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<mods:title id="603AEF1836A89E54C10E303506BF156D">Revision of the fish family Euclichthyidae (Pisces: Gadiformes) with the description of two new species from the Western Pacific</mods:title>
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<mods:namePart id="4366C951A2324D1FDB8FDA3A2D4F5DEF">Pogonoski, John J.</mods:namePart>
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<treatment id="AD3E8781485D3373FF63AAF7D19007F1" ID-DOI="http://doi.org/10.5281/zenodo.3812373" ID-GBIF-Taxon="163417924" ID-Zenodo-Dep="3812373" LSID="urn:lsid:zoobank.org:act:FC3A18E4-A605-437B-A35A-E3E62E04AFF8" httpUri="http://treatment.plazi.org/id/AD3E8781485D3373FF63AAF7D19007F1" lastPageNumber="244" pageId="13" pageNumber="244">
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<emphasis id="17E3EA85485D3373FF63AAF7D38006F0" bold="true" box="[151,496,1238,1265]" pageId="13" pageNumber="244">
<taxonomicName id="E2974D14485D3373FF63AAF7D3FB06F0" authority="Last &amp; Pogonoski, 2020" authorityName="Last &amp; Pogonoski" authorityYear="2020" box="[151,395,1238,1265]" class="Actinopterygii" family="Euclichthyidae" genus="Euclichthys" kingdom="Animalia" order="Gadiformes" pageId="13" pageNumber="244" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="17E3EA85485D3373FF63AAF7D3FB06F0" bold="true" box="[151,395,1238,1265]" italics="true" pageId="13" pageNumber="244">Euclichthys robertsi</emphasis>
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<subSubSection id="6D8D651C485D3373FF63AADBD05C0756" pageId="13" pageNumber="244" type="description">
<paragraph id="25283697485D3373FF63AADBD0290715" blockId="13.[151,601,1238,1300]" box="[151,601,1273,1300]" pageId="13" pageNumber="244">
<figureCitation id="BDAC2A12485D3373FF63AADBD2800715" box="[151,240,1274,1300]" captionStart="FIGURE 1" captionStartId="7.[151,250,1732,1757]" captionTargetBox="[154,1423,191,1686]" captionTargetId="figure@7.[151,1436,179,1704]" captionTargetPageId="7" captionText="FIGURE 1. Illustrations of Euclichthys species:A. E. polynemus; B. E. microdorsalis sp. nov.; C. E. robertsi sp. nov. Copyright S. Barnes." figureDoi="http://doi.org/10.5281/zenodo.3734410" httpUri="https://zenodo.org/record/3734410/files/figure.png" pageId="13" pageNumber="244">Figs 1C</figureCitation>
,
<figureCitation id="BDAC2A12485D3373FF09AADBD3500715" box="[253,288,1274,1300]" captionStart="FIGURE 3" captionStartId="10.[151,250,1938,1963]" captionTargetBox="[151,1435,777,1913]" captionTargetId="figure@10.[151,1435,777,1913]" captionTargetPageId="10" captionText="FIGURE 3. Caudal skeletons of A. Euclichthys microdorsalis sp. nov. (holotype CSIRO H 719-33), B. E. robertsi sp. nov. (holotype MNHN 2014-2945). Abbreviations: see Fig. 2." figureDoi="http://doi.org/10.5281/zenodo.3734416" httpUri="https://zenodo.org/record/3734416/files/figure.png" pageId="13" pageNumber="244">3B</figureCitation>
,
<figureCitation id="BDAC2A12485D3373FED8AADBD3200715" box="[300,336,1274,1300]" captionStart="FIGURE 4" captionStartId="14.[151,250,1946,1971]" captionTargetBox="[229,1358,181,1919]" captionTargetId="figure@14.[229,1359,181,1920]" captionTargetPageId="14" captionText="FIGURE 4. Proximal views of otoliths of A. E. robertsi sp. nov. (MNHN 2014–2945), B. Euclichthys polynemus (CSIRO H 2578-02), and C. E. microdorsalis sp. nov. (CSIRO H 719-34). Abbreviations: ca—cauda, ci—crista inferior, co—collum, cs—crista superior, ex—excisura, os—ostium, pr—pseudo rostrum, ps—pseudo excisura, ro—rostrum, vg—ventral groove." figureDoi="http://doi.org/10.5281/zenodo.3734418" httpUri="https://zenodo.org/record/3734418/files/figure.png" pageId="13" pageNumber="244">4A</figureCitation>
