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<document id="8CAEE2D233B12355C935D8B41D0C99BB" ID-CLB-Dataset="188516" ID-DOI="10.11646/zootaxa.5209.1.1" ID-GBIF-Dataset="51b10d7e-0ef3-496e-9ec5-d4bcd31e9f91" ID-ISSN="1175-5326" ID-Zenodo-Dep="7322341" ID-ZooBank="78B932B7-0194-4DEA-A4FA-40A457ACA662" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668501069430" checkinUser="plazi" docAuthor="Smith, Sarah M., Beaver, Roger A., Pham, Thai Hong &amp; Cognato, Anthony I." docDate="2022" docId="7D518786FFB47108FF7027B77F78F3AD" docLanguage="en" docName="zootaxa.5209.1.1.pdf" docOrigin="Zootaxa 5209 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xyleborinus acanthopteron Smith, Beaver &amp; Cognato 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="19" masterDocId="8168FFFEFFA6711AFFE726327E6CF45C" masterDocTitle="New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="19" updateTime="1699411528783" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="56CE8440219896F3FB21B4B463C818C9">New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae)</mods:title>
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<mods:name id="475F179B849AA7A4D953BA6962E6E762" type="personal">
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<mods:namePart id="D1AE1E3A5C30BD94DD97DC299CAC24C6">Smith, Sarah M.</mods:namePart>
<mods:nameIdentifier id="5CB4BC1D4C0E9B9F7D3690E9CB090652" type="ORCID">0000-0002-5173-3736</mods:nameIdentifier>
<mods:affiliation id="31A0FC99E642D80A59007E9CA4D4126B">Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA. smith 462 @ msu. edu; https: // orcid. org / 0000 - 0002 - 5173 - 3736</mods:affiliation>
<mods:nameIdentifier id="9F7663C617F6DBC52FCC4BFA638ED661" type="email">smith462@msu.edu</mods:nameIdentifier>
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<mods:namePart id="76D56AFEBE74FC2A14F442726E5EA6C9">Beaver, Roger A.</mods:namePart>
<mods:affiliation id="DC97FCA9B9C0B71A9C45CEBE6CE9D3B5">161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand.</mods:affiliation>
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<mods:namePart id="DDA3C7830085511148553F13C4ACCB6D">Pham, Thai Hong</mods:namePart>
<mods:affiliation id="BF77E85BD3C4379F428BCFC0CF11D44C">Mientrung Institute for Scientific Research (MISR), Vietnam National Museum of Nature, Vietnam Academy of Science and Technology (VAST), 321 Huynh Thuc Khang St, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg., East Lansing, MI 48824, USA. cognato @ msu. edu; https: // orcid. org / 0000 - 0001 - 6436 - 2447</mods:affiliation>
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<mods:namePart id="D314C035BAA91B56D5A7439834CCBF0B">Cognato, Anthony I.</mods:namePart>
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<mods:nameIdentifier id="E984345594BEC37A6C70BE22F4DCA9B9" type="email">cognato@msu.edu</mods:nameIdentifier>
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<mods:title id="0099C4A6EC2704757F4D6E2C0D6B4D85">Zootaxa</mods:title>
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<mods:date id="C489D38F1DF970AD3AFA4A652768DA59">2022</mods:date>
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<mods:number id="88FF3E1B0C8F18E8D3511294A3837F6D">2022-11-15</mods:number>
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<treatment id="7D518786FFB47108FF7027B77F78F3AD" ID-DOI="http://doi.org/10.5281/zenodo.7322390" ID-GBIF-Taxon="205078253" ID-Zenodo-Dep="7322390" LSID="urn:lsid:plazi:treatment:7D518786FFB47108FF7027B77F78F3AD" httpUri="http://treatment.plazi.org/id/7D518786FFB47108FF7027B77F78F3AD" lastPageNumber="19" pageId="18" pageNumber="19">
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<emphasis id="C78CEA82FFB47108FF7027B77DFCF5C3" bold="true" box="[151,912,389,416]" pageId="18" pageNumber="19">
