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<document id="CC5D4974C59BBEE35292D4F7BF814C7C" ID-CLB-Dataset="55816" ID-DOI="10.11646/zootaxa.5129.2.6" ID-GBIF-Dataset="5ca92eee-18fb-4a4a-8ae2-ca752e4a3498" ID-ISSN="1175-5326" ID-Zenodo-Dep="6500929" ID-ZooBank="DC6E9533-E5DD-4906-BAC5-662D5EF3ADB8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651144291995" checkinUser="plazi" docAuthor="Li, Yan-Da, Kundrata, Robin, Huang, Di-Ying &amp; Cai, Chen-Yang" docDate="2022" docId="4D2064559441FFEF1AB2FC51FECEF89F" docLanguage="en" docName="zootaxa.5192.2.6.pdf" docOrigin="Zootaxa 5129 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carinibipogonia xiai Li, Kundrata &amp; Cai 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="265" masterDocId="B1191C2D9449FFE71A25FFD7FFDAFFD6" masterDocTitle="First Artematopodidae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Elateroidea)" masterLastPageNumber="271" masterPageNumber="257" pageNumber="265" updateTime="1699329263253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C68B028EA6F6658EE3CE3B0DDAA4699C">First Artematopodidae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Elateroidea)</mods:title>
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<mods:namePart id="EE329A52971135BD273CDD4E513F644C">Li, Yan-Da</mods:namePart>
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<mods:affiliation id="C38513E5550598B19649E7D510C5F5CF">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK &amp; ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X</mods:affiliation>
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<mods:namePart id="2D2E2EFB0EC03C1AC40566F67BC53257">Kundrata, Robin</mods:namePart>
<mods:affiliation id="ECFAA4DDA321788CF7054453B3B4D4EF">Department of Zoology, Faculty of Science, Palacký University, 77900 Olomouc, Czech Republic</mods:affiliation>
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<mods:namePart id="EA2C0800F13A0173CF71911FBC062CA3">Huang, Di-Ying</mods:namePart>
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<mods:affiliation id="282C37727D3F8A522C2267FA50B74FC9">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867</mods:affiliation>
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<mods:namePart id="C970CD00682C8B45D5F14F5678D6A87D">Cai, Chen-Yang</mods:namePart>
<mods:affiliation id="94D2E1FF1D9B084EE54F5F8ED1E8F7E4">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK</mods:affiliation>
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<treatment id="4D2064559441FFEF1AB2FC51FECEF89F" ID-DOI="http://doi.org/10.5281/zenodo.6502349" ID-GBIF-Taxon="194854810" ID-Zenodo-Dep="6502349" LSID="urn:lsid:plazi:treatment:4D2064559441FFEF1AB2FC51FECEF89F" httpUri="http://treatment.plazi.org/id/4D2064559441FFEF1AB2FC51FECEF89F" lastPageNumber="265" pageId="8" pageNumber="265">
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<taxonomicName id="0289AEC09441FFEF1AB2FC51FD55FC49" authority="Li, Kundrata &amp; Cai" authorityName="Li, Kundrata &amp; Cai" authorityYear="2022" box="[151,655,901,928]" class="Insecta" family="Artematopodidae" genus="Carinibipogonia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="265" phylum="Arthropoda" rank="species" species="xiai" status="sp. nov.">
<emphasis id="F7FD09519441FFEF1AB2FC51FE54FC76" box="[151,398,902,928]" italics="true" pageId="8" pageNumber="265">Carinibipogonia xiai</emphasis>
<emphasis id="F7FD09519441FFEF1BB0FC52FD55FC49" bold="true" box="[405,655,901,928]" pageId="8" pageNumber="265">Li, Kundrata &amp; Cai</emphasis>
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<taxonomicNameLabel id="ECCEB42A9441FFEF18B3FC51FD2EFC76" box="[662,756,902,928]" pageId="8" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="8D9386C89441FFEF1AB2FC7DFEC3FC12" box="[151,281,938,964]" pageId="8" pageNumber="265" type="description">
<paragraph id="C536D5439441FFEF1AB2FC7DFEC3FC12" blockId="8.[151,756,901,964]" box="[151,281,938,964]" pageId="8" pageNumber="265">
(