,
<figureCitation id="BDAC2A12485D3373FEA8AAD8D31C0715" box="[348,364,1273,1300]" captionStart="FIGURE 5" captionStartId="15.[151,250,1450,1475]" captionTargetBox="[164,1422,705,1424]" captionTargetId="figure@15.[164,1422,705,1424]" captionTargetPageId="15" captionText="FIGURE 5. Distributions of Euclichthys polynemus (green diamond—holotype, green circles—other specimens examined, green triangles—other material from Stewart &amp; Roberts, 2015), E. microdorsalis sp. nov. (yellow diamond—holotype, yellow circles—paratypes) and E. robertsi sp. nov. (red diamond—holotype, red circles—other specimens examined)." figureDoi="http://doi.org/10.5281/zenodo.3734420" httpUri="https://zenodo.org/record/3734420/files/figure.png" pageId="13" pageNumber="244">5</figureCitation>
,
<figureCitation id="BDAC2A12485D3373FE8CAADBD3F70715" box="[376,391,1274,1300]" captionStart="FIGURE 8" captionStartId="20.[151,250,1148,1173]" captionTargetBox="[151,1436,561,1123]" captionTargetId="figure@20.[151,1436,561,1123]" captionTargetPageId="20" captionText="FIGURE 8. Lateral view of Euclichthys robertsi sp. nov.: holotype MNHN 2014-2945, 202 mm SL, Norfolk Ridge, New Caledonia (fresh)." figureDoi="http://doi.org/10.5281/zenodo.3734426" httpUri="https://zenodo.org/record/3734426/files/figure.png" pageId="13" pageNumber="244">8</figureCitation>
,
<figureCitation id="BDAC2A12485D3373FE60AADBD3C80715" box="[404,440,1274,1300]" captionStart="FIGURE 9" captionStartId="21.[151,250,1572,1597]" captionTargetBox="[177,1411,194,1547]" captionTargetId="figure@21.[177,1411,181,1547]" captionTargetPageId="21" captionText="FIGURE 9. Lateral view of Euclichthys juveniles: A. E. microdorsalis sp. nov., CSIRO H 579-07, 97 mm SL, near Swains and Saumarez Reefs, Queensland (preserved); B. E. polynemus CSIRO H 7725-08, 85 mm SL, off Coffin Bay, South Australia (fresh); C. E. robertsi sp. nov., CSIRO H 6031-03, 108 mm SL, off Lord Howe Island, Tasman Sea (fresh)." figureDoi="http://doi.org/10.5281/zenodo.3734428" httpUri="https://zenodo.org/record/3734428/files/figure.png" pageId="13" pageNumber="244">9C</figureCitation>
;
<tableCitation id="6815032C485D3373FE37AADBD04F0715" box="[451,575,1274,1300]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="6.[151,239,1401,1425]" captionStartId-1="9.[151,239,157,181]" captionText-0="TABLE 1. Meristic characters for Euclichthys species. E. polynemus E. microdorsalis E. robertsi" captionText-1="TABLE 2. Meristic characters of caudal skeleton of Euclichthys species." pageId="13" pageNumber="244">Tables 1–2</tableCitation>
,
<tableCitation id="6815032C485D3373FDBFAAD8D0290715" box="[587,601,1273,1300]" captionStart="TABLE 5" captionStartId="18.[151,239,154,178]" captionText="TABLE 5. Measurements expressed as percentages of standard length (SL) for Euclichthys robertsi sp. nov.: holotype" pageId="13" pageNumber="244">5</tableCitation>
</paragraph>
<paragraph id="25283697485D3373FF63AB1FD05C0756" blockId="13.[151,1437,1342,1520]" box="[151,556,1342,1367]" pageId="13" pageNumber="244">
<taxonomicName id="E2974D14485D3373FF63AB1FD37D0757" authorityName="McCulloch" authorityYear="1926" box="[151,269,1342,1366]" class="Actinopterygii" family="Euclichthyidae" genus="Euclichthys" kingdom="Animalia" order="Gadiformes" pageId="13" pageNumber="244" phylum="Chordata" rank="genus">
<emphasis id="17E3EA85485D3373FF63AB1FD37D0757" box="[151,269,1342,1366]" italics="true" pageId="13" pageNumber="244">Euclichthys</emphasis>
</taxonomicName>
sp.:
