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<document id="589A5DE19B4F6AEF5816EB8E84BBEED9" ID-CLB-Dataset="188516" ID-DOI="10.11646/zootaxa.5209.1.1" ID-GBIF-Dataset="51b10d7e-0ef3-496e-9ec5-d4bcd31e9f91" ID-ISSN="1175-5326" ID-Zenodo-Dep="7322341" ID-ZooBank="78B932B7-0194-4DEA-A4FA-40A457ACA662" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668501069430" checkinUser="plazi" docAuthor="Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong &amp; Cognato, Anthony I." docDate="2022" docId="7D518786FFB77108FF70278B7B41F51D" docLanguage="en" docName="zootaxa.5209.1.1.pdf" docOrigin="Zootaxa 5209 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tricosa hipparion Smith, Beaver &amp; Cognato 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="19" masterDocId="8168FFFEFFA6711AFFE726327E6CF45C" masterDocTitle="New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="18" updateTime="1699411528783" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="FD9F4DF28B1C2EB6B973022CA99C9FDE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F3B42C9B335601A8D0197F3E35786C8E">
<mods:title id="E79CC910065EFCC8AFFF95EEA8DDBB24">New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae)</mods:title>
</mods:titleInfo>
<mods:name id="F4EAEE5B842B6E2FB1512868182139DA" type="personal">
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<mods:namePart id="9CE3E6BB72D33A6B6F59AA5971714577">Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier id="1CC9FB478FCA7E73C494177C71019E16" type="ORCID">0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation id="0DEB42D6897CF682B0C98C0FD91FDB04">Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA. smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736</mods:affiliation>
<mods:nameIdentifier id="1FC74E2CA88333D9E91AB8E5575CE1BF" type="email">smith462@msu.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="C5C294AE72D1426F80FC68E240EF52D9" type="personal">
<mods:role id="EE3A4D94C1C20093331883C5AC361932">
<mods:roleTerm id="4B567FBEFC1E7F16113046DF5C936714">Author</mods:roleTerm>
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<mods:namePart id="5299D613B5E91C77CE0A0539CD91503B">Beaver, Roger A.</mods:namePart>
<mods:affiliation id="F66FE774A00C30B956E2B5DB96854707">161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand.</mods:affiliation>
</mods:name>
<mods:name id="8344AAB3222A6ABAFBF4F427941F8274" type="personal">
<mods:role id="F7D7C00F31377354F9FFDB98C199B994">
<mods:roleTerm id="A6DB1322F59F059AD0ED55446F78B099">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE1C19559C0C6EE275F902F29214CABC">Pham, Thai Hong</mods:namePart>
<mods:affiliation id="55B39A2F3C2B548A19FFF0B13651338C">Mientrung Institute for Scientific Research (MISR), Vietnam National Museum of Nature, Vietnam Academy of Science and Technology (VAST), 321 Huynh Thuc Khang St, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA. cognato @ msu. edu; https: // orcid. org / 0000 - 0001 - 6436 - 2447</mods:affiliation>
</mods:name>
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<mods:namePart id="ABDE00454D92E44DBAB36CD03A4ADE5B">Cognato, Anthony I.</mods:namePart>
<mods:nameIdentifier id="E884EBE7EC89F7C5E25B4536E969650A" type="ORCID">0000-0001-6436-2447</mods:nameIdentifier>
<mods:nameIdentifier id="359D3FA05AA8F3891AEF0B74C68E1FF0" type="email">cognato@msu.edu</mods:nameIdentifier>
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<mods:typeOfResource id="A647AD4CCA9A5E5716E249FE9DCA60E2">text</mods:typeOfResource>