<taxonomicName id="32F84D13FFB47108FF7027B77D47F5C3" authority="Smith, Beaver &amp; Cognato" authorityName="Smith, Beaver &amp; Cognato" authorityYear="2022" box="[151,811,389,416]" class="Insecta" family="Curculionidae" genus="Xyleborinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="acanthopteron" status="sp. nov.">
<emphasis id="C78CEA82FFB47108FF7027B77F8DF5FC" bold="true" box="[151,481,389,416]" italics="true" pageId="18" pageNumber="19">Xyleborinus acanthopteron</emphasis>
Smith, Beaver &amp; Cognato
</taxonomicName>
<taxonomicNameLabel id="DCBF57F9FFB47108FCD527B77DFCF5C3" box="[818,912,389,415]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="BDE2651BFFB47108FF70279B7AE1F5BB" pageId="18" pageNumber="19" type="description">
<paragraph id="F5473690FFB47108FF70279B7F4BF59F" blockId="18.[151,1165,389,487]" box="[151,295,425,451]" pageId="18" pageNumber="19">
(
<figureCitation id="6DC32A15FFB47108FF78279B7F73F59F" box="[159,287,425,451]" captionStart="FIGURES 6172" captionStartId="20.[151,264,1228,1252]" captionTargetBox="[151,1436,562,1204]" captionTargetId="figure-113@20.[151,1436,561,1204]" captionTargetPageId="20" captionText="FIGURES 6172. Lateral, dorsal, posterior oblique and anterior view of Xyleborinus acanthopteron holotype, 2.5 mm (6164), Xyleborinus dumosus holotype, 2.9 mm (6568), Xyleborinus nobuchii holotype, 2.2 mm (6972)." figureDoi="http://doi.org/10.5281/zenodo.7322370" httpUri="https://zenodo.org/record/7322370/files/figure.png" pageId="18" pageNumber="19">Figs 6164</figureCitation>
)
</paragraph>
<paragraph id="F5473690FFB47108FF7027FF7AE1F5BB" blockId="18.[151,1165,389,487]" box="[151,1165,461,487]" pageId="18" pageNumber="19">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF70243D7C91F6CF" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="F5473690FFB47108FF70243D7AF8F633" blockId="18.[151,1437,526,2033]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF70243D7F24F674" bold="true" box="[151,328,526,553]" pageId="18" pageNumber="19">Type material.</emphasis>
<materialsCitation id="45903CCDFFB47108FEB5243D7CE1F610" ID-GBIF-Occurrence="3966310322" collectingDate="2004-10-18" collectingDateMax="2004-10-22" collectingDateMin="2004-10-18" collectionCode="MSUC" collectorName="W. Puranasakul" country="Thailand" location="Doi Pui" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Chiang Mai" typeStatus="holotype">
<typeStatus id="2A438832FFB47108FEB5243D7FD1F675" box="[338,445,527,553]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
, female:
<collectingCountry id="8DEF7600FFB47108FDCD243D7CD2F675" box="[554,702,527,553]" name="Thailand" pageId="18" pageNumber="19">THAILAND</collectingCountry>
,
<collectingRegion id="373CF872FFB47108FD2C243D7D3BF675" box="[715,855,527,553]" country="Thailand" name="Chiang Mai" pageId="18" pageNumber="19">Chiang Mai</collectingRegion>
,
<location id="F027604BFFB47108FC82243D7DADF675" LSID="urn:lsid:plazi:treatment:7D518786FFB47108FF7027B77F78F3AD:F027604BFFB47108FC82243D7DADF675" box="[869,961,527,553]" country="Thailand" name="Doi Pui" pageId="18" pageNumber="19" stateProvince="Chiang Mai">Doi Pui</location>
,
<date id="81461050FFB47108FC37243D7A00F675" box="[976,1132,527,553]" pageId="18" pageNumber="19" value="2004-10-18" valueMax="2004-10-22" valueMin="2004-10-18">
<collectingDate id="9102E9B8FFB47108FC37243D7A00F675" box="[976,1132,527,553]" pageId="18" pageNumber="19" value="2004-10-18" valueMax="2004-10-22" valueMin="2004-10-18">1822.x.2004</collectingDate>
</date>
, TH2,
<collectorName id="580D5346FFB47108FB58243D7B18F675" box="[1215,1396,527,553]" pageId="18" pageNumber="19">W. Puranasakul</collectorName>
,
<collectingMethod id="2CB94E87FFB47108FA65243D7F72F610" pageId="18" pageNumber="19">ex ethanol trap</collectingMethod>
, MSUC_ARC_314133 (