<bibRefCitation id="41064B66485D3373FECAAB1FD0580756" author="Fahay, M. P. &amp; Markle, D. F." box="[318,552,1342,1367]" pageId="13" pageNumber="244" pagination="265 - 283" refId="ref18987" refString="Fahay, M. P. &amp; Markle, D. F. (1984) Gadiformes: development and relationships. In: Moser, H. G. (Ed.), Ontogeny and systematics of fishes. American Society of Ichthyologists and Herpetologists Special Publication, 1, pp. 265 - 283." type="journal article" year="1984">Fahay &amp; Markle, 1984</bibRefCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="25283697485D3373FF63AB7CD6900774" blockId="13.[151,1437,1342,1520]" box="[151,1248,1372,1397]" pageId="13" pageNumber="244">
<taxonomicName id="E2974D14485D3373FF63AB7CD37D0774" ID-CoL="4F8M" authorityName="McCulloch" authorityYear="1926" box="[151,269,1373,1397]" class="Actinopterygii" family="Euclichthyidae" genus="Euclichthys" kingdom="Animalia" order="Gadiformes" pageId="13" pageNumber="244" phylum="Chordata" rank="genus">
<emphasis id="17E3EA85485D3373FF63AB7CD37D0774" box="[151,269,1373,1397]" italics="true" pageId="13" pageNumber="244">Euclichthys</emphasis>
</taxonomicName>
sp. NFZ1:
<bibRefCitation id="41064B66485D3373FE75AB7DD0450774" author="Clark, M. R. &amp; Roberts, C. D. &amp; Williams, A. &amp; Last, P. R." box="[385,565,1372,1397]" pageId="13" pageNumber="244" refId="ref18791" refString="Clark, M. R., Roberts, C. D., Williams, A. &amp; Last, P. R. (2003) Voyage report of a biodiversity survey of seamounts and slopes of the Norfolk Ridge and Lord Howe Rise (NORFANZ). May-June 2003. Unpublished report held in NIWA library, Wellington 43 pp." type="book" year="2003">
Clark
<emphasis id="17E3EA85485D3373FE4BAB7FD39C0774" box="[447,492,1373,1397]" italics="true" pageId="13" pageNumber="244">et al</emphasis>
., 2003
</bibRefCitation>
;
<bibRefCitation id="41064B66485D3373FDCAAB7DD1660774" author="Williams, A. &amp; Gowlett-Holmes, K. &amp; Althaus, F." box="[574,790,1372,1397]" pageId="13" pageNumber="244" refId="ref21172" refString="Williams, A., Gowlett-Holmes, K. &amp; Althaus, F. (2006) Biodiversity survey of seamounts and slopes of the Norfolk Ridge and Lord Howe Rise. Final Report to the Environment and Heritage (National Oceans Office), Hobart, 378 pp." type="book" year="2006">
Williams
<emphasis id="17E3EA85485D3373FD54AB7FD0BD0774" box="[672,717,1373,1397]" italics="true" pageId="13" pageNumber="244">et al</emphasis>
., 2006
</bibRefCitation>
;
<bibRefCitation id="41064B66485D3373FCD4AB7DD67A0774" author="Clark, M. R. &amp; Roberts, C. D." box="[800,1034,1372,1397]" pageId="13" pageNumber="244" refId="ref18737" refString="Clark, M. R. &amp; Roberts, C. D. (2008) Fish and invertebrate biodiversity on the Norfolk Ridge and Lord Howe Rise, Tasman Sea (NORFANZ voyage, 2003). Aquatic Environment and Biodiversity Report No. 28. NIWA, Wellington, 131 pp." type="book" year="2008">Clark &amp; Roberts, 2008</bibRefCitation>
;
<bibRefCitation id="41064B66485D3373FBE0AB7DD6AC0774" author="Zintzen, V. &amp; Roberts, C. D. &amp; Clark, M. R. &amp; Williams, A. &amp; Althaus, F. &amp; Last, P. R." box="[1044,1244,1372,1397]" pageId="13" pageNumber="244" pagination="933 - 947" refId="ref21291" refString="Zintzen, V., Roberts, C. D., Clark, M. R., Williams, A., Althaus, F. &amp; Last, P. R. (2011) Composition, distribution and regional affinities of the deepwater ichthyofauna of the Lord Howe Rise and Norfolk Ridge, south-west Pacific Ocean. Deep Sea Research II, 58, 933 - 947. https: // doi. org / 10.1016 / j. dsr 2.2010.10.049" type="journal article" year="2011">
Zintzen
<emphasis id="17E3EA85485D3373FB9CAB7FD6E40774" box="[1128,1172,1373,1397]" italics="true" pageId="13" pageNumber="244">et al</emphasis>
., 2011
</bibRefCitation>
.