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<mods:title id="5F0C874FF46AD64CFA4300BC721C4A9B">Zootaxa</mods:title>
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<mods:part id="1BB11DC54E4AC1A987ED30014430A19E">
<mods:date id="C6ED035FBF6E71FBAF6F4ACD5644DF66">2022</mods:date>
<mods:detail id="D71C8ED140C0F5D8734C30D5AAC4E9BF" type="pubDate">
<mods:number id="A54B5DD9EC7863D519B52073077D9E20">2022-11-15</mods:number>
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<mods:detail id="8BF625D246A6BDBD06DC1132D6F3AC8D" type="volume">
<mods:number id="67F44DCD72D77225A7886E79590F4C82">5209</mods:number>
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<mods:number id="EFF57FA286EE0707841ABF95C908955C">1</mods:number>
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<mods:start id="5D8171E1F9B38214072C6CCB879AC7A8">1</mods:start>
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<mods:classification id="D3F15152C6FD8B83FB42EE64557FBA4D">journal article</mods:classification>
<mods:identifier id="35339261923C58036884484AC0177F71" type="CLB-Dataset">188516</mods:identifier>
<mods:identifier id="86EE51640B93ADBA1B3D9A7D025E6994" type="DOI">10.11646/zootaxa.5209.1.1</mods:identifier>
<mods:identifier id="92D6DF8776C3ED82A207B6D75DCFBCDC" type="GBIF-Dataset">51b10d7e-0ef3-496e-9ec5-d4bcd31e9f91</mods:identifier>
<mods:identifier id="A1BC3DF8DBFEBA5868595E23943C235A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="7D518786FFB77108FF70278B7B41F51D" ID-DOI="http://doi.org/10.5281/zenodo.7322388" ID-GBIF-Taxon="205078259" ID-Zenodo-Dep="7322388" LSID="urn:lsid:plazi:treatment:7D518786FFB77108FF70278B7B41F51D" httpUri="http://treatment.plazi.org/id/7D518786FFB77108FF70278B7B41F51D" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="BDE2651BFFB7710BFF70278B7D72F58F" box="[151,798,441,468]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="F5473690FFB7710BFF70278B7D72F58F" blockId="17.[151,1186,441,540]" box="[151,798,441,468]" pageId="17" pageNumber="18">
<heading id="AE0F81FCFFB7710BFF70278B7D72F58F" bold="true" box="[151,798,441,468]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="C78CEA82FFB7710BFF70278B7D72F58F" bold="true" box="[151,798,441,468]" pageId="17" pageNumber="18">
<taxonomicName id="32F84D13FFB7710BFF70278B7CD5F58F" authority="Smith, Beaver &amp; Cognato" authorityName="Smith, Beaver &amp; Cognato" authorityYear="2022" box="[151,697,441,468]" class="Insecta" family="Curculionidae" genus="Tricosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hipparion" status="sp. nov.">
<emphasis id="C78CEA82FFB7710BFF70278B7F1CF588" bold="true" box="[151,368,441,468]" italics="true" pageId="17" pageNumber="18">Tricosa hipparion</emphasis>
Smith, Beaver &amp; Cognato
</taxonomicName>
<taxonomicNameLabel id="DCBF57F9FFB7710BFD27278B7D72F58F" box="[704,798,441,467]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB7710BFF7027EF7ACEF640" pageId="17" pageNumber="18" type="description">
<paragraph id="F5473690FFB7710BFF7027EF7F4BF5A4" blockId="17.[151,1186,441,540]" box="[151,295,477,504]" pageId="17" pageNumber="18">
(
<figureCitation id="6DC32A15FFB7710BFF7827EF7F4CF5A4" box="[159,288,477,504]" captionStart="FIGURES 57‒60" captionStartId="17.[151,264,1140,1164]" captionTargetBox="[151,1436,852,1116]" captionTargetId="figure-145@17.[151,1436,851,1116]" captionTargetPageId="17" captionText="FIGURES 57‒60. Tricosa hipparion holotype, 3.1 mm. Dorsal, lateral, posterior oblique and anterior views." figureDoi="http://doi.org/10.5281/zenodo.7322366" httpUri="https://zenodo.org/record/7322366/files/figure.png" pageId="17" pageNumber="18">Figs 57–60</figureCitation>
