F16AF5EF767DB64406" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pereopod 3; B. Pereopod 4; C. Pereopod 5; D. Pereopod 6; E. Pereopod 7. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure4" httpUri="https://binary.pensoft.net/fig/947233" pageId="0" pageNumber="557">4A</figureCitation>
|
||
) length ratio from basis to dactylus 14:6:4.5:4:5.5:3.5; coxa 2.2
|
||
<normalizedToken id="1A5519C53DEE33A764AF05E04E5D4428" originalValue="×">x</normalizedToken>
|
||
as broad as deep, with 2 anterodistal setae; basis length 2.2
|
||
<normalizedToken id="F359E8A6CAB05F83C8EF3465CFB7448A" originalValue="×">x</normalizedToken>
|
||
the width, anterior margin with 10 setae; merus with 3 anterodistal setae and 3 posterodistal plumose setae; carpus with 2 anterodistal setae and 2 posterodistal setae; propodus with 3 anterodistal setae.
|
||
</paragraph>
|
||
<caption id="B9EF01276FFB65CAB08C379212BE201B" doi="10.3897/zse.99.107974.figure4" httpUri="https://binary.pensoft.net/fig/947233" pageId="0" pageNumber="557" start="Figure 4" startId="F4">
|
||
<paragraph id="8654E8CEB502AA3D61EAFC56FB6B3D1B" pageId="0" pageNumber="557">
|
||
<emphasis id="94F6E511B1CEBEA8E8F75278A2B9395F" bold="true" pageId="0" pageNumber="557">Figure 4.</emphasis>
|
||
<taxonomicName id="F0E6C31D687C55A036A72ED27315DD9E" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="7A6D0796740210C3DD736B540AC34DEA" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, male (THNHM -19377).
|
||
<emphasis id="6C3F942401FBB2891104522E324426C0" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Pereopod 3;
|
||
<emphasis id="B650FA9F916C94A9682BF177F6D7B1E2" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Pereopod 4;
|
||
<emphasis id="AC6795120DF291CDD0A238ED3204FD47" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Pereopod 5;
|
||
<emphasis id="B597D47B20271E0849D503BBC289503F" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Pereopod 6;
|
||
<emphasis id="096F1C1E377B4E20ADBE055CA75B0A2A" bold="true" pageId="0" pageNumber="557">E.</emphasis>
|
||
Pereopod 7. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="97A2CED4B0AC5268846025D83FD8BAF7" pageId="0" pageNumber="557">
|
||
<emphasis id="69739B58CA609D034A8B3EE6B9F3133D" bold="true" italics="true" pageId="0" pageNumber="557">Pereopod 4</emphasis>
|
||
(Fig.
|
||
<figureCitation id="DAB0333ED6C33F4F2E3980D4254859B8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pereopod 3; B. Pereopod 4; C. Pereopod 5; D. Pereopod 6; E. Pereopod 7. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure4" httpUri="https://binary.pensoft.net/fig/947233" pageId="0" pageNumber="557">4B</figureCitation>
|
||
) length ratio from basis to dactylus 12.5:6.5:6:4:5:3; coxa 1.8
|
||
<normalizedToken id="40222DB62CF1479482DE5AC104294C0D" originalValue="×">x</normalizedToken>
|
||
as broad as deep, with a anterodistal seta; basis, length 1.9
|
||
<normalizedToken id="D8E12E0B38835A690D5A973C2AA1407B" originalValue="×">x</normalizedToken>
|
||
the width, anterior margin with 8 setae; ischium subequal to merus, with 4 posterodistal setae; merus without ridges, with a posterodistal plumose seta; carpus with an anterodistal seta and 2 posterodistal setae; propodus with 4 posteromarginal setae; dactylus with unguis.
|
||
</paragraph>
|
||
<paragraph id="457ADFE18A18F7AE060B692129036F9A" pageId="0" pageNumber="557">
|
||
<emphasis id="169B226B00A6A93E1531B39F216868EA" bold="true" italics="true" pageId="0" pageNumber="557">Pereopod 5</emphasis>
|
||
(Fig.
|
||
<figureCitation id="40C3BFBB7AD4FC05F54583F949663BC4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pereopod 3; B. Pereopod 4; C. Pereopod 5; D. Pereopod 6; E. Pereopod 7. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure4" httpUri="https://binary.pensoft.net/fig/947233" pageId="0" pageNumber="557">4C</figureCitation>
|
||
) length ratio from basis to dactylus 8:3:4.5:3:5.5: 3; coxa 1.2
|
||
<normalizedToken id="F6D237EFAD6A1ECB1B21F3A27713DECC" originalValue="×">x</normalizedToken>
|
||
as broad as deep; basis globular, length equals to width; ischium subrectangular; merus posterior lobe with 1 plumose seta, smaller anterior lobe with 1 seta; carpus smaller than merus, posterior lobe with fine hair, bearing a seta; propodus subovate; dactylus unguis with 2 accessory hooks.
|
||
</paragraph>
|
||
<paragraph id="DED4D9F85CA9DD8CAEB9CC23074C9C4D" pageId="0" pageNumber="557">
|
||
<emphasis id="5A7283E0CE2BDE06BEF1EC89305F1630" bold="true" italics="true" pageId="0" pageNumber="557">Pereopod 6</emphasis>
|
||
(Fig.
|
||
<figureCitation id="2411E76F4E52FE932499E75D9633AF49" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pereopod 3; B. Pereopod 4; C. Pereopod 5; D. Pereopod 6; E. Pereopod 7. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure4" httpUri="https://binary.pensoft.net/fig/947233" pageId="0" pageNumber="557">4D</figureCitation>
|
||
) length ratio from basis to dactylus 8.5:2:4:3:5:3; coxa lobate, about 1.3
|
||
<normalizedToken id="62A7D36856EA80B3FEA717E6D01F45F6" originalValue="×">x</normalizedToken>
|
||
as broad as deep; basis length 1.6
|
||
<normalizedToken id="BF0D60B52E8D93BC87BE07DDEAD730B0" originalValue="×">x</normalizedToken>
|
||
the width, with setae on posterior margin; ischium subrectangular, length 1.25
|
||
<normalizedToken id="DDC1E4E0110840535CBC44EA0EA92D06" originalValue="×">x</normalizedToken>
|
||
width; merus 2.1
|
||
<normalizedToken id="8E24DFDEAD24FE0D19143B8F49B961C9" originalValue="×">x</normalizedToken>
|
||
as long as broad with distal setae on both sides; carpus as long as broad, with 3 long setae on anterodistal corner and 2 long setae on posterodistal corner; propodus with 2 anterodistal long setae and 1 short marginal seta; dactylus unguis with 2 accessory hooks, with 1 seta on anterior side.
|
||
</paragraph>
|
||
<paragraph id="40DD1DBFD39D84CBC1B8585C4AED5875" pageId="0" pageNumber="557">
|
||
<emphasis id="65581A605CC85244F0CB9C0E6CD7895B" bold="true" italics="true" pageId="0" pageNumber="557">Pereopod 7</emphasis>
|
||
(Fig.
