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<document id="3E66D6558A872F693043FB264DAAFADB" ID-DOI="10.11646/zootaxa.3887.1.1" ID-GBIF-Dataset="9f78f2e6-ed95-4639-b784-e027d0451853" ID-ISSN="1175-5326" ID-Zenodo-Dep="287460" ID-ZooBank="A305AADA-ED80-4894-B8C4-2091F2F35544" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460973699321" checkinUser="plazi" docAuthor="Marshall, S. A., Luk, S. P. L. &amp; Dong, H." docDate="2014" docId="7D468E3E1816FFCD37DDFC72FEBEF8EB" docLanguage="en" docName="zt03887p036.pdf" docOrigin="Zootaxa 3887 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Howickia tangata Marshall, sp. nov." docType="treatment" docUuid="37DBA59E-DCBA-45D4-9836-203A2D87218E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="28" masterDocId="817FF646180DFFD6374AFFF8FFE6FF85" masterDocTitle="A revision of the New Zealand species of Howickia Richards" masterLastPageNumber="36" masterPageNumber="1" pageNumber="28" updateTime="1698559501540" updateUser="plazi">
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<mods:titleInfo id="69AF8FEFD431993DF7AE596A9759689E">
<mods:title id="1631D48F3C8925ADAA1D6A3A9989BD79">A revision of the New Zealand species of Howickia Richards</mods:title>
</mods:titleInfo>
<mods:name id="794FA353C1F6784B52A4B2EEBEFAAA52" type="personal">
<mods:role id="BE3ACEBA5CB5A653502727244AB02FDB">
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</mods:role>
<mods:namePart id="6397C535EEBA754AC225EFE1D24859C7">Marshall, S. A.</mods:namePart>
</mods:name>
<mods:name id="215F615D7D2C8C6C2D759FA833FCE0CA" type="personal">
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<mods:namePart id="06807685213B6605026C382C33195A6C">Luk, S. P. L.</mods:namePart>
</mods:name>
<mods:name id="46462829ED2F46B0FA46B7F8992C864A" type="personal">
<mods:role id="E312FEB10E6F0B8CDACE7071C1092400">
<mods:roleTerm id="5832611927382153A3343AFFBDF7156D">Author</mods:roleTerm>
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<mods:namePart id="11E5781FC42077A131C2B1F2C7A3F820">Dong, H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0CD793329FA3FA7CAE7B15D3C153A7EF">text</mods:typeOfResource>
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<mods:title id="2A376BB38F028A408D1DB031076D28E7">Zootaxa</mods:title>
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<mods:part id="486A4FE48F4913B0815704C1C398E4FD">
<mods:date id="555940A94B931BE21C3AA2B19A130BF9">2014</mods:date>
<mods:detail id="9E5D465A38AD23C522B9C9B2994C2A6D" type="volume">
<mods:number id="A08451DCC9343B1D2B616315A7BED5DF">3887</mods:number>
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<mods:identifier id="6171DE8EEF32DF9E8532FBA220A0AEBE" type="DOI">10.11646/zootaxa.3887.1.1</mods:identifier>
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<treatment id="7D468E3E1816FFCD37DDFC72FEBEF8EB" ID-DOI="http://doi.org/10.5281/zenodo.6129805" ID-GBIF-Taxon="127670818" ID-Zenodo-Dep="6129805" LSID="urn:lsid:zoobank.org:act:37DBA59E-DCBA-45D4-9836-203A2D87218E" httpUri="http://treatment.plazi.org/id/7D468E3E1816FFCD37DDFC72FEBEF8EB" lastPageNumber="28" pageId="27" pageNumber="28">
<subSubSection id="BDF56CA31816FFCD37DDFC72FE64FC43" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="F5503F281816FFCD37DDFC72FDACFC21" blockId="27.[151,586,906,966]" box="[151,586,906,932]" pageId="27" pageNumber="28">
<heading id="AE1888441816FFCD37DDFC72FDACFC21" bold="true" box="[151,586,906,932]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="C79BE33A1816FFCD37DDFC72FDACFC21" bold="true" box="[151,586,906,932]" pageId="27" pageNumber="28">
<taxonomicName id="32EF44AB1816FFCD37DDFC72FE03FC21" ID-CoL="9HNHG" authority="Marshall" authorityName="Marshall" box="[151,485,906,932]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tangata" status="sp. nov.">
<emphasis id="C79BE33A1816FFCD37DDFC72FE8BFC21" bold="true" box="[151,365,906,932]" italics="true" pageId="27" pageNumber="28">Howickia tangata</emphasis>