<collectionCode id="93E9AE55FFB47108FDC824007CE9F610" box="[559,645,562,588]" pageId="18" pageNumber="19">MSUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="45903CCDFFB47108FD7124007AFCF633" ID-GBIF-Occurrence="3966310465" collectingDate="2004-11" collectingDateMax="2005-01" collectingDateMin="2004-11" collectionCode="MSUC" collectorName="S. A. Dole" country="Thailand" pageId="18" pageNumber="19" specimenCount="2" stateProvince="Chiang Mai" typeStatus="paratype">
<typeStatus id="2A438832FFB47108FD7124007D6AF610" box="[662,774,562,588]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
, as
<typeStatus id="2A438832FFB47108FCC924007DFCF610" box="[814,912,562,588]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
except:
<date id="81461050FFB47108FC0A24007A28F610" box="[1005,1092,562,588]" pageId="18" pageNumber="19" value="2004-11">
<collectingDate id="9102E9B8FFB47108FC0A24007A28F610" box="[1005,1092,562,588]" pageId="18" pageNumber="19" value="2004-11">xi.2004</collectingDate>
</date>
,
<collectorName id="580D5346FFB47108FBA824007AACF610" box="[1103,1216,562,588]" pageId="18" pageNumber="19">S.A. Dole</collectorName>
, THA 149, MSUC_ ARC_314137 (1,
<collectionCode id="93E9AE55FFB47108FEB824677FDFF633" box="[351,435,597,623]" pageId="18" pageNumber="19">MSUC</collectionCode>
),
<date id="81461050FFB47108FE2124677C63F62C" box="[454,527,597,624]" pageId="18" pageNumber="19" value="2005-01">
<collectingDate id="9102E9B8FFB47108FE2124677C63F62C" box="[454,527,597,624]" pageId="18" pageNumber="19" value="2005-01">i.2005</collectingDate>
</date>
,
<collectorName id="580D5346FFB47108FDFC24677CE0F633" box="[539,652,597,623]" pageId="18" pageNumber="19">S.A. Dole</collectorName>
, THA 193, MSUC_ARC_314138 (1,
<collectionCode id="93E9AE55FFB47108FBD324677AE4F633" box="[1076,1160,597,623]" pageId="18" pageNumber="19">MSUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F5473690FFB47108FF20244A7C91F6CF" blockId="18.[151,1437,526,2033]" box="[199,765,632,659]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF20244A7F2CF6CE" bold="true" box="[199,320,632,658]" pageId="18" pageNumber="19">GenBank.</emphasis>
<materialsCitation id="45903CCDFFB47108FEA0244A7C65F6CE" ID-GBIF-Occurrence="3966310376" accessionNumber="OP607261" box="[327,521,632,658]" country="Chad" location="CAD" pageId="18" pageNumber="19" specimenCount="1">
<collectingCountry id="8DEF7600FFB47108FEA0244A7FEAF6CE" box="[327,390,632,658]" name="Chad" pageId="18" pageNumber="19">CAD</collectingCountry>
:
<accessionNumber id="EAABAB73FFB47108FE76244A7C65F6CE" box="[401,521,632,658]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OP607261" pageId="18" pageNumber="19">OP607261</accessionNumber>
</materialsCitation>
and COI:
<materialsCitation id="45903CCDFFB47108FD99244A7C96F6CF" ID-GBIF-Occurrence="3966310443" accessionNumber="OP617825" box="[638,762,632,659]" country="Chad" pageId="18" pageNumber="19" specimenCount="1">
<accessionNumber id="EAABAB73FFB47108FD99244A7C96F6CF" box="[638,762,632,659]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/OP617825" pageId="18" pageNumber="19" type="GenBank">OP617825</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF2024A97AA3F73A" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="F5473690FFB47108FF2024A97D8CF743" blockId="18.[151,1437,526,2033]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF2024A97F2EF6E9" bold="true" box="[199,322,667,693]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
This species is distinguished by the posterior margin of elytra broadly rounded; declivital face with interstriae 1 and 2 unarmed by tubercles; declivital face moderately sulcate, striae punctate; lateral declivital margins elevated, bearing 68 pairs of small to moderate, narrow, sharply pointed and recurved spines (often asymmetric) on basal 2/3 of declivity; and sulcate area of declivity shagreened, subshining.