|
||
<figureCitation id="D538DA02DB6454E5B4899F941F4F94BF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pereopod 3; B. Pereopod 4; C. Pereopod 5; D. Pereopod 6; E. Pereopod 7. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure4" httpUri="https://binary.pensoft.net/fig/947233" pageId="0" pageNumber="557">4E</figureCitation>
|
||
) length ratio from basis to dactylus 9:3:5:3:5:3; coxa lobate, about 1.6
|
||
<normalizedToken id="9C464ACE7701D2EE07951C41E41E9B45" originalValue="×">x</normalizedToken>
|
||
as broad as deep; basis length 2.25
|
||
<normalizedToken id="C7CF485E94614C9322B81DFE6ED269D2" originalValue="×">x</normalizedToken>
|
||
the width, with setae on anterior margin; ischium subrectangular, length 1.5
|
||
<normalizedToken id="524CA466E8D80D7C21C2B072E148160F" originalValue="×">x</normalizedToken>
|
||
the width; merus 2
|
||
<normalizedToken id="64B3FB38A304D5E8BD9052B7D6FC0FF4" originalValue="×">x</normalizedToken>
|
||
as long as broad with distal long setae on both sides; carpus as long as broad broad, with 1 marginal seta and 3 long setae on anterodistal corner and 3 long setae on posterodistal corner; propodus with 3 marginal setae on both sides and 8 long posterodistal setae; dactylus unguis with 2 accessory hooks.
|
||
</paragraph>
|
||
<paragraph id="A7C7E5BF30951263A003E205E9AF9071" pageId="0" pageNumber="557">
|
||
<emphasis id="1A8FEA5D6F2BDDF5D6921AD9CB507A85" bold="true" pageId="0" pageNumber="557">Pleon.</emphasis>
|
||
Pleopods 1 to 3 decreasing in size.
|
||
<emphasis id="F90476AA6D3FF6876164A7013A9B4323" bold="true" italics="true" pageId="0" pageNumber="557">Pleopod 1</emphasis>
|
||
(Fig.
|
||
<figureCitation id="209868CB2EBA90FB4537A7141223A4B7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">5A</figureCitation>
|
||
) peduncle bearing 2 retinacula, without setae; outer ramus slightly shorter than inner ramus, with 6 articles; outer ramus broader than inner ramus, with 2 articles, first article is the largest.
|
||
<emphasis id="2A48E7B1C63A50ED88201E24C85393E8" bold="true" italics="true" pageId="0" pageNumber="557">Pleopod 2</emphasis>
|
||
(Fig.
|
||
<figureCitation id="8707D769B6C3F4E95DBB386DB1AA86EB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">5B</figureCitation>
|
||
) peduncle with 2 retinacula, without setae; outer ramus 3.1
|
||
<normalizedToken id="F4324441EE2CA50A04609AF224724707" originalValue="×">x</normalizedToken>
|
||
as long as inner ramus, with a single article; inner ramus reduced, 1.6
|
||
<normalizedToken id="FEA186EBB5F0AC8887ED7A61A9658B51" originalValue="×">x</normalizedToken>
|
||
as long as broad, 1 plumose seta and single article.
|
||
<emphasis id="4BFB796C1EDE775D25833B4FA7EE9DE8" bold="true" italics="true" pageId="0" pageNumber="557">Pleopod 3</emphasis>
|
||
(Fig.
|
||
<figureCitation id="58810F6F487543E586AB6146AFB4E3C9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">5C</figureCitation>
|
||
) peduncle with 2 retinacula, without plumose setae distomarginally; outer ramus 3
|
||
<normalizedToken id="EF1FB3E8BBDD55806D9EF0A37C34AFF3" originalValue="×">x</normalizedToken>
|
||
as long as inner ramus; single article; inner ramus reduced, about 1.75
|
||
<normalizedToken id="BBABF854991D838EECB7303B48C4A96B" originalValue="×">x</normalizedToken>
|
||
as long as broad, single article with 1 plumose seta.
|
||
</paragraph>
|
||
<caption id="F2172247BC9BF3F6EC9F4151090175F3" doi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557" start="Figure 5" startId="F5">
|
||
<paragraph id="C51EA40FA3E64B85FBEB1D4001E771AB" pageId="0" pageNumber="557">
|
||
<emphasis id="4E2BD3F28C4F0E6DC8399697B037881C" bold="true" pageId="0" pageNumber="557">Figure 5.</emphasis>
|
||
<taxonomicName id="64105E8BD70DB12FFAF61FDB13714F23" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="7018B258356758E229604733F9A752E8" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, male (THNHM -19377).
|
||
<emphasis id="E6D4CDF4519A586BA074B86344CD6443" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Pleopod 1;
|
||
<emphasis id="991A770251310941CAA49199FEB26B00" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Pleopod 2;
|
||
<emphasis id="C259155D13152775945B687BA6EEAA50" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Pleopod 3;
|
||
<emphasis id="83172CC8060F605BBCBDCC2049732AF8" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Uropods 1-3 and Telson;
|
||
<emphasis id="84D100D7A9271A01555E466401F64D9F" bold="true" pageId="0" pageNumber="557">E.</emphasis>
|
||
Uropod 1. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E28AE338F52F3E9AE684D060F09668D3" doi="10.3897/zse.99.107974.figure6" httpUri="https://binary.pensoft.net/fig/947235" pageId="0" pageNumber="557" start="Figure 6" startId="F6">
|
||
<paragraph id="4A1A45655F1EAF0C9482218E52424087" pageId="0" pageNumber="557">
|
||
<emphasis id="6E3565284A401398555429ACEB564079" bold="true" pageId="0" pageNumber="557">Figure 6.</emphasis>
|
||
<taxonomicName id="4526DE97184F88141A979A515DE93672" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="C4E1058645CDDC4F8946FF9CA5867345" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, male (THNHM -19377).
|
||
<emphasis id="44E23160748B7725387BB2D3653F7F9D" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Left mandible;
|
||
<emphasis id="A917C699F6D6BB48D7134CC926A34FC5" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Right mandible;
|
||
<emphasis id="BD10AD527FC67E928B9490851A801704" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Maxilliped;
|
||
<emphasis id="557A9B4EB1C78C6090CA7A887FBD473E" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Maxilla 2;
|
||
<emphasis id="C70CB08D844712749EE8BFBFFEFC4FC6" bold="true" pageId="0" pageNumber="557">E.</emphasis>
|
||
Maxilla 1;
|
||
<emphasis id="FF21FFB608398D3BDD06A7F2839E51AB" bold="true" pageId="0" pageNumber="557">F.</emphasis>
|
||
Upper lip;
|
||
<emphasis id="CE6D1619F1F78C436DA8B594A86A1758" bold="true" pageId="0" pageNumber="557">G</emphasis>
|
||
. Lower lip. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="44AB0EB380A4A1615031B30470200F8F" pageId="0" pageNumber="557">
|
||
<emphasis id="4447FEE17D6FE241DD7DF48E08763531" bold="true" pageId="0" pageNumber="557">Urosome.</emphasis>
|
||
(Fig.