)
</paragraph>
<paragraph id="F5473690FFB7710BFF7024337ACEF640" blockId="17.[151,1186,441,540]" box="[151,1186,513,540]" pageId="17" pageNumber="18">
http://zoobank.org/
<uri id="81693A92FFB7710BFE9324337ACEF640" box="[372,1186,513,540]" pageId="17" pageNumber="18">
urn:lsid:zoobank.org:act:
<uuid id="815E0C45FFB7710BFD6124337ACEF640" box="[646,1186,513,540]" pageId="17" pageNumber="18">024CA706-3D66-4B20-9CA6-5E5B543A4DE2</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB7710BFF70247B7C7CF6F0" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="F5473690FFB7710BFF70247B7CD1F6DB" blockId="17.[151,1436,585,828]" pageId="17" pageNumber="18">
<emphasis id="C78CEA82FFB7710BFF70247B7F26F63F" bold="true" box="[151,330,585,611]" pageId="17" pageNumber="18">Type material.</emphasis>
<materialsCitation id="45903CCDFFB7710BFEB1247B7CD1F6DB" ID-GBIF-Occurrence="3966310412" collectingDate="2006-11-11" collectionCode="NMNH" collectorName="J. Hulcr" country="Malaysia" location="Danum Valley" municipality="Rhino Ridge" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="2A438832FFB7710BFEB1247B7FADF63F" box="[342,449,585,611]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
, female:
<collectingCountry id="8DEF7600FFB7710BFDD6247B7CA9F63F" box="[561,709,585,611]" name="Malaysia" pageId="17" pageNumber="18">MALAYSIA</collectingCountry>
,
<collectingRegion id="373CF872FFB7710BFD32247B7D70F63F" box="[725,796,585,611]" country="Malaysia" name="Sabah" pageId="17" pageNumber="18">Sabah</collectingRegion>
,
<collectingMunicipality id="1523ACEAFFB7710BFCCB247B7DAEF63F" box="[812,962,585,611]" pageId="17" pageNumber="18">Rhino Ridge</collectingMunicipality>
,
<location id="F027604BFFB7710BFC3524787A15F63F" LSID="urn:lsid:plazi:treatment:7D518786FFB77108FF70278B7B41F51D:F027604BFFB7710BFC3524787A15F63F" box="[978,1145,585,612]" country="Malaysia" municipality="Rhino Ridge" name="Danum Valley" pageId="17" pageNumber="18" stateProvince="Sabah">Danum Valley</location>
,
<date id="81461050FFB7710BFB6E247B7B6EF63F" box="[1161,1282,585,611]" pageId="17" pageNumber="18" value="2006-11-11">
<collectingDate id="9102E9B8FFB7710BFB6E247B7B6EF63F" box="[1161,1282,585,611]" pageId="17" pageNumber="18" value="2006-11-11">11.xi.2006</collectingDate>
</date>
,
<collectorName id="580D5346FFB7710BFAF524787B1EF63F" box="[1298,1394,585,612]" pageId="17" pageNumber="18">J. Hulcr</collectorName>
, ex
<taxonomicName id="32F84D13FFB7710BFF70245C7F0BF6D4" authority=", SAX" authorityName="SAX" box="[151,359,621,648]" class="Magnoliopsida" family="Burseraceae" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="family">Burseraceae, SAX</taxonomicName>
503 [DNA voucher] (
<collectionCode id="93E9AE55FFB7710BFDB2245F7CDDF6DB" box="[597,689,621,647]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F5473690FFB7710BFF2024A37C7CF6F0" blockId="17.[151,1436,585,828]" box="[199,528,657,684]" pageId="17" pageNumber="18">
<emphasis id="C78CEA82FFB7710BFF2024A37F2CF6F7" bold="true" box="[199,320,657,683]" pageId="17" pageNumber="18">GenBank.</emphasis>
<collectingCountry id="8DEF7600FFB7710BFEA024A37FEAF6F7" box="[327,390,657,683]" name="Chad" pageId="17" pageNumber="18">CAD</collectingCountry>
:
<accessionNumber id="EAABAB73FFB7710BFE7624A37C60F6F0" box="[401,524,657,684]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OP607245" pageId="17" pageNumber="18">OP607245</accessionNumber>
.