</paragraph>
<paragraph id="F5473690FFB47108FF20251A7AA3F73A" blockId="18.[151,1437,526,2033]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF20251A7FECF71E" bold="true" box="[199,384,808,834]" pageId="18" pageNumber="19">Similar species.</emphasis>
<taxonomicName id="32F84D13FFB47108FE60251A7C1BF71E" authority="Smith &amp; Beaver &amp; Pham &amp; Cognato, 2022" authorityName="Smith &amp; Beaver &amp; Pham &amp; Cognato" authorityYear="2022" box="[391,631,808,834]" class="Insecta" family="Curculionidae" genus="Xyleborinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dumosus" status="sp. nov.">
<emphasis id="C78CEA82FFB47108FE60251A7C1BF71E" box="[391,631,808,834]" italics="true" pageId="18" pageNumber="19">Xyleborinus dumosus</emphasis>
</taxonomicName>
<emphasis id="C78CEA82FFB47108FD98251A7CB4F71E" bold="true" box="[639,728,808,834]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="DCBF57F9FFB47108FD98251A7CB4F71E" box="[639,728,808,834]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="32F84D13FFB47108FD01251A7B15F71E" authority="Smith, Beaver &amp; Cognato, 2020" authorityName="Smith, Beaver &amp; Cognato" authorityYear="2020" box="[742,1401,808,835]" class="Insecta" family="Curculionidae" genus="Xyleborinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="echinopterus">
<emphasis id="C78CEA82FFB47108FD01251A7A6AF71E" box="[742,1030,808,834]" italics="true" pageId="18" pageNumber="19">Xyleborinus echinopterus</emphasis>
<bibRefCitation id="91694B61FFB47108FBEA251A7B15F71E" author="Cognato, A. I. &amp; Smith, S. M. &amp; Beaver, R. A." box="[1037,1401,808,835]" pageId="18" pageNumber="19" pagination="540 - 554" refId="ref26083" refString="Cognato, A. I., Smith, S. M. &amp; Beaver, R. A. (2020 a) Two new genera of Oriental xyleborine ambrosia beetles (Coleoptera, Curculionidae: Scolytinae). Zootaxa, 4722 (6), 540 - 554. https: // doi. org / 10.11646 / zootaxa. 4722.6.2" type="journal article" year="2020">Smith, Beaver &amp; Cognato, 2020</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="32F84D13FFB47108FA63251B7C7EF73A" authority="(Murayama, 1931)" baseAuthorityName="Murayama" baseAuthorityYear="1931" class="Insecta" family="Curculionidae" genus="Xyleborinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="octiesdentatus">
<emphasis id="C78CEA82FFB47108FA63251B7F5BF739" italics="true" pageId="18" pageNumber="19">X. octiesdentatus</emphasis>
(
<bibRefCitation id="91694B61FFB47108FEA1257E7C66F73A" author="Murayama, J." box="[326,522,844,870]" pageId="18" pageNumber="19" pagination="39 - 62" refId="ref26999" refString="Murayama, J. (1931) Revision des familles des Ipides et Platypides (Coleopteres) de l'ile de Quelpart. Annotationes Zoologicae Japonenses, 13, 39 - 62, 2 pls." type="journal article" year="1931">Murayama, 1931</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="32F84D13FFB47108FDF9257E7D8BF73A" authority="Park &amp; Smith, 2020" authorityName="Park &amp; Smith" authorityYear="2020" box="[542,999,843,870]" class="Insecta" family="Curculionidae" genus="Xyleborinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kwangruengensis">
<emphasis id="C78CEA82FFB47108FDF9257E7C93F739" box="[542,767,843,869]" italics="true" pageId="18" pageNumber="19">X. kwangruengensis</emphasis>