|
||
<figureCitation id="8560F6AA3757E161FA0B7DABE440DE51" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">5D</figureCitation>
|
||
)
|
||
<emphasis id="17AF61CC9256E89D347B3F518FC46683" bold="true" italics="true" pageId="0" pageNumber="557">Uropod 1</emphasis>
|
||
(Fig.
|
||
<figureCitation id="8F7A1DCD529B67AA40A8F86E48080C3C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">5E</figureCitation>
|
||
) biramous; peduncle 2.1
|
||
<normalizedToken id="D4DA634CA88F978A92C0AD9F03594AEC" originalValue="×">x</normalizedToken>
|
||
as long as wide, with 3 dorsodistal setae; outer ramus shorter than peduncle, 1.5
|
||
<normalizedToken id="50EA61FC3FF570FEBA8FB15CDE34975E" originalValue="×">x</normalizedToken>
|
||
longer than inner ramus, with outer row of spinules and 1 large apical robust seta, inner ramus with a large apical robust seta, corona of short robust setae surrounding large terminal robust seta.
|
||
</paragraph>
|
||
<paragraph id="2A6ECBCE2F48E0469900747D3F2704BD" pageId="0" pageNumber="557">
|
||
<emphasis id="5E4CF2EE239C743299F66470F651A864" bold="true" italics="true" pageId="0" pageNumber="557">Uropod 2</emphasis>
|
||
(Fig.
|
||
<figureCitation id="09DE07440A2DB6D4A7C9C318966DA7CB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">5D</figureCitation>
|
||
) uniramous; peduncle 3.75
|
||
<normalizedToken id="D3D1692F111AD287FEAACD1DC6E8D3E0" originalValue="×">x</normalizedToken>
|
||
as long as broad, 6.3
|
||
<normalizedToken id="96131EE5060E1AA313F902968EB3AAEB" originalValue="×">x</normalizedToken>
|
||
as long as vestigial ramus; ramus with 1 row of short setae and a terminal seta.
|
||
<emphasis id="B9F57940574442CAE5B52668AA2BC5AA" bold="true" italics="true" pageId="0" pageNumber="557">Uropod 3</emphasis>
|
||
(Fig.
|
||
<figureCitation id="F9D1765B0AA573317EAEBFE242AEC5E9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">5D</figureCitation>
|
||
) uniramous; peduncle 2.7
|
||
<normalizedToken id="9473798A6DD9379865ED82A34758B868" originalValue="×">x</normalizedToken>
|
||
as long as wide, with one seta on inner margin; ramus vestigial, inconspicuously small with 2 hooks.
|
||
<emphasis id="E84FDDA660F641EF8D9AA158C98511E0" bold="true" italics="true" pageId="0" pageNumber="557">Telson</emphasis>
|
||
(Fig.
|
||
<figureCitation id="2986E23958D4986018921492B07EB3D4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Cerapus rivulus sp. nov., holotype, male (THNHM - 19377). A. Pleopod 1; B. Pleopod 2; C. Pleopod 3; D. Uropods 1 - 3 and Telson; E. Uropod 1. Scale bars: 0.1 mm." figureDoi="10.3897/zse.99.107974.figure5" httpUri="https://binary.pensoft.net/fig/947234" pageId="0" pageNumber="557">5D</figureCitation>
|
||
) deeply cleft, 0.5
|
||
<normalizedToken id="62D095C1D94D1A850194AA224C41C85A" originalValue="×">x</normalizedToken>
|
||
the length of uropod 3, each lobe with 19 or 20 recurved hooks in two rows and an apical seta.
|
||
</paragraph>
|
||
<paragraph id="702B93AAF99B8C28916A90B79916E2BB" pageId="0" pageNumber="557">
|
||
<emphasis id="89A258AD65D6A92A31AB469D2814213A" bold="true" pageId="0" pageNumber="557">Female</emphasis>
|
||
(Figs
|
||
<figureCitation id="7AC0766517032E2D998B588AD31DA296" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Habitus of the female paratype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure7" httpUri="https://binary.pensoft.net/fig/947236" pageId="0" pageNumber="557">7</figureCitation>
|
||
-
|
||
<figureCitation id="739FF04E9DAEAC961170C4124EDA4B85" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Maxilliped; B. Maxilla 2; C. Maxilla 1; D. Upper lip; E. Lower lip; F. Left mandible; G. Right mandible. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure10" httpUri="https://binary.pensoft.net/fig/947239" pageId="0" pageNumber="557">10</figureCitation>
|
||
) (Sexually dimorphic characters). Based on allotype (THNHM-iv-19379), length 3.5 mm (Fig.
|
||
<figureCitation id="BF66597A43AB82669FDBE9600CDAE36D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Habitus of the female paratype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure7" httpUri="https://binary.pensoft.net/fig/947236" pageId="0" pageNumber="557">7A</figureCitation>
|
||
). Similar to male, except for the following:
|
||
<emphasis id="5DF1EF528E600FAF62D16B7CE688E3EB" bold="true" italics="true" pageId="0" pageNumber="557">Antenna 1</emphasis>
|
||
(Fig.
|
||
<figureCitation id="34AC977E5E8631361D4F35001B62E763" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Habitus of the female paratype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure7" httpUri="https://binary.pensoft.net/fig/947236" pageId="0" pageNumber="557">7B</figureCitation>
|
||
) with fewer setae and fewer aesthetascs.
|
||
<emphasis id="AE4AD658D5F11586FFADC31EB120AD16" bold="true" italics="true" pageId="0" pageNumber="557">Antenna 2</emphasis>
|
||
(Fig.
|
||
<figureCitation id="16D2CFC8ECFE76A7D2710E35A5B46068" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Habitus of the female paratype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure7" httpUri="https://binary.pensoft.net/fig/947236" pageId="0" pageNumber="557">7C</figureCitation>
|
||
) more slender than in male.
|
||
<emphasis id="389F00AD55569F2680AFEC61879C114F" bold="true" italics="true" pageId="0" pageNumber="557">Gnathopod 1</emphasis>
|
||
(Fig.
|
||
<figureCitation id="19442E26ECFA970299AAC26724A922D8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Habitus of the female paratype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure7" httpUri="https://binary.pensoft.net/fig/947236" pageId="0" pageNumber="557">7D, E</figureCitation>
|
||
) dactylus posterior margin serrated, with one spine.
|
||
<emphasis id="B0765A24F6EB23A736F847D9AC80261E" bold="true" italics="true" pageId="0" pageNumber="557">Gnathopod 2</emphasis>
|
||
(Fig.
|
||
<figureCitation id="2C435319BFB92F3249161393BA746D19" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Habitus of the female paratype; B. Antenna 1; C. Antenna 2; D. Gnathopod 1; E. Palm of gnathopod 1; F. Gnathopod 2; G. Palm of gnathopod 2. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure7" httpUri="https://binary.pensoft.net/fig/947236" pageId="0" pageNumber="557">7F, G</figureCitation>
|
||
) slightly larger than gnathopod 1; carpus triangular, posterodistal corner with plumose setae and normal setae; propodus palm not serrated, with plumose setae and simple setae; dactylus posterior margin crenulated.