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB7710BFF2024877F53F767" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="F5473690FFB7710BFF2024877F24F6AF" blockId="17.[151,1436,585,828]" pageId="17" pageNumber="18">
<emphasis id="C78CEA82FFB7710BFF2024877F2EF693" bold="true" box="[199,322,693,719]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
This species is distinguished by the elytral disc broadly, deeply transversely impressed with a saddlelike depression.
</paragraph>
<paragraph id="F5473690FFB7710BFF2024CF7F53F767" blockId="17.[151,1436,585,828]" pageId="17" pageNumber="18">
<emphasis id="C78CEA82FFB7710BFF2024CF7FEFF74B" bold="true" box="[199,387,765,791]" pageId="17" pageNumber="18">Similar species.</emphasis>
<taxonomicName id="32F84D13FFB7710BFE6B24CC7D96F74B" authority="Cognato, Smith &amp; Beaver, 2020 a" authorityName="Cognato, Smith &amp; Beaver" authorityYear="2020" box="[396,1018,765,792]" class="Insecta" family="Curculionidae" genus="Tricosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="C78CEA82FFB7710BFE6B24CC7C02F74B" box="[396,622,766,791]" italics="true" pageId="17" pageNumber="18">Tricosa cattienensis</emphasis>
<bibRefCitation id="91694B61FFB7710BFD9F24CF7D96F74B" author="Cognato, A. I. &amp; Smith, S. M. &amp; Beaver, R. A." box="[632,1018,765,792]" pageId="17" pageNumber="18" pagination="540 - 554" refId="ref26083" refString="Cognato, A. I., Smith, S. M. &amp; Beaver, R. A. (2020 a) Two new genera of Oriental xyleborine ambrosia beetles (Coleoptera, Curculionidae: Scolytinae). Zootaxa, 4722 (6), 540 - 554. https: // doi. org / 10.11646 / zootaxa. 4722.6.2" type="journal article" year="2020">Cognato, Smith &amp; Beaver, 2020a</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="32F84D13FFB7710BFBEF24CC7F57F767" authority="Cognato, Smith &amp; Beaver, 2020 a" authorityName="Cognato, Smith &amp; Beaver" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Tricosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="indochinensis">
<emphasis id="C78CEA82FFB7710BFBEF24CC7AACF74B" box="[1032,1216,765,791]" italics="true" pageId="17" pageNumber="18">T. indochinensis</emphasis>
<bibRefCitation id="91694B61FFB7710BFB2D24CF7F57F767" author="Cognato, A. I. &amp; Smith, S. M. &amp; Beaver, R. A." pageId="17" pageNumber="18" pagination="540 - 554" refId="ref26083" refString="Cognato, A. I., Smith, S. M. &amp; Beaver, R. A. (2020 a) Two new genera of Oriental xyleborine ambrosia beetles (Coleoptera, Curculionidae: Scolytinae). Zootaxa, 4722 (6), 540 - 554. https: // doi. org / 10.11646 / zootaxa. 4722.6.2" type="journal article" year="2020">Cognato, Smith &amp; Beaver, 2020a</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="A1876618FFB7710BFF7022467A8BF0D0" ID-DOI="http://doi.org/10.5281/zenodo.7322366" ID-Zenodo-Dep="7322366" box="[151,1255,1140,1164]" httpUri="https://zenodo.org/record/7322366/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,264,1140,1164]" targetBox="[151,1436,852,1116]" targetPageId="17">
<paragraph id="F5473690FFB7710BFF7022467A8BF0D0" blockId="17.[151,1255,1140,1164]" box="[151,1255,1140,1164]" pageId="17" pageNumber="18">
<emphasis id="C78CEA82FFB7710BFF7022467F38F0D0" bold="true" box="[151,340,1140,1164]" pageId="17" pageNumber="18">FIGURES 57‒60.</emphasis>
<taxonomicName id="32F84D13FFB7710BFEBD22477C7CF0D0" authorityName="Smith, Beaver &amp; Cognato" authorityYear="2022" box="[346,528,1140,1164]" class="Insecta" family="Curculionidae" genus="Tricosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hipparion">
<emphasis id="C78CEA82FFB7710BFEBD22477C7CF0D0" box="[346,528,1140,1164]" italics="true" pageId="17" pageNumber="18">Tricosa hipparion</emphasis>
</taxonomicName>
holotype, 3.1 mm. Dorsal, lateral, posterior oblique and anterior views.