<collectingCountry id="8DF87FB81816FFCD3639FC72FE03FC21" box="[371,485,906,932]" name="Marshall Islands" pageId="27" pageNumber="28">Marshall</collectingCountry>
</taxonomicName>
<taxonomicNameLabel id="DCA85E411816FFCD36A6FC72FDACFC21" box="[492,586,906,932]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F5503F281816FFCD37DDFC55FE64FC43" blockId="27.[151,586,906,966]" box="[151,386,941,966]" pageId="27" pageNumber="28">
(
<figureCitation id="6DD423AD1816FFCD37D5FC55FEC0FC43" box="[159,294,941,966]" captionStart="FIGURES 15 18" captionStartId="7.[151,264,1775,1798]" captionTargetBox="[151,1436,339,1746]" captionTargetId="figure@7.[151,1436,339,1746]" captionTargetPageId="7" captionText="FIGURES 15 18. Howickia myersi group. Howickia tangata Marshall, sp. n.: 15, habitus; 16, head. Howickia myersi (Richards): 17, live individual in situ; 18, habitus." httpUri="https://zenodo.org/record/287463/files/figure.png" pageId="27" pageNumber="28">Figs. 15, 16</figureCitation>
,
<figureCitation id="6DD423AD1816FFCD3678FC55FE9CFC43" box="[306,378,941,966]" captionStart="FIGURES 66 69" captionStartId="29.[151,264,1887,1910]" captionText="FIGURES 66 69. Howickia tangata Marshall, sp. n.: 66, epandrium, posterior; 67, epandrium, left lateral; 68, male sternite 5; 69, aedeagus and associated structures." pageId="27" pageNumber="28">6669</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BDF56CA31816FFCD37DDFC0DFE7BF983" pageId="27" pageNumber="28" type="description">
<paragraph id="F5503F281816FFCD37DDFC0DFE22FBB7" blockId="27.[151,1437,1013,1902]" pageId="27" pageNumber="28">
<emphasis id="C79BE33A1816FFCD37DDFC0DFECFFB8B" bold="true" box="[151,297,1013,1038]" pageId="27" pageNumber="28">Description.</emphasis>
Head: Eye 3.0X genal height. Thorax: Scutellum 3.0X as wide as long; basal and apical marginal bristles subequal in length.
</paragraph>
<paragraph id="F5503F281816FFCD378DFBC5FB2EFAD7" blockId="27.[151,1437,1013,1902]" pageId="27" pageNumber="28">Male abdomen: Sternite 5 twice as long as sternite 4, with a pale, strongly spinulose medial patch and a straight posteromedial comb of 13 basally contiguous but distally widely spaced flat bristles. Subanal plate broad and complete with a straight ventral margin, not strongly differentiated from lower/outer part of cercus; lower parts of cerci widely separated and projecting below subanal plate. Surstylus with 3 lobes: a broad anterior lobe without large bristles; a small, pedunculate ventral lobe with a stout posteromedially curved bristle; and a large posterior lobe with several stout bristles including a very large ventral one. Distiphallus very narrow, tubular. Postgonites broad on basal part, abruptly constricted to a narrow and weakly clubbed distal part. Basiphallus frame-like, projecting posteriorly at right angle to distiphallus base for a distance equal to distiphallus base.</paragraph>
<paragraph id="F5503F281816FFCD378DFAA5FE7BF983" blockId="27.[151,1437,1013,1902]" pageId="27" pageNumber="28">Female abdomen (described from undissected abdomen with everted ovipositor): Tergite 8 divided into narrow bare dorsal sclerite and larger posteriorly setulose and setose lateral sclerites. Epiproct longitudinally divided into two sclerites, each with 1 small bristle and several small setulae; epiproct separate from the strongly sclerotized, elongate, narrow, setose cerci. Sternite 8 small and bare. Hypoproct large, mostly bare, setulose and setose only along posterior margin.</paragraph>
</subSubSection>
<subSubSection id="BDF56CA31816FFCD378DF9E9FEA8F95B" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="F5503F281816FFCD378DF9E9FEA8F95B" blockId="27.[151,1437,1013,1902]" pageId="27" pageNumber="28">
<emphasis id="C79BE33A1816FFCD378DF9E9FE93F9AF" bold="true" box="[199,373,1553,1578]" pageId="27" pageNumber="28">
<typeStatus id="2A54818A1816FFCD378DF9E9FEE4F9AF" box="[199,258,1553,1578]" pageId="27" pageNumber="28">Type</typeStatus>
material.