|
||
<emphasis id="46C94FCC6758CFD5F55BC84692493546" bold="true" italics="true" pageId="0" pageNumber="557">Pereopod 5</emphasis>
|
||
(Fig.
|
||
<figureCitation id="D993E2973B382B8906AA4A871A1D7F77" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cerapus rivulus sp. nov., paratype, female (THNHM - 19379). A. Pereopod 3; B. Pereopod 4; C. Pereopod 5; D. Pereopod 6; E. Pereopod 7. Scale bars: 0.2 mm." figureDoi="10.3897/zse.99.107974.figure8" httpUri="https://binary.pensoft.net/fig/947237" pageId="0" pageNumber="557">8C</figureCitation>
|
||
) coxa larger, about 2.7
|
||
<normalizedToken id="54C676E299D7B2BCD9584958E5E2EAF4" originalValue="×">x</normalizedToken>
|
||
as broad as deep, about 6.3
|
||
<normalizedToken id="AED881A36E68A7F22F4351E873AA422F" originalValue="×">x</normalizedToken>
|
||
longer than the basis width; basis slightly globular; dactylus with 2 accessory hooks; with a pair of oostegites on each segment from 4 to 6.
|
||
</paragraph>
|
||
<caption id="F9EFFED4064084EE86020D77743ACFA3" doi="10.3897/zse.99.107974.figure7" httpUri="https://binary.pensoft.net/fig/947236" pageId="0" pageNumber="557" start="Figure 7" startId="F7">
|
||
<paragraph id="DE2FF1B262C3CC79099031600E04891B" pageId="0" pageNumber="557">
|
||
<emphasis id="C0662269AA8C3654AE65B52172FE833B" bold="true" pageId="0" pageNumber="557">Figure 7.</emphasis>
|
||
<taxonomicName id="9C97871B5AD8708ABE927204FD1DC918" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="F8871E56CA0A952D03D3046BA616C165" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, female (THNHM -19379).
|
||
<emphasis id="F6EAF7F3D19DFA790C7D8A1110A953FB" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Habitus of the female paratype;
|
||
<emphasis id="38F7AA7767B227E1C9817E0E0D299CBC" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Antenna 1;
|
||
<emphasis id="F0963CA65F493CA5B47D3119B88DCF09" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Antenna 2;
|
||
<emphasis id="0C5EB03D809CAD78E45DF512DDE1E673" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Gnathopod 1;
|
||
<emphasis id="54AB0B0A4181A192B36286CEA89000A6" bold="true" pageId="0" pageNumber="557">E.</emphasis>
|
||
Palm of gnathopod 1;
|
||
<emphasis id="0D221D9E34E8175181AB02666BBE06DE" bold="true" pageId="0" pageNumber="557">F.</emphasis>
|
||
Gnathopod 2;
|
||
<emphasis id="87A7D1875A952C2D02413832627302F6" bold="true" pageId="0" pageNumber="557">G.</emphasis>
|
||
Palm of gnathopod 2. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFC68D349399584DECB7968808BA850B" doi="10.3897/zse.99.107974.figure8" httpUri="https://binary.pensoft.net/fig/947237" pageId="0" pageNumber="557" start="Figure 8" startId="F8">
|
||
<paragraph id="AC0214FBDFC7A8EAB918ACE4214A16AE" pageId="0" pageNumber="557">
|
||
<emphasis id="5A5773C90C547BC33281D78E25239EDB" bold="true" pageId="0" pageNumber="557">Figure 8.</emphasis>
|
||
<taxonomicName id="D8357E600502E6E365F8DCB995662290" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="991F66C63D05E6C7E8C150079033ACCA" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, female (THNHM -19379).
|
||
<emphasis id="CF0ADBD035E3535F1B409EDE0C3D529E" bold="true" pageId="0" pageNumber="557">A</emphasis>
|
||
. Pereopod 3;
|
||
<emphasis id="79C8789FF3AFEBD5EE6C659EE367B5B6" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Pereopod 4;
|
||
<emphasis id="0C357698694A3438219ECCB5AB3AF44A" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Pereopod 5;
|
||
<emphasis id="3B67DFDFB5C55111ECBC1736D8EC1794" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Pereopod 6;
|
||
<emphasis id="14F20D75A87FC0A25BBACC8D5D3B9EE4" bold="true" pageId="0" pageNumber="557">E.</emphasis>
|
||
Pereopod 7. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BB309FAFC36C9FC03C84843550D4DE6B" doi="10.3897/zse.99.107974.figure9" httpUri="https://binary.pensoft.net/fig/947238" pageId="0" pageNumber="557" start="Figure 9" startId="F9">
|
||
<paragraph id="850868D798C8C57A81DDAB581ED0C384" pageId="0" pageNumber="557">
|
||
<emphasis id="5C7BA50E563537AE4F626471EDDDFFD0" bold="true" pageId="0" pageNumber="557">Figure 9.</emphasis>
|
||
<taxonomicName id="54494936A4BEDC0BAD63A92C1B7FF201" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="BA2D1ED773D1F0F63056B000A5D4F1FD" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, female (THNHM -19379).
|
||
<emphasis id="25347F51B31BAB7139D789C48D70C8D9" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Pleopod 1;
|
||
<emphasis id="B37EC229FE5CE8794E7A33AAECA879AB" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Pleopod 2;
|
||
<emphasis id="8EA281829DC213FC52196A1A4DDC0A65" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Pleopod 3;
|
||
<emphasis id="A3F8A21D0760EE03385F05699DCEB933" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Uropods 1-3 and Telson. Scale bars: 0.1 mm (
|
||
<emphasis id="97EF2D61778B51D9AB0E4B93B1FA788C" bold="true" pageId="0" pageNumber="557">A-C</emphasis>
|
||
); 0.2 mm (
|
||
<emphasis id="1CC920824EAA8FAAC6ACE77A0CD9FA58" bold="true" pageId="0" pageNumber="557">D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C806BE6A31619D936CFCAF65602C2279" doi="10.3897/zse.99.107974.figure10" httpUri="https://binary.pensoft.net/fig/947239" pageId="0" pageNumber="557" start="Figure 10" startId="F10">
|
||
<paragraph id="DA40AD378AF80CF9E68647125DCFA08F" pageId="0" pageNumber="557">
|
||
<emphasis id="850CB693FADAFB9B614C6CD8229FBEEB" bold="true" pageId="0" pageNumber="557">Figure 10.</emphasis>
|
||
<taxonomicName id="2DB0BFA0DD89176E59BDB7D580401685" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="A608A8A325CC22AFF04C4522AEFDC50A" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, female (THNHM -19379).