<figureCitation id="5DB2C9C69441FFEF1ABAFC7DFECBFC12" box="[159,273,938,964]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="9.[151,250,1943,1967]" captionStartId-1="10.[151,250,1322,1346]" captionStartId-2="11.[151,250,1503,1527]" captionTargetBox-0="[190,1398,181,1919]" captionTargetBox-1="[151,1436,181,1298]" captionTargetBox-2="[151,1436,181,1480]" captionTargetId-0="figure-18@9.[190,1398,181,1919]" captionTargetId-1="figure-22@10.[151,1436,181,1298]" captionTargetId-2="figure-18@11.[151,1436,181,1480]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 8. General habitus of Carinibipogonia xiai Li, Kundrata &amp; Cai sp. nov., holotype, NIGP179432, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 1.5 mm." captionText-1="FIGURE 9. Details of Carinibipogonia xiai Li, Kundrata &amp; Cai gen. et sp. nov., holotype, NIGP179432, under widefield fluorescence. A, Head and prothorax, dorsal view. B, Head and prothorax, ventral view. C, Hind wing, dorsal view. D, Abdomen, ventral view. E, Elytron, ventral view.Abbreviations: an, antenna; el, elytron; lbp, labial palp; md, mandible; mt, mentum; mxp, maxillary palp; pn, pronotum; ps, prosternum; ptb, protibia; R, radial cell; v15, ventrites 15. Scale bars: 500 μm." captionText-2="FIGURE 10. Details of Carinibipogonia xiai Li, Kundrata &amp; Cai gen. et sp. nov., holotype, NIGP179432, under confocal microscopy. A, Head, ventral view. B, Prothorax, ventral view, showing the paired longitudinal ridges (arrowhead). C, Fore and mid legs. D, Elytral apex, showing the interlocking tongue (arrowhead). E, Abdominal apex, ventral view. F, Head, dorsal view. G, Antenna, dorsal view. H, Elytron, dorsal view. I, Ovipositor, dorsal view. Abbreviations: a111, antennomeres 111; cl, clypeus; gs, gonostylus; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; mt, mentum; ps, prosternum; ptb, protibia; pts, protarsus; v5, ventrite 5. Scale bars: 300 μm." figureDoi-0="http://doi.org/10.5281/zenodo.6500948" figureDoi-1="http://doi.org/10.5281/zenodo.6500950" figureDoi-2="http://doi.org/10.5281/zenodo.6500952" httpUri-0="https://zenodo.org/record/6500948/files/figure.png" httpUri-1="https://zenodo.org/record/6500950/files/figure.png" httpUri-2="https://zenodo.org/record/6500952/files/figure.png" pageId="8" pageNumber="265">Figs 810</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8D9386C89441FFEF1AB2FC26FD0CFBDA" box="[151,726,1009,1036]" pageId="8" pageNumber="265" type="materials_examined">
<paragraph id="C536D5439441FFEF1AB2FC26FD0CFBDA" blockId="8.[151,1437,1009,1865]" box="[151,726,1009,1036]" pageId="8" pageNumber="265">
<emphasis id="F7FD09519441FFEF1AB2FC26FEDCFBDD" bold="true" box="[151,262,1009,1035]" pageId="8" pageNumber="265">Material.</emphasis>
<materialsCitation id="75E1DF1E9441FFEF1B28FC25FD0CFBDA" ID-GBIF-Occurrence="3757619302" box="[269,726,1010,1036]" collectionCode="NIGP" pageId="8" pageNumber="265" specimenCode="NIGP179432" specimenCount="1" typeStatus="holotype">
<typeStatus id="1A326BE19441FFEF1B28FC25FEA2FBDA" box="[269,376,1010,1036]" pageId="8" pageNumber="265" type="holotype">Holotype</typeStatus>
, female,
<specimenCode id="952F7D389441FFEF1BFBFC25FDA9FBDA" box="[478,627,1010,1036]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="8" pageNumber="265">NIGP179432</specimenCode>
(
<collectionCode id="A3984D869441FFEF18A7FC25FD10FBDA" box="[642,714,1010,1036]" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="8" pageNumber="265">NIGP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8D9386C89441FFEF1AE2FBC1FDCBFB82" pageId="8" pageNumber="265" type="etymology">
<paragraph id="C536D5439441FFEF1AE2FBC1FDCBFB82" blockId="8.[151,1437,1009,1865]" pageId="8" pageNumber="265">
<emphasis id="F7FD09519441FFEF1AE2FBC1FE94FBE6" bold="true" box="[199,334,1046,1072]" pageId="8" pageNumber="265">Etymology.</emphasis>
The species is named after Mr. Fang-Yuan Xia, who kindly allowed us to study many valuable fossil specimens in his collection.
</paragraph>
</subSubSection>
<subSubSection id="8D9386C89441FFEF1AE2FB8AFB38FB4A" pageId="8" pageNumber="265" type="distribution">
<paragraph id="C536D5439441FFEF1AE2FB8AFB38FB4A" blockId="8.[151,1437,1009,1865]" pageId="8" pageNumber="265">
<emphasis id="F7FD09519441FFEF1AE2FB8AFE15FBA1" bold="true" box="[199,463,1117,1144]" pageId="8" pageNumber="265">Locality and horizon.</emphasis>
Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="074D1BA19441FFEF1AB2FB55FEF0FB4A" box="[151,298,1154,1180]" country="Myanmar" name="Kachin" pageId="8" pageNumber="265">Kachin State</collectingRegion>
,
<collectingCountry id="BD9E95D39441FFEF1B10FB55FE7CFB4A" box="[309,422,1154,1180]" name="Myanmar" pageId="8" pageNumber="265">Myanmar</collectingCountry>
; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.