</paragraph>
</caption>
<subSubSection id="BDE2651BFFB77108FF2022887AD6F489" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="F5473690FFB77108FF2022887AD6F489" blockId="17.[151,1437,1210,2029]" lastBlockId="18.[151,1436,151,321]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="C78CEA82FFB7710BFF2022887FADF088" bold="true" box="[199,449,1210,1236]" pageId="17" pageNumber="18">Description (female).</emphasis>
<quantity id="32009B75FFB7710BFE2022897C71F089" box="[455,541,1211,1237]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="17" pageNumber="18" unit="mm" value="3.1">3.1 mm</quantity>
long (n = 1);
<quantity id="32009B75FFB7710BFD5322897D7BF089" box="[692,791,1211,1237]" metricMagnitude="-3" metricUnit="m" metricValue="2.81" pageId="17" pageNumber="18" unit="mm" value="2.81">2.81 mm</quantity>
× as long as wide. Head and pronotum ferruginous, elytra dark brown, legs and antennae light brown.
<emphasis id="C78CEA82FFB7710BFD6122ED7CA3F0A5" bold="true" box="[646,719,1247,1273]" pageId="17" pageNumber="18">Head:</emphasis>
epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes; surface shagreened, dull, punctate; punctures above epistoma small, fine, shallow, punctures larger, deeper on frons. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum slightly impressed below genae, narrowly triangular. Scape regularly thick, shorter than club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 as long as pedicel. Club approximately circular and flat,
<typeStatus id="2A438832FFB7710BFDA923A17C12F1F1" box="[590,638,1427,1453]" pageId="17" pageNumber="18">type</typeStatus>
3; segment 1 corneous, transverse on anterior face, occupying basal ~1/3; segment 2 narrow, corneous; segments 1 and 2 present on posterior face.
<emphasis id="C78CEA82FFB7710BFBF123857AE3F18D" bold="true" box="[1046,1167,1463,1489]" pageId="17" pageNumber="18">Pronotum</emphasis>
1.09 × as long as wide. In dorsal view elongate and parallel sided,
<typeStatus id="2A438832FFB7710BFD6323E97CD8F1A9" box="[644,692,1499,1525]" pageId="17" pageNumber="18">type</typeStatus>
7, sides parallel in basal 2/3, rounded anteriorly; anterior margin with a row of six serrations. In lateral view elongate, disc longer than anterior slope,
<typeStatus id="2A438832FFB7710BFBB623CD7AEDF245" box="[1105,1153,1535,1561]" pageId="17" pageNumber="18">type</typeStatus>
7, summit low, on apical 2/5. Anterior slope with densely spaced narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, erect hair-like setae. Disc subshiny, alutaceous, finely punctate, finely setose, setae short, erect, hair-like, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded.
<emphasis id="C78CEA82FFB7710BFDF820BC7CFBF2F4" bold="true" box="[543,663,1678,1704]" pageId="17" pageNumber="18">Scutellum</emphasis>
small, triangular, flat, flush with elytra.