<bibRefCitation id="91694B61FFB47108FCE025797D8BF73A" author="Park, S. &amp; Smith, S. M. &amp; Cognato, A. I. &amp; Beaver, R. A." box="[775,999,843,870]" pageId="18" pageNumber="19" pagination="210 - 228" refId="ref27196" refString="Park, S., Smith, S. M., Cognato, A. I. &amp; Beaver, R. A. (2020) Catalogue of Korean xyleborine ambrosia beetles (Coleoptera: Curculionidae) with seven new species. Journal of Asia-Pacific Biodiversity, 13, 210 - 228. https: // doi. org / 10.1016 / j. japb. 2020.01.002" type="journal article" year="2020">Park &amp; Smith, 2020</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="91694B61FFB47108FBF325797AABF73A" author="Park, S. &amp; Smith, S. M. &amp; Cognato, A. I. &amp; Beaver, R. A." box="[1044,1223,843,870]" pageId="18" pageNumber="19" pagination="210 - 228" refId="ref27196" refString="Park, S., Smith, S. M., Cognato, A. I. &amp; Beaver, R. A. (2020) Catalogue of Korean xyleborine ambrosia beetles (Coleoptera: Curculionidae) with seven new species. Journal of Asia-Pacific Biodiversity, 13, 210 - 228. https: // doi. org / 10.1016 / j. japb. 2020.01.002" type="journal article" year="2020">
Park
<emphasis id="C78CEA82FFB47108FBA9257E7A13F739" box="[1102,1151,843,869]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. 2020
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF20255C7BF9F33E" pageId="18" pageNumber="19" type="description">
<paragraph id="F5473690FFB47108FF20255C7BF9F33E" blockId="18.[151,1437,526,2033]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF20255C7FAEF7D4" bold="true" box="[199,450,878,904]" pageId="18" pageNumber="19">Description (female).</emphasis>
<quantity id="32009B75FFB47108FE2E255C7C73F7D5" box="[457,543,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="mm" value="2.5">2.5 mm</quantity>
long (mean =
<quantity id="32009B75FFB47108FD24255C7D70F7D5" box="[707,796,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="mm" value="2.5">2.5 mm</quantity>
; n = 3); 2.78 × as long as wide. Body, legs, and antennae uniformly brown.
<emphasis id="C78CEA82FFB47108FE8125A07FC3F7F0" bold="true" box="[358,431,914,940]" pageId="18" pageNumber="19">Head:</emphasis>
epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, alutaceous, dull, densely, coarsely punctate, setose; punctures each a long, erect hair-like seta. Eyes shallowly emarginate just above antennal insertion, upper part smaller than lower part. Submentum large, distinctly triangular, deeply impressed. Antennal scape regularly thick, as long as club. Pedicel as wide as scape, as long as funicle. Funicle 4-segmented, segment 1 shorter than pedicel. Club longer than wide, obliquely truncate,
<typeStatus id="2A438832FFB47108FAAA222D7B11F065" box="[1357,1405,1055,1081]" pageId="18" pageNumber="19">type</typeStatus>
1; segment 1 corneous, encircling anterior face; segment 2 narrow, concave, corneous on anterior face only; sutures absent on posterior face.
<emphasis id="C78CEA82FFB47108FE5022577C5CF023" bold="true" box="[439,560,1125,1151]" pageId="18" pageNumber="19">Pronotum</emphasis>
1.11 × as long as wide. In dorsal view basic and parallel-sided
<typeStatus id="2A438832FFB47108FAEA22547B51F0DC" box="[1293,1341,1126,1152]" pageId="18" pageNumber="19">type</typeStatus>
2, sides parallel in basal 2/3, rounded anteriorly; anterior margin without serrations. In lateral view elongate, disc longer than anterior slope,
<typeStatus id="2A438832FFB47108FE63229E7FD8F09A" box="[388,436,1196,1222]" pageId="18" pageNumber="19">type</typeStatus>
7, summit low, on apical 2/5. Anterior slope with densely spaced narrow asperities, becoming lower and more strongly transverse towards summit, bearing long, fine, semi-recumbent, hair-like setae. Disc subshiny, alutaceous, finely punctate, finely setose, setae short, erect, hair-like, some longer hair-like setae at margins. Lateral margins obliquely costate. Base transverse, posterior angles acutely rounded.