</paragraph>
</subSubSection>
<subSubSection id="8D9386C89441FFEF1AE2FB72FDDDFB16" box="[199,519,1189,1216]" pageId="8" pageNumber="265" type="diagnosis">
<paragraph id="C536D5439441FFEF1AE2FB72FDDDFB16" blockId="8.[151,1437,1009,1865]" box="[199,519,1189,1216]" pageId="8" pageNumber="265">
<emphasis id="F7FD09519441FFEF1AE2FB72FE98FB69" bold="true" box="[199,322,1189,1215]" pageId="8" pageNumber="265">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="8D9386C89441FFEF1AE2FB1EFECEF89F" pageId="8" pageNumber="265" type="description">
<paragraph id="C536D5439441FFEF1AE2FB1EFBA9FB32" blockId="8.[151,1437,1009,1865]" box="[199,1139,1225,1252]" pageId="8" pageNumber="265">
<emphasis id="F7FD09519441FFEF1AE2FB1EFE83FB35" bold="true" box="[199,345,1225,1251]" pageId="8" pageNumber="265">Description.</emphasis>
Body moderately broad,
<quantity id="027178A69441FFEF1853FB1DFD16FB32" box="[630,716,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="8" pageNumber="265" unit="mm" value="4.4">4.4 mm</quantity>
long,
<quantity id="027178A69441FFEF1937FB1DFCB2FB32" box="[786,872,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="265" unit="mm" value="1.7">1.7 mm</quantity>
wide; surface punctate.
</paragraph>
<paragraph id="C536D5439441FFEF1AE2FB39FEC1FA6A" blockId="8.[151,1437,1009,1865]" pageId="8" pageNumber="265">
Eyes well-developed, without interfacetal setae. Antennal insertions dorsally exposed. Subantennal groove absent. Frontoclypeal suture absent; anterior clypeal margin multidentate. Antennomeres 24 simple; antennomere 5 moderately serrate; antennomeres 610 strongly serrate, about as long as wide. Mandibles with one apical and one subapical teeth. Maxillary palps 4-segmented; apical palpomere seemingly cylindrical, not clearly expanded apically. Labial palps 3-segmented; apical palpomere widened apically.
<collectingRegion id="074D1BA19441FFEF19FDFAA9FBEDFA4E" box="[984,1079,1406,1432]" country="Myanmar" name="Chin" pageId="8" pageNumber="265">Mentum</collectingRegion>
subtriangular, apically slightly emarginate.
</paragraph>
<paragraph id="C536D5439441FFEF1AE2FA11FB90F9FE" blockId="8.[151,1437,1009,1865]" pageId="8" pageNumber="265">Pronotal disc subtrapezoidal, widest at base; surface without transverse groove. Prosternum in front coxae transverse, with paired incomplete ridges; ridges not extending to anterior half of prosternum; deep pits absent; prosternal process with subparallel sides. Procoxal cavities broadly open posteriorly.</paragraph>
<paragraph id="C536D5439441FFEF1AE2F9E5FAB0F99A" blockId="8.[151,1437,1009,1865]" box="[199,1386,1586,1612]" pageId="8" pageNumber="265">Elytra elongate; surface of each elytron with 10 puncture rows; epipleura developed only in anterior third.</paragraph>
<paragraph id="C536D5439441FFEF1AE2F981FACAF942" blockId="8.[151,1437,1009,1865]" pageId="8" pageNumber="265">Legs slender; tibiae slightly longer than respective femora, with two tibial spurs; tarsomeres 1 and 5 elongate; tarsomeres 3 and 4 with well-developed, bilobed membranous ventral process. Pretarsal claws simple.</paragraph>
<paragraph id="C536D5439441FFEF1AE2F949FEFFF90A" blockId="8.[151,1437,1009,1865]" pageId="8" pageNumber="265">Hind wing well-developed. Radial cell elongate, about 4.2 times as long as wide; inner posterobasal angle almost right.</paragraph>
<paragraph id="C536D5439441FFEF1AE2F931FECEF89F" blockId="8.[151,1437,1009,1865]" pageId="8" pageNumber="265">Abdominal ventrites with margins denticulate. Sutures between all ventrites complete and distinct; suture between ventrites 4 and 5 only weakly curved. Ventrite 5 about 1.3 times as long as ventrite 4 (might be affected by distortion).</paragraph>
</subSubSection>
</treatment>
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