|
||
<emphasis id="1F72C6A838EA0404542279D0B85D1ACD" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Maxilliped;
|
||
<emphasis id="64EF51265EA48FC829D5B651B31E074E" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Maxilla 2;
|
||
<emphasis id="439DFA4B485FD646DD84C75FFAA7ABD8" bold="true" pageId="0" pageNumber="557">C.</emphasis>
|
||
Maxilla 1;
|
||
<emphasis id="83E186506E9587F560521E3F7AD0495E" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Upper lip;
|
||
<emphasis id="E31D9B92390B31C2201E5527E3A4E08F" bold="true" pageId="0" pageNumber="557">E.</emphasis>
|
||
Lower lip;
|
||
<emphasis id="246A8BE6F7FB3969C4E3A664C7E24C10" bold="true" pageId="0" pageNumber="557">F.</emphasis>
|
||
Left mandible;
|
||
<emphasis id="DD7AAC6B5164CD6D713E9355D918C602" bold="true" pageId="0" pageNumber="557">G.</emphasis>
|
||
Right mandible. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="5CAA087202EB5E726240689E85DDD046" pageId="0" pageNumber="557" type="remarks">
|
||
<paragraph id="D599676FC2FA74E15F0CEC098CDD47FB" pageId="0" pageNumber="557">Remarks.</paragraph>
|
||
<paragraph id="99D4F11FFDE55B2444ECC7E3EDCE262D" pageId="0" pageNumber="557">
|
||
<taxonomicName id="DF9B07B7523EF902B3650B02588B67B6" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="83703646265321E220CF36E8FF7E10B4" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName id="4602B2A8E38ABEFBF742D477A4B9714A" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. nudus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="nudus">
|
||
<emphasis id="A8FF0800C9175D61DBE9D181E72B943D" italics="true" pageId="0" pageNumber="557">C. nudus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DEE04754832558324385A5F92B1693FA" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longirostris" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longirostris">
|
||
<emphasis id="8BEF09125474E473BEB5BB9D743A23A8" italics="true" pageId="0" pageNumber="557">C. longirostris</emphasis>
|
||
</taxonomicName>
|
||
which has synapomorphic character states, such as: (1) pereonites 1-2 without constriction; (2) pereopod 5 merus without plumose seta on posterior lobe; (3) pereopod 6 coxa without setae on ventral margin; (4) uropod 1 without hook on ventrodistal margin. They are distinguished by the following: (1) head subequal in length to pereonites 1 + 2 [
|
||
<taxonomicName id="5B363E3FC4636AFB9F78220074E31FB2" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. nudus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="nudus">
|
||
<emphasis id="D05937F8B36617271D06371A81118F18" italics="true" pageId="0" pageNumber="557">C. nudus</emphasis>
|
||
</taxonomicName>
|
||
longer than pereonites 1 + 2,
|
||
<taxonomicName id="AB59324BFC32DF202477E90996B445F0" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longirostris" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longirostris">
|
||
<emphasis id="8DC266D8B18B56CC0E8A0C8152DE0B24" italics="true" pageId="0" pageNumber="557">C. longirostris</emphasis>
|
||
</taxonomicName>
|
||
subequal to pereonites 1 + 2]; (2) gnathopod 2 propodus twice as long as wide [
|
||
<taxonomicName id="ED4505A234C325DE3F60E31F7C35A75D" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. nudus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="nudus">
|
||
<emphasis id="4D25952DE47B70ABFBFF954E771E4D2A" italics="true" pageId="0" pageNumber="557">C. nudus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EBEDC39F9A2706D8930416683CE9B663" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longirostris" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longirostris">
|
||
<emphasis id="780A6673EE34EDC2AD61B33D8A1EDCB3" italics="true" pageId="0" pageNumber="557">C. longirostris</emphasis>
|
||
</taxonomicName>
|
||
less than twice as long as wide]; (3) telson deeply cleft (> 50%) [
|
||
<taxonomicName id="AF8FBFB9DFF3721CFB9D8AABF9EA6ECC" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. nudus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="nudus">
|
||
<emphasis id="B89635CD9873C77F591C3492667D19C3" italics="true" pageId="0" pageNumber="557">C. nudus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9F04716C1FE1D711333AB9DCC4B3E25A" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longirostris" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longirostris">
|
||
<emphasis id="CA9F46B4491E0174344525643DAA783A" italics="true" pageId="0" pageNumber="557">C. longirostris</emphasis>
|
||
</taxonomicName>
|
||
semi-cleft]; (4) telsonic lobe with 19-20 hooks in two transverse rows [
|
||
<taxonomicName id="B2158D25B173807B97370DA60EDA1A61" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. nudus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="nudus">
|
||
<emphasis id="3DBB929A4F9DA81803C18DE4196D702A" italics="true" pageId="0" pageNumber="557">C. nudus</emphasis>
|
||
</taxonomicName>
|
||
with ten recurved hooks on three transverse rows and
|
||
<taxonomicName id="34C8DA2A7A96756F8EAAE9F822D3CE62" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longirostris" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longirostris">
|
||
<emphasis id="A853B39D8B47B4802E801DAD4664A335" italics="true" pageId="0" pageNumber="557">C. longirostris</emphasis>
|
||
</taxonomicName>
|
||
with eight recurved hooks on two transverse rows.
|
||
</paragraph>
|
||
<paragraph id="CBB76C29EABB5515D0AC44082786B6E3" pageId="0" pageNumber="557">
|
||
This study brings the number of identified
|
||
<taxonomicName id="4EE74468DE3D0545F0E910BF387D6AA3" authorityName="Say" authorityYear="1817" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9BB8458439415F4F11DAC48625B96FA1" italics="true" pageId="0" pageNumber="557">Cerapus</emphasis>
|
||
</taxonomicName>
|
||
species from Southeast Asia to seven.