<emphasis id="C78CEA82FFB47108FAC323277BF0F173" bold="true" box="[1316,1436,1301,1327]" pageId="18" pageNumber="19">Scutellum</emphasis>
minute, conical, disconnected from elytra, surrounded by dense mycangial tuft of setae.
<emphasis id="C78CEA82FFB47108FB99230A7ABFF10E" bold="true" box="[1150,1235,1336,1362]" pageId="18" pageNumber="19">Elytra:</emphasis>
1.67 × as long as wide, elytra 1.50 × as long as pronotum. Base transverse, medially emarginate near scutellum and mycangial tuft, edge oblique, humeral angles angulate, parallel-sided in basal 3/4, then broadly rounded to apex. Disc occupying basal 2/3, smooth, shiny, glabrous, unarmed; striae not impressed, glabrous, with moderate punctures separated by 12 diameters of a puncture; interstriae flat, sparsely finely biseriate punctate, punctures 1/2 as large as those of striae, each bearing erect hair-like setae, setae approximately as long as width of interstriae 2. Declivital face steeply rounded, moderately sulcate between interstriae 3, shagreened, subshiny, striae and interstriae moderately setose, setae long, semi-erect hair-like, as long as the distance between suture and interstriae 3 on declivital margins and semi-recumbent and no longer than an interstrial width on declivital face; three striae present, striae 1 and 2 subcontiguous, confused; strial punctures larger, deeper than on disc; interstriae 1 and 2 flat, armed by two and three denticles, respectively on declivital base, unarmed on face; interstriae 2 very narrow; interstriae 3 forming declivital margin, distinctly elevated, 68 pairs of small to moderate, narrow sharply pointed and recurved spines (often asymmetric) on basal 2/3. Posterolateral margin rounded, denticulate from interstriae 37.
<emphasis id="C78CEA82FFB47108FB0B20ED7B41F2A5" bold="true" box="[1260,1325,1759,1785]" pageId="18" pageNumber="19">Legs:</emphasis>
procoxae contiguous, prosternal coxal piece inflated. Protibiae obliquely triangular, broadest at apical 1/3; posterior face smooth; apical 1/3 of outer margin with seven moderate socketed denticles, their length approximately as long as basal width. Meso- and metatibiae flattened, outer margin evenly rounded with eight moderate socketed denticles.
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF2021597FBEF3D9" box="[199,466,1899,1925]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="F5473690FFB47108FF2021597FBEF3D9" blockId="18.[151,1437,526,2033]" box="[199,466,1899,1925]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF2021597F0DF3D9" bold="true" box="[199,353,1899,1925]" pageId="18" pageNumber="19">Distribution.</emphasis>
<collectingCountry id="8DEF7600FFB47108FE8021597FA2F3D9" box="[359,462,1899,1925]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF2021BD7FBFF3F5" box="[199,467,1935,1961]" pageId="18" pageNumber="19" type="biology_ecology">
<paragraph id="F5473690FFB47108FF2021BD7FBFF3F5" blockId="18.[151,1437,526,2033]" box="[199,467,1935,1961]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF2021BD7F3AF3F5" bold="true" box="[199,342,1935,1961]" pageId="18" pageNumber="19">Host plants.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="BDE2651BFFB47108FF2021817F78F3AD" pageId="18" pageNumber="19" type="etymology">
<paragraph id="F5473690FFB47108FF2021817F78F3AD" blockId="18.[151,1437,526,2033]" pageId="18" pageNumber="19">
<emphasis id="C78CEA82FFB47108FF2021817F22F391" bold="true" box="[199,334,1971,1997]" pageId="18" pageNumber="19">Etymology.</emphasis>
G.
<emphasis id="C78CEA82FFB47108FE9021817FB0F391" box="[375,476,1971,1997]" italics="true" pageId="18" pageNumber="19">acanthos</emphasis>
- = thorn,
<emphasis id="C78CEA82FFB47108FDAA21817CF2F390" box="[589,670,1971,1996]" italics="true" pageId="18" pageNumber="19">-pteron</emphasis>
= wing. In reference to the spines on the elytral declivity. A noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>