</emphasis>
<typeStatus id="2A54818A1816FFCD3636F9E9FE02F9AF" box="[380,484,1553,1578]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
(male,
<collectionCode id="93FEA7ED1816FFCD3573F9EAFD6EF9AF" LSID="urn:lsid:biocol.org:col:34070" box="[569,648,1554,1578]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="27" pageNumber="28">NZAC</collectionCode>
, debu00134615):
<collectingCountry id="8DF87FB81816FFCD3418F9EAFC0DF9AF" box="[850,1003,1553,1578]" name="New Zealand" pageId="27" pageNumber="28">New Zealand</collectingCountry>
, North Island, Putara Valley,
<quantity id="321792CD1816FFCD327EF9EAFA9DF9AF" box="[1332,1403,1553,1578]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="28" unit="km" value="10.0">10 km</quantity>
W Eketahuna,
<quantity id="321792CD1816FFCD366BF9CEFE96F9C8" box="[289,368,1590,1614]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="28" unit="m" value="400.0">400 m</quantity>
, pan traps,
<date id="815119E81816FFCD3548F9CDFD50F9CB" box="[514,694,1589,1614]" pageId="27" pageNumber="28" value="1999-01-14" valueMax="1999-01-16" valueMin="1999-01-14">1416.Jan.1999</date>
, S.A.
<collectingCountry id="8DF87FB81816FFCD344DF9CDFC88F9CB" box="[775,878,1589,1614]" name="Marshall Islands" pageId="27" pageNumber="28">Marshall</collectingCountry>
.
<typeStatus id="2A54818A1816FFCD3434F9CEFC16F9CB" box="[894,1008,1590,1614]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
: same as
<typeStatus id="2A54818A1816FFCD3321F9CDFB2BF9CB" box="[1131,1229,1589,1614]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(
<specimenCount id="E3E9F4A11816FFCD33AFF9CEFAD1F9CB" box="[1253,1335,1589,1614]" pageId="27" pageNumber="28" type="male">1 male</specimenCount>
,
<collectionCode id="93FEA7ED1816FFCD3203F9CEFA7EF9C8" box="[1353,1432,1590,1613]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="27" pageNumber="28">DEBU</collectionCode>
, debu00134616); same as
<typeStatus id="2A54818A1816FFCD368CF9A1FDCEF9F7" box="[454,552,1625,1650]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
but
<date id="815119E81816FFCD352CF9A1FD0BF9F7" box="[614,749,1625,1650]" pageId="27" pageNumber="28" value="1999-01-15">15.Jan.1999</date>
(
<specimenCount id="E3E9F4A11816FFCD344FF9A2FCB1F9F7" box="[773,855,1625,1650]" pageId="27" pageNumber="28" type="male">1 male</specimenCount>
,
<collectionCode id="93FEA7ED1816FFCD3423F9A2FC5CF9F7" box="[873,954,1626,1650]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="27" pageNumber="28">DEBU</collectionCode>
); North Island, Tongariro National Park, southeast corner,
<taxonomicName id="32EF44AB1816FFCD3610F985FDF8F910" box="[346,542,1661,1685]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="fusca">
<emphasis id="C79BE33A1816FFCD3610F985FDF8F910" box="[346,542,1661,1685]" italics="true" pageId="27" pageNumber="28">Nothofagus fusca</emphasis>
</taxonomicName>
bush, baited pit traps,
<date id="815119E81816FFCD3451F985FC27F913" box="[795,961,1661,1686]" pageId="27" pageNumber="28" value="1971-01-04" valueMax="1971-01-26" valueMin="1971-01-04">426.Jan.1971</date>
, H.A. Oliver (