|
||
<taxonomicName id="E961B1EC4619F4C4D2ABC95F220609C6" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="292E4BA544441B504FDA353AF2A563FD" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be separated from Southeast Asian congeners by a combination of characteristics as follows (other species in paratheses): absence of constriction between pereonite 1 and 2 (present in
|
||
<taxonomicName id="247704B0501269B886158C6E9B5C7870" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. yuyatalay" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="yuyatalay">
|
||
<emphasis id="8808F020073F8E75F03B5CCBD5F33AE8" italics="true" pageId="0" pageNumber="557">C. yuyatalay</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5F37EFA091146450CE9548052B4571D4" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. bumbumiensis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="bumbumiensis">
|
||
<emphasis id="ECB264F64678A34EEAD2BCD1067F9CFF" italics="true" pageId="0" pageNumber="557">C. bumbumiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="CFB5F7FBFB98F6E4479D286EA6F5E654" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longicervicum" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longicervicum">
|
||
<emphasis id="80C51E7C9B9098E084A4BA3D37BA6AEE" italics="true" pageId="0" pageNumber="557">C. longicervicum</emphasis>
|
||
</taxonomicName>
|
||
); antenna 1 peduncular article 1 shorter than article 3 (longer in
|
||
<taxonomicName id="79343A51E14E1EFE3208246C4E0C351B" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. chaomai" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="chaomai">
|
||
<emphasis id="EC1912AB635BBA2BA6E9D621FBF7E807" italics="true" pageId="0" pageNumber="557">C. chaomai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E6736B7592BD494E418BE2CA8256B5B6" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. bumbumiensis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="bumbumiensis">
|
||
<emphasis id="9ED96ECAD34684E1783ED3FB8DF252A2" italics="true" pageId="0" pageNumber="557">C. bumbumiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7502966277E65AA80B262CE4889116AC" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longicervicum" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longicervicum">
|
||
<emphasis id="8C946A1567B5B51F76D8CD00834AB175" italics="true" pageId="0" pageNumber="557">C. longicervicum</emphasis>
|
||
</taxonomicName>
|
||
); gnathopod 2 propodus with tooth in male (without tooth in
|
||
<taxonomicName id="B192172821E5BDFD949A6E32D22D111C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. chaomai" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="chaomai">
|
||
<emphasis id="D84DB56B2B6BB62993B7B065E80B55EC" italics="true" pageId="0" pageNumber="557">C. chaomai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="454A52D4D596F75584D7C0CE500CF157" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. yuyatalay" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="yuyatalay">
|
||
<emphasis id="F18E07F757E3F824CED6B0AA432A9386" italics="true" pageId="0" pageNumber="557">C. yuyatalay</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F414AC5B0F323BC8756D07054E477473" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. bumbumiensis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="bumbumiensis">
|
||
<emphasis id="A27D48D6CEBBAACFD45E1DE4040B2C2C" italics="true" pageId="0" pageNumber="557">C. bumbumiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2BF636837EF48E79E50491DA58EF7AEE" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longicervicum" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longicervicum">
|
||
<emphasis id="8EE9D5C7EC115D3C1AD72D05D901F18E" italics="true" pageId="0" pageNumber="557">C. longicervicum</emphasis>
|
||
</taxonomicName>
|
||
); telson fused with pleonite 3 (not fused in
|
||
<taxonomicName id="A55CA2CEFEBCE2914AC8D494A9CF1044" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. yuyatalay" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="yuyatalay">
|
||
<emphasis id="A46166D231E7028E025B2C276ECF1B7C" italics="true" pageId="0" pageNumber="557">C. yuyatalay</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9470ED6400AEC236CADEA5479F2E4786" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. bumbumiensis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="bumbumiensis">
|
||
<emphasis id="FB400EF04C81DBAD74C9A3DA192DD33D" italics="true" pageId="0" pageNumber="557">C. bumbumiensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EB6B45FDF6D6FA890BD48A5DBB906CFC" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. longicervicum" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="longicervicum">
|
||
<emphasis id="A8C7D9BAEFCEC60786F4E567E44973EE" italics="true" pageId="0" pageNumber="557">C. longicervicum</emphasis>
|
||
</taxonomicName>
|
||
); telson with more than ten hooks (fewer than ten in
|
||
<taxonomicName id="2DE3B5CB33CCCC9BAD189522AE790CB5" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. bumbumiensis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="bumbumiensis">
|
||
<emphasis id="0226860E0DC46F439AD8861E8F45FFD8" italics="true" pageId="0" pageNumber="557">C. bumbumiensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D096DDBCE5C4EE0082CA47A1EFC04ACE" pageId="0" pageNumber="557">
|
||
<taxonomicName id="F14D4B72EA8FC822C8AFDE60FA316CFA" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. tubularis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="tubularis">
|
||
<emphasis id="55230261C450236CDE61494DE6314E28" italics="true" pageId="0" pageNumber="557">C. tubularis</emphasis>
|
||
</taxonomicName>
|
||
was described from Long Island Sound, New York, U.S.A. and after that reported in Japan by
|
||
<bibRefCitation id="457A4DA3C1F34EE29829118A2A8024CF" DOI="https://doi.org/10.5134/175915" author="Morino, H" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="557" refId="B20" refString="Morino, H, 1976. On two new forms of Cerapus tubularis, a tube dwelling Amphipoda, from shallow waters of Japan. Publications of the Seto Marine Biological Laboratory 23(1/2): 179-189. https://doi.org/10.5134/175915" title="On two new forms of Cerapus tubularis, a tube dwelling Amphipoda, from shallow waters of Japan. Publications of the Seto Marine Biological Laboratory 23 (1 / 2): 179 - 189." url="https://doi.org/10.5134/175915" year="1976">Morino (1976)</bibRefCitation>
|
||
. However,
|
||
<bibRefCitation id="7B54A81C9A792648D58EEDA9901A4314" DOI="https://doi.org/10.1080/00222938900770711" author="Lowry, JK" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="557" pagination="1341 - 135" refId="B14" refString="Lowry, JK, Berents, PB, 1989. A redescription of Cerapus tubularis Say, 1818, based on material of the first reviewer, S.I. Smith, 1880, (Crustacea: Amphipoda: Corophioidea). Journal of Natural History 23: 1341 - 135, DOI: https://doi.org/10.1080/00222938900770711" title="A redescription of Cerapus tubularis Say, 1818, based on material of the first reviewer, S. I. Smith, 1880, (Crustacea: Amphipoda: Corophioidea)." url="https://doi.org/10.1080/00222938900770711" volume="23" year="1989">Lowry and Berents (1989)</bibRefCitation>
|
||
re-described
|
||
<taxonomicName id="050E461C8FCF2AA4241C7DDBCC1575E1" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. tubularis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="tubularis">
|
||
<emphasis id="DB8D6DE55E5EE4FE8E4EE2B8E4381745" italics="true" pageId="0" pageNumber="557">C. tubularis</emphasis>
|
||
</taxonomicName>
|
||
and excluded the record of
|
||
<bibRefCitation id="234B41A79EE3FC0132FD43A5366D098B" DOI="https://doi.org/10.5134/175915" author="Morino, H" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="557" refId="B20" refString="Morino, H, 1976. On two new forms of Cerapus tubularis, a tube dwelling Amphipoda, from shallow waters of Japan. Publications of the Seto Marine Biological Laboratory 23(1/2): 179-189. https://doi.org/10.5134/175915" title="On two new forms of Cerapus tubularis, a tube dwelling Amphipoda, from shallow waters of Japan. Publications of the Seto Marine Biological Laboratory 23 (1 / 2): 179 - 189." url="https://doi.org/10.5134/175915" year="1976">Morino (1976)</bibRefCitation>
|
||
. Lastly,
|
||
<bibRefCitation id="4443CC1DCC260723FEAA7401A11B7872" author="White, K" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="557" pagination="139 - 142" refId="B31" refString="White, K, 2015. A checklist of Amphipoda (Crustacea) collected from the mudflats of Pulau Ubin, Singapore. The Raffles Bulletin of Zoology 31: 139 - 142" title="A checklist of Amphipoda (Crustacea) collected from the mudflats of Pulau Ubin, Singapore." volume="31" year="2015">White (2015)</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="3BDAFB49ACEF4FAE6433D780B44A4C05" authorityName="Say" authorityYear="1817" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="609F1FB5E99F60B52C8FA1EA9730522C" italics="true" pageId="0" pageNumber="557">Cerapus</emphasis>
|
||
</taxonomicName>
|
||
sp. in Singapore and noted that the amphipod resembles
|
||
<taxonomicName id="F8FFAD090C7EB82BADE39108FD89E73D" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. tubularis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="tubularis">
|
||
<emphasis id="AA06E26DA4E6EAC53D04596283FCC6A7" italics="true" pageId="0" pageNumber="557">C. tubularis</emphasis>
|
||
</taxonomicName>
|
||
and possibly was introduced to Singapore. The presence of
|
||
<taxonomicName id="173DB0E58268DF6BDD09A63F60F3C267" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. tubularis" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="tubularis">
|
||
<emphasis id="43B923886E46BADF16305088EA53863E" italics="true" pageId="0" pageNumber="557">C. tubularis</emphasis>
|
||
</taxonomicName>
|
||
in the eastern Pacific and South China Sea is still unclear and needs clarification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="902E23D8F334A31719B80B0A52418597" pageId="0" pageNumber="557" type="distribution">
|
||
<paragraph id="E988506507704284EFEAEA1A9528CB27" pageId="0" pageNumber="557">Geographic distribution.</paragraph>
|
||
<paragraph id="EDB14F8AD518D0EEA29BF3EC99956C5A" pageId="0" pageNumber="557">
|
||
The
|
||
<taxonomicName id="4ABC145C73B8712159DDACD284EB5911" authorityName="Say" authorityYear="1817" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8E35556A701DCF1CDC8534EAD594AB9D" italics="true" pageId="0" pageNumber="557">Cerapus</emphasis>
|
||
</taxonomicName>
|
||
are distributed worldwide and mainly inhabit marine and brackish water (Fig.