<emphasis id="C78CEA82FFB7710BFB9520BD7AABF2F5" bold="true" box="[1138,1223,1679,1705]" pageId="17" pageNumber="18">Elytra:</emphasis>
1.73 × as long as wide, 1.58 × as long as pronotum. Base transverse, margins oblique; humeral angles rounded; attenuate at apical half, apex broadly rounded. Disc broadly, deeply transversely impressed with a saddle-like depression, surface smooth, shining, unarmed; striae seriate punctate, punctures separated by less than one diameter of a puncture, glabrous; interstrial minutely uniseriate punctate, widely spaced, each with a long, erect seta 1.5 × as long as interstriae 2. Declivity occupying ~2/5 of elytra, gradual, face flat, opalescent; striae impressed, strial punctures as large as those of disc; interstriae impunctate, interstriae tubercles arranged in an alternating pattern of a small denticle followed by a large denticle, with the apices curved ventrad; interstrial setae as long or twice as long as those of disc. Posterolateral margins rounded and granulate.
<emphasis id="C78CEA82FFB7710BFCA9219D7DFCF395" bold="true" box="[846,912,1967,1993]" pageId="17" pageNumber="18">Legs:</emphasis>
procoxae contiguous, prosternal posterocoxal piece conical, slightly inflated. Protibiae distinctly triangular, broadest at apical half, posterior face flat, unarmed; five large denticles present on outer margin of apical third. Meso- and metatibiae obliquely triangular, flattened, posterior face unarmed; metatibiae bearing eight moderate socketed denticles on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF2026EC7C58F4A5" box="[199,564,222,249]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="F5473690FFB47108FF2026EC7C58F4A5" blockId="18.[151,1436,151,321]" box="[199,564,222,249]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF2026EC7F0DF4A4" bold="true" box="[199,353,222,248]" pageId="18" pageNumber="19">Distribution.</emphasis>
<collectingCountry id="8DEF7600FFB47108FE8026ED7FA2F4A5" box="[359,462,223,249]" name="Malaysia" pageId="18" pageNumber="19">Malaysia</collectingCountry>
(
<collectingRegion id="373CF872FFB47108FE3A26ED7C4BF4A5" box="[477,551,223,249]" country="Malaysia" name="Sabah" pageId="18" pageNumber="19">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF2027317D09F541" box="[199,869,259,285]" pageId="18" pageNumber="19" type="biology_ecology">
<paragraph id="F5473690FFB47108FF2027317D09F541" blockId="18.[151,1436,151,321]" box="[199,869,259,285]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF2027317F3AF541" bold="true" box="[199,342,259,285]" pageId="18" pageNumber="19">Host plants.</emphasis>
The
<typeStatus id="2A438832FFB47108FE7727317F9EF541" box="[400,498,259,285]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
was collected from
<taxonomicName id="32F84D13FFB47108FD3227317D0EF541" box="[725,866,259,285]" class="Magnoliopsida" family="Burseraceae" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="family">Burseraceae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF2027157B41F51D" box="[199,1325,295,321]" pageId="18" pageNumber="19" type="etymology">
<paragraph id="F5473690FFB47108FF2027157B41F51D" blockId="18.[151,1436,151,321]" box="[199,1325,295,321]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF2027157F22F51D" bold="true" box="[199,334,295,321]" pageId="18" pageNumber="19">Etymology.</emphasis>
G.
<taxonomicName id="32F84D13FFB47108FE9027157F89F51D" authorityName="Smith, Beaver &amp; Cognato" authorityYear="2022" box="[375,485,295,321]" class="Insecta" family="Curculionidae" genus="Tricosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hipparion">
<emphasis id="C78CEA82FFB47108FE9027157F89F51D" box="[375,485,295,321]" italics="true" pageId="18" pageNumber="19">hipparion</emphasis>
</taxonomicName>
= little horse. In reference to the saddle-shaped elytra. Noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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