<specimenCount id="E3E9F4A11816FFCD332AF986FB5CF913" box="[1120,1210,1661,1686]" pageId="27" pageNumber="28" type="male">4 males</specimenCount>
,
<specimenCount id="E3E9F4A11816FFCD338FF986FAD2F913" box="[1221,1332,1661,1686]" pageId="27" pageNumber="28" type="female">3 females</specimenCount>
,
<collectionCode id="93FEA7ED1816FFCD3275F986FA76F913" LSID="urn:lsid:biocol.org:col:34070" box="[1343,1424,1662,1686]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="27" pageNumber="28">NZAC</collectionCode>
); Kaimanewa Road end, NW Kaimanawa Ranges meat-baited pit traps,
<date id="815119E81816FFCD348DF95AFB63F93F" box="[967,1157,1698,1722]" pageId="27" pageNumber="28" value="1971-01-23" valueMax="1971-01-31" valueMin="1971-01-23">2331.Jan. 1971</date>
, H.A. Oliver (
<specimenCount id="E3E9F4A11816FFCD327BF95AFA66F93F" box="[1329,1408,1697,1722]" pageId="27" pageNumber="28" type="male">1 male</specimenCount>
,
<specimenCount id="E3E9F4A11816FFCD32C4F95AFF00F95B" pageId="27" pageNumber="28" type="female">1 female</specimenCount>
,
<collectionCode id="93FEA7ED1816FFCD37BBF93EFEA4F95B" LSID="urn:lsid:biocol.org:col:34070" box="[241,322,1734,1758]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="27" pageNumber="28">NZAC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="BDF56CA31816FFCD378DF911FBBCF8A3" pageId="27" pageNumber="28" type="etymology">
<paragraph id="F5503F281816FFCD378DF911FBBCF8A3" blockId="27.[151,1437,1013,1902]" pageId="27" pageNumber="28">
<emphasis id="C79BE33A1816FFCD378DF911FEA8F887" bold="true" box="[199,334,1769,1794]" pageId="27" pageNumber="28">Etymology.</emphasis>
The specific epithet is from the name for the ceremonial Maori war club called the Toki Pou
<taxonomicName id="32EF44AB1816FFCD37DDF8F6FF15F8A3" box="[151,243,1806,1830]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="34" phylum="Arthropoda" rank="species" species="tangata">Tangata</taxonomicName>
. It is applied to this species because of its elaborate and well-armed surstylus.
</paragraph>
</subSubSection>
<subSubSection id="BDF56CA31816FFCD378DF8C9FEBEF8EB" pageId="27" pageNumber="28" type="discussion">
<paragraph id="F5503F281816FFCD378DF8C9FEBEF8EB" blockId="27.[151,1437,1013,1902]" pageId="27" pageNumber="28">
<emphasis id="C79BE33A1816FFCD378DF8C9FEA9F8CF" bold="true" box="[199,335,1841,1866]" pageId="27" pageNumber="28">Comments.</emphasis>
The male genitalia of this species are unmistakable, but
<taxonomicName id="32EF44AB1816FFCD3486F8C9FB75F8CC" box="[972,1171,1841,1865]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tangata">
<emphasis id="C79BE33A1816FFCD3486F8C9FB75F8CC" box="[972,1171,1841,1865]" italics="true" pageId="27" pageNumber="28">Howickia tangata</emphasis>
</taxonomicName>
resembles
<taxonomicName id="32EF44AB1816FFCD3258F8CAFA99F8CC" box="[1298,1407,1842,1865]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="myersi">
<emphasis id="C79BE33A1816FFCD3258F8CAFA99F8CC" box="[1298,1407,1842,1865]" italics="true" pageId="27" pageNumber="28">H. myersi</emphasis>
</taxonomicName>
in external features.
</paragraph>
</subSubSection>
</treatment>
</document>