|
||
<figureCitation id="3DAAA577FE584178EBE975646EE64ED9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of Cerapus spp." figureDoi="10.3897/zse.99.107974.figure11" httpUri="https://binary.pensoft.net/fig/947240" pageId="0" pageNumber="557">11</figureCitation>
|
||
) (
|
||
<bibRefCitation id="5593C0A3FA36A67E39137CFED4998929" DOI="https://doi.org/10.1080/00222939100770931" author="Lowry, JK" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="557" pagination="1461 - 1467" refId="B19" refString="Lowry, JK, Thomas, JD, 1991. A new species of Cerapus from Cudjoe Channel, Lower Florida Keys, USA, with notes on male behaviour (Crustacea: Amphipoda: Corophioidea). Journal of Natural History 25 (6): 1461 - 1467, DOI: https://doi.org/10.1080/00222939100770931" title="A new species of Cerapus from Cudjoe Channel, Lower Florida Keys, USA, with notes on male behaviour (Crustacea: Amphipoda: Corophioidea)." url="https://doi.org/10.1080/00222939100770931" volume="25" year="1991">Lowry and Thomas 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="274E02587C55557696CB1C6E42042D62" author="Lowry, JK" journalOrPublisher="Phuket Marine Biological Center Special Publication" pageId="0" pageNumber="557" pagination="189 - 196" refId="B16" refString="Lowry, JK, Berents, PB, 2002. The genus Cerapus in the Andaman Sea (Crustacea, Amphipoda, Ischyroceridae). Phuket Marine Biological Center Special Publication 23: 189 - 196" title="The genus Cerapus in the Andaman Sea (Crustacea, Amphipoda, Ischyroceridae)." volume="23" year="2002">Lowry and Berents 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9F09D1A460C13941CA126F642BF51209" DOI="https://doi.org/10.11646/zootaxa.4441.3.4" author="Drumm, DT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="557" pagination="495 - 510" refId="B8" refString="Drumm, DT, 2018. Two new species of Cerapus (Crustacea: Amphipoda: Ischyroceridae) from the Northwest Atlantic and Gulf of Mexico. Zootaxa 4441 (3): 495 - 510, DOI: https://doi.org/10.11646/zootaxa.4441.3.4" title="Two new species of Cerapus (Crustacea: Amphipoda: Ischyroceridae) from the Northwest Atlantic and Gulf of Mexico." url="https://doi.org/10.11646/zootaxa.4441.3.4" volume="4441" year="2018">Drumm 2018</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="3F02EE6650C462EC1E94DB58C2C06864" DOI="https://doi.org/10.11646/zootaxa.4802.3.7" author="Nurshazwan, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="557" pagination="519 - 533" refId="B22" refString="Nurshazwan, J, Ahmad-Zaki, AB, Azman, BAR, 2020. A new species of Cerapus (Amphipoda: Senticaudata: Ischyroceridae) from Pulau Bum Bum, Sabah, Malaysia, with an identification key to Cerapus species. Zootaxa 4802 (3): 519 - 533, DOI: https://doi.org/10.11646/zootaxa.4802.3.7" title="A new species of Cerapus (Amphipoda: Senticaudata: Ischyroceridae) from Pulau Bum Bum, Sabah, Malaysia, with an identification key to Cerapus species." url="https://doi.org/10.11646/zootaxa.4802.3.7" volume="4802" year="2020">Nurshazwan et al. 2020</bibRefCitation>
|
||
). Interestingly,
|
||
<taxonomicName id="FF7833F7166A8910EAC94055D448CEF1" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="1B493649D8C07D433D4DB41275F25F51" italics="true" pageId="0" pageNumber="557">C. rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. was found in freshwater and brackish water (salinity less than 3 ppt), so this is the first report for freshwater habitats. They were dominant in the coarse filter substrate used for benthos sampling because the tubes are retained on the coarse filter.
|
||
</paragraph>
|
||
<caption id="F08721FDD1975823776BA9320EFF6631" doi="10.3897/zse.99.107974.figure11" httpUri="https://binary.pensoft.net/fig/947240" pageId="0" pageNumber="557" start="Figure 11" startId="F11">
|
||
<paragraph id="0766546D383D0BB8CD58E547AD754C65" pageId="0" pageNumber="557">
|
||
<emphasis id="DE8488E5656276A3511E57BFBBAFB7D6" bold="true" pageId="0" pageNumber="557">Figure 11.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="DF692EF9ADDC80B82438C6321F8A5FCE" authorityName="Say" authorityYear="1817" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9652163F881C6C8F322DECB4BC7E102" italics="true" pageId="0" pageNumber="557">Cerapus</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="CBE78DE5B12CA3A17FB5F578758FF94B" pageId="0" pageNumber="557" type="notes">
|
||
<paragraph id="80A5725D8B2AE2B03CFB37B177865B5D" pageId="0" pageNumber="557">Biological notes.</paragraph>
|
||
<paragraph id="1E0F202130A1212856BCE3E84A787727" pageId="0" pageNumber="557">
|
||
Generally, tube-building amphipods build tubes using amphipod silk which is observed as silk strands adhering to the tip of the dactylar surface close to the pore on P3 and P4 (
|
||
<bibRefCitation id="30903828B03C5D02FBA5D83E45BF40EA" DOI="https://doi.org/10.1163/193724011X615532" author="Kronenberger, K" journalOrPublisher="Journal of Crustacean Biology" pageId="0" pageNumber="557" pagination="191 - 202" refId="B11" refString="Kronenberger, K, Moore, PG, Halcrow, K, Vollrath, F, 2012. Spinning a Marine Silk for the Purpose of Tube-Building. Journal of Crustacean Biology 32 (2): 191 - 202, DOI: https://doi.org/10.1163/193724011X615532" title="Spinning a Marine Silk for the Purpose of Tube-Building." url="https://doi.org/10.1163/193724011X615532" volume="32" year="2012">Kronenberger et al. 2012</bibRefCitation>
|
||
).