</paragraph>
<paragraph id="25283697485D3373FF63AB5DD1BE07B2" blockId="13.[151,1437,1342,1520]" pageId="13" pageNumber="244">
<taxonomicName id="E2974D14485D3373FF63AB5DD30F0795" ID-CoL="3BTWM" authorityName="McCulloch" authorityYear="1926" box="[151,383,1404,1428]" class="Actinopterygii" family="Euclichthyidae" genus="Euclichthys" kingdom="Animalia" order="Gadiformes" pageId="13" pageNumber="244" phylum="Chordata" rank="species" species="polynemus">
<emphasis id="17E3EA85485D3373FF63AB5DD30F0795" box="[151,383,1404,1428]" italics="true" pageId="13" pageNumber="244">Euclichthys polynemus</emphasis>
</taxonomicName>
(non McCulloch):
<bibRefCitation id="41064B66485D3373FDB6AB5AD08A0795" author="Fricke, R. &amp; Kulbicki, M. &amp; Wantiez, L." box="[578,762,1403,1428]" pageId="13" pageNumber="244" pagination="341 - 463" refId="ref19091" refString="Fricke, R., Kulbicki, M. &amp; Wantiez, L. (2011) Checklist of the fishes of New Caledonia, and their distribution in the Southwest Pacific Ocean (Pisces). Stuttgarter Beitrage zur Naturkunde, A (4), 341 - 463." type="journal article" year="2011">
Fricke
<emphasis id="17E3EA85485D3373FD7DAB5CD0C50795" box="[649,693,1404,1428]" italics="true" pageId="13" pageNumber="244">et al</emphasis>
., 2011
</bibRefCitation>
(in part);
<bibRefCitation id="41064B66485D3373FCABAB5AD6E40795" author="Grandperrin, R. &amp; Lehodey, P." box="[863,1172,1403,1428]" pageId="13" pageNumber="244" pagination="1 - 40" refId="ref19412" refString="Grandperrin, R. &amp; Lehodey, P. (1992) Campagne BERYX 2 de peche au chalut de fond sur trois monts sous-marins du Sud-East de la Zone Economique de Nouvelle Caledonie (N. O. &quot; Alis &quot;, 22 - 31 octobre 1991). ORSTOM Rapport de Missions: Sciences de la Mer: Biologie marine, 11, 1 - 40." type="journal article" year="1992">Grandperrin &amp; Lehodey, 1992</bibRefCitation>
;
<bibRefCitation id="41064B66485D3373FB6AAB5AD75A0795" author="Markle, D. F." box="[1182,1322,1403,1428]" pageId="13" pageNumber="244" pagination="59 - 88" refId="ref20132" refString="Markle, D. F. (1989) Aspects of character homology and phylogeny of the Gadiformes. In: Cohen, D. M. (Ed.), Papers on the systematics of gadiform fishes, 32. Natural History Museum Los Angeles County, Los Angeles, California, pp. 59 - 88." type="book chapter" year="1989">Markle, 1989</bibRefCitation>
;
<bibRefCitation id="41064B66485D3373FAC0AB5AD33B07B2" author="Roberts, C. D. &amp; Paulin, C. D." pageId="13" pageNumber="244" pagination="43 - 50" refId="ref20843" refString="Roberts, C. D. &amp; Paulin, C. D. (1997) First record of the Eucla cod, Euclichthys polynemus McCulloch, (Teleostei, Paracanthopterygii, Euclichthyidae) from New Caledonia, southwest Pacific Ocean, with notes on morphological characters. In: Seret, B. (Ed.), Resultats des Campagnes MUSORSTOM, 17. Memoires du Museum national d'Histoire naturelle, 174, pp. 43 - 50." type="journal article" year="1997">Roberts &amp; Paulin, 1997</bibRefCitation>
;
<bibRefCitation id="41064B66485D3373FEA1ABBAD07D07B2" author="Hoese, D. F. &amp; Bray, D. J. &amp; Paxton, J. R. &amp; Allen, G. R." box="[341,525,1434,1459]" pageId="13" pageNumber="244" pagination="1 - 670" refId="ref19528" refString="Hoese, D. F., Bray, D. J., Paxton, J. R. &amp; Allen, G. R. (2006) Fishes. In: Beesley, P. L. &amp; Wells, A. (Eds.), Zoological Catalogue of Australia, 35. Part 1. ABRS &amp; CSIRO Publishing, Collingwood., pp. 1 - 670." type="book chapter" year="2006">
Hoese
<emphasis id="17E3EA85485D3373FE6EABBAD3B707B3" box="[410,455,1434,1458]" italics="true" pageId="13" pageNumber="244">et al</emphasis>
., 2006
</bibRefCitation>
(in part);
<bibRefCitation id="41064B66485D3373FD86ABBAD11F07B2" author="Stewart, A. L. &amp; Roberts, C. D." box="[626,879,1434,1459]" pageId="13" pageNumber="244" pagination="745 - 746" refId="ref21039" refString="Stewart, A. L. &amp; Roberts, C. D. (2015) 102 Family Euclichthyidae. In: Roberts, C. D., Stewart, A. L. &amp; Struthers, C. D. (Eds.), The Fishes of New Zealand, 3, pp. 745 - 746." type="journal article" year="2015">Stewart &amp; Roberts, 2015</bibRefCitation>
(in part).
</paragraph>
<paragraph id="25283697485D3373FF63AB98D11F07D0" blockId="13.[151,1437,1342,1520]" box="[151,879,1464,1489]" pageId="13" pageNumber="244">
<taxonomicName id="E2974D14485D3373FF63AB98D37D07D0" ID-CoL="4F8M" authorityName="McCulloch" authorityYear="1926" box="[151,269,1465,1489]" class="Actinopterygii" family="Euclichthyidae" genus="Euclichthys" kingdom="Animalia" order="Gadiformes" pageId="13" pageNumber="244" phylum="Chordata" rank="genus">
<emphasis id="17E3EA85485D3373FF63AB98D37D07D0" box="[151,269,1465,1489]" italics="true" pageId="13" pageNumber="244">Euclichthys</emphasis>
</taxonomicName>
sp. 2 [NSW, in National Fish Collection]:
<bibRefCitation id="41064B66485D3373FD4AAB9BD11B07D0" author="Rees, A. J. J. &amp; Yearsley, G. K. &amp; Gowlett-Holmes, K. &amp; Pogonoski, J." box="[702,875,1465,1489]" pageId="13" pageNumber="244" refId="ref20714" refString="Rees, A. J. J., Yearsley, G. K., Gowlett-Holmes, K. &amp; Pogonoski, J. (2020) Codes for Australian Aquatic Biota. CSIRO Marine and Atmospheric Research, World Wide Web electronic publication, 1999 onwards. Available from: http: // www. cmar. csiro. au / caab / (accessed 10 January 2020)" type="url" year="2020">
Rees
<emphasis id="17E3EA85485D3373FD01AB9BD15907D0" box="[757,809,1465,1489]" italics="true" pageId="13" pageNumber="244">et al.</emphasis>
, 2020
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6D8D651C485D3373FF63ABF9D19007F1" box="[151,992,1495,1520]" pageId="13" pageNumber="244" type="description">
<paragraph id="25283697485D3373FF63ABF9D19007F1" blockId="13.[151,1437,1342,1520]" box="[151,992,1495,1520]" pageId="13" pageNumber="244">
<taxonomicName id="E2974D14485D3373FF63ABF9D37D07F1" authorityName="McCulloch" authorityYear="1926" box="[151,269,1496,1520]" class="Actinopterygii" family="Euclichthyidae" genus="Euclichthys" kingdom="Animalia" order="Gadiformes" pageId="13" pageNumber="244" phylum="Chordata" rank="genus">
<emphasis id="17E3EA85485D3373FF63ABF9D37D07F1" box="[151,269,1496,1520]" italics="true" pageId="13" pageNumber="244">Euclichthys</emphasis>
</taxonomicName>
sp. 3 [Lord Howe Rise, in National Fish Collection]:
<bibRefCitation id="41064B66485D3373FCDBABF9D1AC07F1" author="Rees, A. J. J. &amp; Yearsley, G. K. &amp; Gowlett-Holmes, K. &amp; Pogonoski, J." box="[815,988,1496,1520]" pageId="13" pageNumber="244" refId="ref20714" refString="Rees, A. J. J., Yearsley, G. K., Gowlett-Holmes, K. &amp; Pogonoski, J. (2020) Codes for Australian Aquatic Biota. CSIRO Marine and Atmospheric Research, World Wide Web electronic publication, 1999 onwards. Available from: http: // www. cmar. csiro. au / caab / (accessed 10 January 2020)" type="url" year="2020">
Rees
<emphasis id="17E3EA85485D3373FC93ABF8D1EA07F1" box="[871,922,1496,1520]" italics="true" pageId="13" pageNumber="244">et al.</emphasis>
, 2020
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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