|
||
<taxonomicName id="B5D28B895875ABF43E819472CDF73009" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="735DFEC2C23E87F7BDF5CEF57B748725" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. also presents this characteristic (Fig.
|
||
<figureCitation id="286146510526C2890C19A0693A740FC8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distal end of P 3 and P 4 of Cerapus rivulus sp. nov. Red dash line represents distal end of P 3 dactylus with amphipod silk opening and yellow dash line represents distal end of P 4 dactylus with amphipod silk opening." figureDoi="10.3897/zse.99.107974.figure12" httpUri="https://binary.pensoft.net/fig/947241" pageId="0" pageNumber="557">12</figureCitation>
|
||
). The outer surface of the tube consists of uniformly coarse sediment and detritus with a mix of fragmented fibres and algae (Fig.
|
||
<figureCitation id="FA7FF67D37B3E7BEA832F32A23A2F915" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cerapus rivulus sp. nov. A. Holotype male lateral (THNHM - 19377); B. A specimen in its tube with protruding head and antennae; C. Paratype female front (THNHM- 19379)." figureDoi="10.3897/zse.99.107974.figure2" httpUri="https://binary.pensoft.net/fig/947231" pageId="0" pageNumber="557">2</figureCitation>
|
||
) (small tubes of juvenile attached to the larger tube) (Fig.
|
||
<figureCitation id="8A73DBE7089485E81B9AE8B0FBB0F0A1" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Cerapus rivulus sp. nov. A. Showing a mix of fragmented fibres and algae on its tube; B. Showing small tubes of juvenile attached on the larger tube; C. Uniformly coarse sediment on the outer surface of the tube; D. Ultrastructure of uniformly coarse sediment on the outer surface of the tube (scanning electron microscope); E. Ultrastructure of fine network of amphipod silks on the inner surface of the tube (scanning electron microscope)." figureDoi="10.3897/zse.99.107974.figure13" httpUri="https://binary.pensoft.net/fig/947242" pageId="0" pageNumber="557">13A-D</figureCitation>
|
||
); the inner surface, on the other hand, exhibits a fine network of amphipod silk. Laboratory observations on
|
||
<taxonomicName id="299BF993A87B9CD5CBE396FE0800BC6B" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="E1FCB0CFDDB7B8DFB69F9505A3E9B268" italics="true" pageId="0" pageNumber="557">C. rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.: feeding behaviour, both antennae 1 and 2 were used to gather food and move chunks of organic material back inside their tube. From the gut content analysis, the major food items consist of fungus, algae, diatoms, organic substances and protozoa. Moreover, mating behaviour showed that males wander about the habitat investigating the tube of female individuals. During these encounters, males try to contact females with their antennae. After that, females withdraw the posterior part of the body into their tubes, while males use their urosome to stimulate the ventral surface of the female pleosome. These situations are assumed to be a possible prelude to mating (Fig.
|
||
<figureCitation id="204AD51562194E2F2F844C253514A111" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Series of mating of Cerapus rivulus sp. nov. (created from video of Suppl. material 1)." figureDoi="10.3897/zse.99.107974.figure14" httpUri="https://binary.pensoft.net/fig/947243" pageId="0" pageNumber="557">14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="D34FDA03F960749DFC0C6B936958A1F0" doi="10.3897/zse.99.107974.figure12" httpUri="https://binary.pensoft.net/fig/947241" pageId="0" pageNumber="557" start="Figure 12" startId="F12">
|
||
<paragraph id="84314DA60D7D8BEA4713FB5FBAAB87B0" pageId="0" pageNumber="557">
|
||
<emphasis id="43CA09949EB240CD9564548DCEF9BE53" bold="true" pageId="0" pageNumber="557">Figure 12.</emphasis>
|
||
Distal end of P3 and P4 of
|
||
<taxonomicName id="081FCAA931B9B66556D7E942CBEBE941" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="E77293BA43ECEAAAA4DD88BEB8764EC2" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Red dash line represents distal end of P3 dactylus with amphipod silk opening and yellow dash line represents distal end of P4 dactylus with amphipod silk opening.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CD83BBAA91A8FF6AAC3C93F282D85664" doi="10.3897/zse.99.107974.figure13" httpUri="https://binary.pensoft.net/fig/947242" pageId="0" pageNumber="557" start="Figure 13" startId="F13">
|
||
<paragraph id="8CCA9A2FF93FB172B0D8E702EB6E6616" pageId="0" pageNumber="557">
|
||
<emphasis id="9B4C3C866275F11A012B788CC84EFCD2" bold="true" pageId="0" pageNumber="557">Figure 13.</emphasis>
|
||
<taxonomicName id="7DF4A9C64D7362B7827F53A9C3F94486" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="B1D9B1E99F3FDFAC3EDECACDA83FD751" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="5D72BAF5D76ECBB5A9699DAFC1C9347C" bold="true" pageId="0" pageNumber="557">A.</emphasis>
|
||
Showing a mix of fragmented fibres and algae on its tube;
|
||
<emphasis id="57EB96628B9A736C13EE0B389F3293BF" bold="true" pageId="0" pageNumber="557">B.</emphasis>
|
||
Showing small tubes of juvenile attached on the larger tube;
|
||
<emphasis id="FCC4BF000FF50D1C3D34D90841D8F345" bold="true" pageId="0" pageNumber="557">C</emphasis>
|
||
. Uniformly coarse sediment on the outer surface of the tube;
|
||
<emphasis id="5CB29CA3651ADDDAAA8075AD13ED7985" bold="true" pageId="0" pageNumber="557">D.</emphasis>
|
||
Ultrastructure of uniformly coarse sediment on the outer surface of the tube (scanning electron microscope);
|
||
<emphasis id="573961D0D86CD694A261C25C99B9377E" bold="true" pageId="0" pageNumber="557">E.</emphasis>
|
||
Ultrastructure of fine network of amphipod silks on the inner surface of the tube (scanning electron microscope).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9D747B50379DEF7C6C456EC6DB222D1E" doi="10.3897/zse.99.107974.figure14" httpUri="https://binary.pensoft.net/fig/947243" pageId="0" pageNumber="557" start="Figure 14" startId="F14">
|
||
<paragraph id="EF1BE22181D78A036182064DD8D897FE" pageId="0" pageNumber="557">
|
||
<emphasis id="00DB78C272DAB5999DB3D8578FE79482" bold="true" pageId="0" pageNumber="557">Figure 14.</emphasis>
|
||
Series of mating of
|
||
<taxonomicName id="C86D7600DDC5E023220A52E94BCAB369" authorityName="Katnoum & Keetapithchayakul & Rahim & Wongkamhaeng" authorityYear="2023" class="Malacostraca" family="Ischyroceridae" genus="Cerapus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerapus rivulus" order="Amphipoda" pageId="0" pageNumber="557" phylum="Arthropoda" rank="species" species="rivulus">
|
||
<emphasis id="DE20ACCA91A08CE67057E517248DE43B" italics="true" pageId="0" pageNumber="557">Cerapus rivulus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (created from video of Suppl. material 1).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |