279 lines
33 KiB
XML
279 lines
33 KiB
XML
<document id="3F51E911F592B575FDE41C73834CC125" ID-CLB-Dataset="28086" ID-DOI="10.11646/zootaxa.4504.2.3" ID-GBIF-Dataset="c4c5a6db-ede2-4797-a9bb-fd7f7df6fed3" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606266" ID-ZooBank="DA9CF61F-0398-4E71-8CE5-3A77FEB00CB0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553581558909" checkinUser="plazi" docAuthor="Enushchenko, Ilya V." docDate="2018" docId="4411879EFFF4B03EFF76FB6B21F9FB64" docLanguage="en" docName="zootaxa.4504.2.3.pdf" docOrigin="Zootaxa 4504 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gyrophaena fontanedai Enushchenko 2018, sp.n." docType="treatment" docVersion="7" lastPageNumber="214" masterDocId="B828FFE6FFF7B03BFFE1FFA42073FFC4" masterDocTitle="Five new species of the tribe Gyrophaenina Kraatz 1830 (Coleoptera: Staphylinidae: Aleocharinae) from the southern part of the United States" masterLastPageNumber="224" masterPageNumber="209" pageNumber="212" updateTime="1698712732609" updateUser="ExternalLinkService">
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<mods:title id="F2F21DAC23BEF757FEE2E5C574B1D617">Five new species of the tribe Gyrophaenina Kraatz 1830 (Coleoptera: Staphylinidae: Aleocharinae) from the southern part of the United States</mods:title>
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<mods:name id="187EAAF708548A01906DA4D7710657D2" type="personal">
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<mods:namePart id="CED3154B5BFB7135064D8A53AE825192">Enushchenko, Ilya V.</mods:namePart>
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<mods:title id="2A077AF48177620A6C1A8C581D718570">Zootaxa</mods:title>
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<mods:part id="950B31B9DC37183B71682C7E2D801838">
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<mods:date id="2E5E5606A6315F3668439685F12F7E2E">2018</mods:date>
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<mods:number id="354D582EB22B79210BE3893F6D2BD4D7">2018-10-24</mods:number>
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<mods:number id="1C5347DDCCC1666CE90041C84CE53EC1">4504</mods:number>
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<mods:identifier id="BC1A4C69B3DC96B4D460B79E93B4F3C5" type="DOI">10.11646/zootaxa.4504.2.3</mods:identifier>
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<treatment id="4411879EFFF4B03EFF76FB6B21F9FB64" ID-DOI="http://doi.org/10.5281/zenodo.5987759" ID-GBIF-Taxon="154494951" ID-Zenodo-Dep="5987759" LSID="urn:lsid:plazi:treatment:4411879EFFF4B03EFF76FB6B21F9FB64" httpUri="http://treatment.plazi.org/id/4411879EFFF4B03EFF76FB6B21F9FB64" lastPageId="5" lastPageNumber="214" pageId="3" pageNumber="212">
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<subSubSection id="84A26503FFF4B038FF76FB6B237FFB2E" box="[151,780,1231,1258]" pageId="3" pageNumber="212" type="nomenclature">
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<paragraph id="CC073688FFF4B038FF76FB6B237FFB2E" blockId="3.[151,780,1231,1291]" box="[151,780,1231,1258]" pageId="3" pageNumber="212">
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<heading id="974F81E4FFF4B038FF76FB6B237FFB2E" bold="true" box="[151,780,1231,1258]" fontSize="11" level="1" pageId="3" pageNumber="212" reason="1">
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<emphasis id="FECCEA9AFFF4B038FF76FB6B237FFB2E" bold="true" box="[151,780,1231,1258]" pageId="3" pageNumber="212">
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<taxonomicName id="0BB84D0BFFF4B038FF76FB6B22B7FB2E" ID-CoL="3HZYX" authority="Enushchenko" authorityName="Enushchenko" authorityYear="2018" box="[151,708,1231,1258]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="fontanedai" status="sp. nov.">
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<emphasis id="FECCEA9AFFF4B038FF76FB6B215EFB2D" bold="true" box="[151,301,1231,1257]" italics="true" pageId="3" pageNumber="212">Gyrophaena</emphasis>
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(s. str.)
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<emphasis id="FECCEA9AFFF4B038FE6EFB6B2262FB2D" bold="true" box="[399,529,1231,1257]" italics="true" pageId="3" pageNumber="212">fontanedai</emphasis>
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Enushchenko
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</taxonomicName>
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,
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<taxonomicNameLabel id="E5FF57E1FFF4B038FD30FB74237FFB2E" box="[721,780,1232,1258]" pageId="3" pageNumber="212" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="84A26503FFF4B038FF76FB562153FACF" box="[151,288,1266,1291]" pageId="3" pageNumber="212" type="description">
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<paragraph id="CC073688FFF4B038FF76FB562153FACF" blockId="3.[151,780,1231,1291]" box="[151,288,1266,1291]" pageId="3" pageNumber="212">
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(
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<figureCitation id="54832A0DFFF4B038FF7EFB562164FACF" box="[159,279,1266,1291]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1929,1952]" captionTargetBox="[186,1376,203,1903]" captionTargetId="figure@4.[180,1414,193,1919]" captionTargetPageId="4" captionText="FIGURES 6–10. Gyrophaena fontanedai sp.n.: 6—habitus, 7—aedeagus, lateral view (Holotype), 8—aedeagus, lateral view (Paratype), 9—male abdominal tergite VIII, dorsal view (Holotype), 10—male abdominal tergite VIII, dorsal view (Paratype). Scale bars: 1 mm (6), 0.1 mm (7–10)." httpUri="https://zenodo.org/record/2946795/files/figure.png" pageId="3" pageNumber="212">Figs. 6–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="84A26503FFF4B038FF76FA9E25FFF8C7" pageId="3" pageNumber="212" type="materials_examined">
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<paragraph id="CC073688FFF4B038FF76FA9E23F2FA27" blockId="3.[151,1436,1338,2011]" pageId="3" pageNumber="212">
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<emphasis id="FECCEA9AFFF4B038FF76FA9E21CBFA97" bold="true" box="[151,440,1338,1363]" pageId="3" pageNumber="212">Type material examined</emphasis>
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:
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<typeStatus id="1303882AFFF4B038FE26FA9E2247FA97" box="[455,564,1338,1363]" pageId="3" pageNumber="212" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="DABEFD01FFF4B038FDDEFA9F221AFA96" box="[575,617,1339,1363]" pageId="3" pageNumber="212" type="male">1 ♂</specimenCount>
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[Specimen from was re-glued by me on white rectangular card; a plastic plate with a preparation of the aedeagus and abdominal segments VIII in Canada balsam was pinned under the card with specimen]:
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<collectingCountry id="B4AF7618FFF4B038FEB4FA2621FEFA5F" box="[341,397,1410,1435]" name="United States of America" pageId="3" pageNumber="212">
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<emphasis id="FECCEA9AFFF4B038FEB4FA2621FEFA5F" bold="true" box="[341,397,1410,1435]" pageId="3" pageNumber="212">USA</emphasis>
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</collectingCountry>
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: ‘ALA[bama]: Shalby Co[unty]; | Helena 684B |
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<date id="B8061048FFF4B038FC27FA26245BFA5F" box="[966,1064,1410,1435]" pageId="3" pageNumber="212" value="1959-01-03">I-3-1959</date>
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| leg. H.R. Steeves Jr.’, ‘CNHM 1962 | H.G. Steeves| Gen. Coleop. Colln. | Acc. Z–13, 050’ ‘Tree Hole’, ‘
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<typeStatus id="1303882AFFF4B038FC34FA03241DFA7B" box="[981,1134,1447,1471]" pageId="3" pageNumber="212" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="0BB84D0BFFF4B038FB62FA02257DFA7A" authorityName="Mannerheim" authorityYear="1830" box="[1155,1294,1446,1470]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="genus">
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<emphasis id="FECCEA9AFFF4B038FB62FA02257DFA7A" box="[1155,1294,1446,1470]" italics="true" pageId="3" pageNumber="212">Gyrophaena</emphasis>
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</taxonomicName>
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<taxonomicName id="0BB84D0BFFF4B038FAC5FA0225EFFA7A" authority="Enushchenko, 2018" authorityName="Enushchenko" authorityYear="2018" box="[1316,1436,1446,1470]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="fontanedai" status="sp. nov.">
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<emphasis id="FECCEA9AFFF4B038FAC5FA0225EFFA7A" box="[1316,1436,1446,1470]" italics="true" pageId="3" pageNumber="212">fontanedai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E5FF57E1FFF4B038FF76FA6E20BDFA27" box="[151,206,1482,1507]" pageId="3" pageNumber="212" rank="species">
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<emphasis id="FECCEA9AFFF4B038FF76FA6E20BDFA27" bold="true" box="[151,206,1482,1507]" pageId="3" pageNumber="212">sp.n.</emphasis>
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</taxonomicNameLabel>
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| Enushchenko I.V. | 2018 des.’ <red printed label> (FMNH).
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</paragraph>
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<paragraph id="CC073688FFF4B038FF26FA4B25FFF8C7" blockId="3.[151,1436,1338,2011]" pageId="3" pageNumber="212">
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<typeStatus id="1303882AFFF4B038FF26FA4B214AF9C3" box="[199,313,1519,1543]" pageId="3" pageNumber="212" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="DABEFD01FFF4B038FEA3FA4B2118F9C2" box="[322,363,1519,1543]" pageId="3" pageNumber="212" type="male">1 ♂</specimenCount>
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||
[Specimen from was re-glued by me on white rectangular card; a plastic card with a preparation of the abdominal tergite VIII in
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<collectingCountry id="B4AF7618FFF4B038FDF8F9B6221EF9EF" box="[537,621,1554,1579]" name="Canada" pageId="3" pageNumber="212">Canada</collectingCountry>
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balsam was pinned under the card with specimen; aedeagus is lost]: ‘
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<collectingCountry id="B4AF7618FFF4B038FF7DF993209EF98B" box="[156,237,1591,1615]" name="United States of America" pageId="3" pageNumber="212">U.S.A.</collectingCountry>
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<collectingRegion id="0E7CF86AFFF4B038FF15F9922136F98B" box="[244,325,1590,1615]" country="United States of America" name="Florida" pageId="3" pageNumber="212">Florida</collectingRegion>
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| Monroe Co[unty].; Lignum | Vitae Key. 14:III:1 968 | leg. S. Peck, Ber.: #109’, ‘FM(HD) #68-15, | Ber.: hardwood | forest litter.
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<quantity id="0B409B6DFFF4B038FE1BF9FF223AF9B7" box="[506,585,1626,1651]" metricMagnitude="1" metricUnit="kg" metricValue="1.270058636" pageId="3" pageNumber="212" unit="lb" value="28.0">28 lbs</quantity>
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.’, ‘
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<typeStatus id="1303882AFFF4B038FD84F9FF2286F9B7" box="[613,757,1627,1651]" pageId="3" pageNumber="212" type="paratype">PARATYPE</typeStatus>
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||
<taxonomicName id="0BB84D0BFFF4B038FCEFF9FF23EAF9B7" authorityName="Mannerheim" authorityYear="1830" box="[782,921,1627,1651]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="genus">
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<emphasis id="FECCEA9AFFF4B038FCEFF9FF23EAF9B7" box="[782,921,1627,1651]" italics="true" pageId="3" pageNumber="212">Gyrophaena</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="0BB84D0BFFF4B038FC50F9FF245BF9B7" authority="Enushchenko, 2018" authorityName="Enushchenko" authorityYear="2018" box="[945,1064,1627,1651]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="fontanedai" status="sp. nov.">
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<emphasis id="FECCEA9AFFF4B038FC50F9FF245BF9B7" box="[945,1064,1627,1651]" italics="true" pageId="3" pageNumber="212">fontanedai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E5FF57E1FFF4B038FBD2F9FE2419F9B7" box="[1075,1130,1626,1651]" pageId="3" pageNumber="212" rank="species">
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<emphasis id="FECCEA9AFFF4B038FBD2F9FE2419F9B7" bold="true" box="[1075,1130,1626,1651]" pageId="3" pageNumber="212">sp.n.</emphasis>
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</taxonomicNameLabel>
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| Enushchenko I.V. | 2018 des.’ <red printed label> (FMNH);
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<specimenCount id="DABEFD01FFF4B038FDD1F9DB222FF952" box="[560,604,1663,1687]" pageId="3" pageNumber="212" type="male">1 ♂</specimenCount>
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[Specimen was cleaned and reglued by me on white rectangular card; a plastic card with a preparation of the abdominal tergite VIII and aedeagus in
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<collectingCountry id="B4AF7618FFF4B038FC1BF906243DF97F" box="[1018,1102,1698,1723]" name="Canada" pageId="3" pageNumber="212">Canada</collectingCountry>
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balsam was pinned under the card with specimen]: ‘
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<taxonomicName id="0BB84D0BFFF4B038FE43F962225EF91A" authorityName="Mannerheim" authorityYear="1830" box="[418,557,1734,1758]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="genus">
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<emphasis id="FECCEA9AFFF4B038FE43F962225EF91A" box="[418,557,1734,1758]" italics="true" pageId="3" pageNumber="212">Gyrophaena</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="0BB84D0BFFF4B038FDAAF96222BEF91A" box="[587,717,1734,1758]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="breviuscula">
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<emphasis id="FECCEA9AFFF4B038FDAAF96222BEF91A" box="[587,717,1734,1758]" italics="true" pageId="3" pageNumber="212">breviuscula</emphasis>
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</taxonomicName>
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| Motch.[ulskiy] | Am.[erica] bor.[ealis]’ <green handwritten label>, ‘
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<typeStatus id="1303882AFFF4B038FF0EF94F21F3F8C7" box="[239,384,1771,1795]" pageId="3" pageNumber="212" type="paratype">PARATYPE</typeStatus>
|
||
|
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||
<taxonomicName id="0BB84D0BFFF4B038FE72F94F226DF8C7" authorityName="Mannerheim" authorityYear="1830" box="[403,542,1771,1795]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="genus">
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<emphasis id="FECCEA9AFFF4B038FE72F94F226DF8C7" box="[403,542,1771,1795]" italics="true" pageId="3" pageNumber="212">Gyrophaena</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="0BB84D0BFFF4B038FDD0F94F22DBF8C7" authority="Enushchenko, 2018" authorityName="Enushchenko" authorityYear="2018" box="[561,680,1771,1795]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="fontanedai" status="sp. nov.">
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<emphasis id="FECCEA9AFFF4B038FDD0F94F22DBF8C7" box="[561,680,1771,1795]" italics="true" pageId="3" pageNumber="212">fontanedai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5FF57E1FFF4B038FD51F94E229BF8C7" box="[688,744,1770,1795]" pageId="3" pageNumber="212" rank="species">
|
||
<emphasis id="FECCEA9AFFF4B038FD51F94E229BF8C7" bold="true" box="[688,744,1770,1795]" pageId="3" pageNumber="212">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
| Enushchenko I.V. | 2018 des.’ <red printed label> (ZMM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="84A26503FFF4B03EFF26F8AA24CBFD34" lastPageId="5" lastPageNumber="214" pageId="3" pageNumber="212" type="description">
|
||
<paragraph id="CC073688FFF4B03EFF26F8AA2400FF3C" blockId="3.[151,1436,1338,2011]" lastBlockId="5.[151,1437,151,1184]" lastPageId="5" lastPageNumber="214" pageId="3" pageNumber="212">
|
||
<emphasis id="FECCEA9AFFF4B038FF26F8AA2122F8E3" bold="true" box="[199,337,1806,1831]" pageId="3" pageNumber="212">Description</emphasis>
|
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. Body red-yellow to red-brown; length
|
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<quantity id="0B409B6DFFF4B038FCF4F8AA230EF8E2" box="[789,893,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="212" unit="mm" value="1.5">1.50 mm</quantity>
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. Habitus as in
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<figureCitation id="54832A0DFFF4B038FBC2F8AA241BF8E3" box="[1059,1128,1806,1831]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1929,1952]" captionTargetBox="[186,1376,203,1903]" captionTargetId="figure@4.[180,1414,193,1919]" captionTargetPageId="4" captionText="FIGURES 6–10. Gyrophaena fontanedai sp.n.: 6—habitus, 7—aedeagus, lateral view (Holotype), 8—aedeagus, lateral view (Paratype), 9—male abdominal tergite VIII, dorsal view (Holotype), 10—male abdominal tergite VIII, dorsal view (Paratype). Scale bars: 1 mm (6), 0.1 mm (7–10)." httpUri="https://zenodo.org/record/2946795/files/figure.png" pageId="3" pageNumber="212">Fig. 6</figureCitation>
|
||
. Antennae, mouthparts and legs yellow-red or red-brown. Body with distinct reticulate microsculpture, weaker on abdomen. Head almost twice as wide as long; vertex with small, sparse punctures; median area impunctate. Antennomeres IV and V as wide as long, quadrate to slightly elongate, antennomeres VI–X subquadrate to feebly transverse, apical antennomere flattened, conical, 1.5 times longer than wide. Pronotum 1.6 times as wide as long and 1.4 times as wide as head; microsculpture as that on head, with two median rows consisting of three small, shallow, weakly defined punctures; basal portion of pronotum impunctate. Elytra approximately as long as pronotum, almost twice as wide as long; with distinct reticulate microsculpture, surface covered by indistinct, small punctures and welldefined tubercles, most densely in the middle part and in the posterior margins of elytra.
|
||
</paragraph>
|
||
<caption id="98C76600FFF3B03FFF76F82D2267F820" httpUri="https://zenodo.org/record/2946795/files/figure.png" pageId="4" pageNumber="213" startId="4.[151,264,1929,1952]" targetBox="[186,1376,203,1903]" targetPageId="4">
|
||
<paragraph id="CC073688FFF3B03FFF76F82D2267F820" blockId="4.[151,1436,1929,2020]" pageId="4" pageNumber="213">
|
||
<emphasis id="FECCEA9AFFF3B03FFF76F82D2132F85B" bold="true" box="[151,321,1929,1952]" pageId="4" pageNumber="213">FIGURES 6–10</emphasis>
|
||
.
|
||
<taxonomicName id="0BB84D0BFFF3B03FFEAEF82E2233F864" authority="Enushchenko, 2018" authorityName="Enushchenko" authorityYear="2018" box="[335,576,1930,1952]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="213" phylum="Arthropoda" rank="genus" status="sp. nov.">
|
||
<emphasis id="FECCEA9AFFF3B03FFEAEF82E2233F864" box="[335,576,1930,1952]" italics="true" pageId="4" pageNumber="213">Gyrophaena fontanedai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5FF57E1FFF3B03FFDA9F82E2209F85B" box="[584,634,1930,1951]" pageId="4" pageNumber="213" rank="species">
|
||
<emphasis id="FECCEA9AFFF3B03FFDA9F82E2209F85B" bold="true" box="[584,634,1930,1951]" pageId="4" pageNumber="213">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 6—habitus, 7—aedeagus, lateral view (Holotype), 8—aedeagus, lateral view (Paratype), 9—male abdominal tergite VIII, dorsal view (Holotype), 10—male abdominal tergite VIII, dorsal view (Paratype). Scale bars: 1 mm (6), 0.1 mm (7–10).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CC073688FFF2B03EFF26FEA02480FE43" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="214">
|
||
Male. Apical margin of abdominal tergite VIII with four teeth, with lateral pair rather stout and slightly curved inwards and medial pair blunt, equal to margin carinae (
|
||
<figureCitation id="54832A0DFFF2B03EFCC9FE83231CFE84" box="[808,879,295,320]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1929,1952]" captionTargetBox="[186,1376,203,1903]" captionTargetId="figure@4.[180,1414,193,1919]" captionTargetPageId="4" captionText="FIGURES 6–10. Gyrophaena fontanedai sp.n.: 6—habitus, 7—aedeagus, lateral view (Holotype), 8—aedeagus, lateral view (Paratype), 9—male abdominal tergite VIII, dorsal view (Holotype), 10—male abdominal tergite VIII, dorsal view (Paratype). Scale bars: 1 mm (6), 0.1 mm (7–10)." httpUri="https://zenodo.org/record/2946795/files/figure.png" pageId="5" pageNumber="214">Fig. 9</figureCitation>
|
||
). Ventral plate of aedeagus long, widely curved latero-ventrally, with acute apex (
|
||
<figureCitation id="54832A0DFFF2B03EFDF9FEE8222DFEA0" box="[536,606,332,357]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1929,1952]" captionTargetBox="[186,1376,203,1903]" captionTargetId="figure@4.[180,1414,193,1919]" captionTargetPageId="4" captionText="FIGURES 6–10. Gyrophaena fontanedai sp.n.: 6—habitus, 7—aedeagus, lateral view (Holotype), 8—aedeagus, lateral view (Paratype), 9—male abdominal tergite VIII, dorsal view (Holotype), 10—male abdominal tergite VIII, dorsal view (Paratype). Scale bars: 1 mm (6), 0.1 mm (7–10)." httpUri="https://zenodo.org/record/2946795/files/figure.png" pageId="5" pageNumber="214">Fig. 7</figureCitation>
|
||
); apical part of median lobe very short; dorsal projection of internal sac long but distinctly shorter than ventral plate, inclined to the ventral plate, slightly widening to apex.
|
||
</paragraph>
|
||
<paragraph id="CC073688FFF2B03EFF26FE3021E3FE69" blockId="5.[151,1437,151,1184]" box="[199,400,404,429]" pageId="5" pageNumber="214">Female unknown.</paragraph>
|
||
<paragraph id="CC073688FFF2B03EFF26FE132217FD09" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="214">
|
||
<emphasis id="FECCEA9AFFF2B03EFF26FE1321D8FE14" bold="true" box="[199,427,439,464]" pageId="5" pageNumber="214">Comparative notes</emphasis>
|
||
. Based on the morphological features both of habitus and aedeagus,
|
||
<taxonomicName id="0BB84D0BFFF2B03EFB22FE1C2528FE14" authority="Enushchenko, 2018" authorityName="Enushchenko" authorityYear="2018" box="[1219,1371,440,464]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="fontanedai" status="sp. nov.">
|
||
<emphasis id="FECCEA9AFFF2B03EFB22FE1C2528FE14" box="[1219,1371,440,464]" italics="true" pageId="5" pageNumber="214">G. fontanedai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5FF57E1FFF2B03EFA85FE1325E8FE14" box="[1380,1435,439,464]" pageId="5" pageNumber="214" rank="species">
|
||
<emphasis id="FECCEA9AFFF2B03EFA85FE1325E8FE14" bold="true" box="[1380,1435,439,464]" pageId="5" pageNumber="214">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from all known Nearctic species and is more similar to the Euro-asiatic
|
||
<taxonomicName id="0BB84D0BFFF2B03EFBC3FE782517FE30" authority="(Gyllenhal 1810)" authorityName="Gyllenhal" authorityYear="1810" baseAuthorityName="Gyllenhal" baseAuthorityYear="1810" box="[1058,1380,476,501]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="nitidula">
|
||
<emphasis id="FECCEA9AFFF2B03EFBC3FE7824EAFE30" box="[1058,1177,476,500]" italics="true" pageId="5" pageNumber="214">G. nitidula</emphasis>
|
||
(Gyllenhal 1810)
|
||
</taxonomicName>
|
||
and, especially by the distinct reticulate microsculpture of the body, shapes of antennomeres IV–IX and shape of the aedeagus to
|
||
<taxonomicName id="0BB84D0BFFF2B03EFEC0FD8022E4FDF9" authority="Semenov 2015" authorityName="Semenov" authorityYear="2015" box="[289,663,548,573]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="pseudonitidula">
|
||
<emphasis id="FECCEA9AFFF2B03EFEC0FD802195FDF8" box="[289,486,548,572]" italics="true" pageId="5" pageNumber="214">G. pseudonitidula</emphasis>
|
||
<bibRefCitation id="A8294B79FFF2B03EFE0DFD8022E4FDF9" author="Semenov, V. B. & Egorov, L. V. & Vinogradova, E. Yu." box="[492,663,548,573]" pageId="5" pageNumber="214" refId="ref8070" refString="Semenov, V. B., Egorov, L. V. & Vinogradova, E. Yu. (2015) Annotirovannyi spisok stafilinid (Insecta, Coleoptera, Staphylinidae) Chuvashskoi respubliki [The staphylinid beetles annotated checklist of the Chuvash Republic]. Novoye vremya, Cheboksary, 146 pp. [in Russian]" type="book" year="2015">Semenov 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
, described recently from the Chuvash Republic of
|
||
<collectingCountry id="B4AF7618FFF2B03EFB2EFD802569FDF9" box="[1231,1306,548,573]" name="Russia" pageId="5" pageNumber="214">Russia</collectingCountry>
|
||
and known from a vast range extending from the European part of
|
||
<collectingCountry id="B4AF7618FFF2B03EFD1AFDE32335FDA4" box="[763,838,583,608]" name="Russia" pageId="5" pageNumber="214">Russia</collectingCountry>
|
||
to the Russian Far East (
|
||
<bibRefCitation id="A8294B79FFF2B03EFBBAFDEC253EFDA4" author="Semenov, V. B. & Egorov, L. V. & Vinogradova, E. Yu." box="[1115,1357,583,608]" pageId="5" pageNumber="214" refId="ref8070" refString="Semenov, V. B., Egorov, L. V. & Vinogradova, E. Yu. (2015) Annotirovannyi spisok stafilinid (Insecta, Coleoptera, Staphylinidae) Chuvashskoi respubliki [The staphylinid beetles annotated checklist of the Chuvash Republic]. Novoye vremya, Cheboksary, 146 pp. [in Russian]" type="book" year="2015">
|
||
Semenov
|
||
<emphasis id="FECCEA9AFFF2B03EFB2AFDED248FFDA4" box="[1227,1276,584,608]" italics="true" pageId="5" pageNumber="214">et al</emphasis>
|
||
., 2015
|
||
</bibRefCitation>
|
||
). From both these species,
|
||
<taxonomicName id="0BB84D0BFFF2B03EFE96FDC82262FD40" authority="Enushchenko, 2018" authorityName="Enushchenko" authorityYear="2018" box="[375,529,620,644]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="fontanedai" status="sp. nov.">
|
||
<emphasis id="FECCEA9AFFF2B03EFE96FDC82262FD40" box="[375,529,620,644]" italics="true" pageId="5" pageNumber="214">G. fontanedai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E5FF57E1FFF2B03EFDFAFDC82221FD41" box="[539,594,620,645]" pageId="5" pageNumber="214" rank="species">
|
||
<emphasis id="FECCEA9AFFF2B03EFDFAFDC82221FD41" bold="true" box="[539,594,620,645]" pageId="5" pageNumber="214">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs by the significantly smaller body (see the key below) and by the shape of elongated teeth on the apical margin of abdominal tergite VIII of male (these teeth in
|
||
<taxonomicName id="0BB84D0BFFF2B03EFB35FD342538FD6C" baseAuthorityName="Gyllenhal" baseAuthorityYear="1810" box="[1236,1355,656,680]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="nitidula">
|
||
<emphasis id="FECCEA9AFFF2B03EFB35FD342538FD6C" box="[1236,1355,656,680]" italics="true" pageId="5" pageNumber="214">G. nitidula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0BB84D0BFFF2B03EFA65FD34214EFD08" authorityName="Semenov" authorityYear="2015" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="pseudonitidula">
|
||
<emphasis id="FECCEA9AFFF2B03EFA65FD34214EFD08" italics="true" pageId="5" pageNumber="214">G. pseudonitidula</emphasis>
|
||
</taxonomicName>
|
||
are significantly smaller).
|
||
</paragraph>
|
||
<paragraph id="CC073688FFF2B03EFF26FD7C24CBFD34" blockId="5.[151,1437,151,1184]" box="[199,1208,727,752]" pageId="5" pageNumber="214">
|
||
For illustrations of
|
||
<taxonomicName id="0BB84D0BFFF2B03EFE7FFD7C2267FD34" baseAuthorityName="Gyllenhal" baseAuthorityYear="1810" box="[414,532,728,752]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="nitidula">
|
||
<emphasis id="FECCEA9AFFF2B03EFE7FFD7C2267FD34" box="[414,532,728,752]" italics="true" pageId="5" pageNumber="214">G. nitidula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0BB84D0BFFF2B03EFDABFD7C2363FD34" authorityName="Semenov" authorityYear="2015" box="[586,784,728,752]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="pseudonitidula">
|
||
<emphasis id="FECCEA9AFFF2B03EFDABFD7C2363FD34" box="[586,784,728,752]" italics="true" pageId="5" pageNumber="214">G. pseudonitidula</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="A8294B79FFF2B03EFCA0FD7C2446FD34" author="Semenov, V. B. & Egorov, L. V. & Vinogradova, E. Yu." box="[833,1077,727,752]" pageId="5" pageNumber="214" refId="ref8070" refString="Semenov, V. B., Egorov, L. V. & Vinogradova, E. Yu. (2015) Annotirovannyi spisok stafilinid (Insecta, Coleoptera, Staphylinidae) Chuvashskoi respubliki [The staphylinid beetles annotated checklist of the Chuvash Republic]. Novoye vremya, Cheboksary, 146 pp. [in Russian]" type="book" year="2015">
|
||
Semenov
|
||
<emphasis id="FECCEA9AFFF2B03EFC51FD7D2391FD34" box="[944,994,728,752]" italics="true" pageId="5" pageNumber="214">et al</emphasis>
|
||
. (2015
|
||
</bibRefCitation>
|
||
:
|
||
<figureCitation id="54832A0DFFF2B03EFBDEFD7324D9FD34" box="[1087,1194,727,752]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1953,1976]" captionTargetBox="[249,1371,206,1914]" captionTargetId="figure@2.[210,1377,195,1929]" captionTargetPageId="2" captionText="FIGURES 1–5. Gyrophaena affinifera sp.n.: 1—habitus, 2—aedeagus, lateral view, 3—male abdominal tergite VIII, dorsal view, 4—female abdominal tergite VIII, dorsal view, 5—female abdominal sternite, dorsal view. Scale bars: 1 mm (1), 0.1 mm (2–5)." httpUri="https://zenodo.org/record/2946786/files/figure.png" pageId="5" pageNumber="214">Figs. 2, 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="84A26503FFF2B03EFF26FD582082FC44" pageId="5" pageNumber="214" type="etymology">
|
||
<paragraph id="CC073688FFF2B03EFF26FD582082FC44" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="214">
|
||
<emphasis id="FECCEA9AFFF2B03EFF26FD582135FCD1" bold="true" box="[199,326,764,789]" pageId="5" pageNumber="214">Etymology</emphasis>
|
||
. The new species is dedicated to the Spanish shipwreck survivor Hernando de Escalante Fontaneda (
|
||
<date id="B8061048FFF2B03EFF7EFCBB2152FCFC" box="[159,289,799,824]" pageId="5" pageNumber="214" value="1536" valueMax="1575">1536–1575</date>
|
||
, dates uncertain) who lived among the Indians of
|
||
<collectingRegion id="0E7CF86AFFF2B03EFCB0FCBB23D1FCFC" box="[849,930,799,824]" country="United States of America" name="Florida" pageId="5" pageNumber="214">Florida</collectingRegion>
|
||
for 17 years. His memoir, written in 1575, is one of the most valuable contemporary contributions to the history and ethnography of the life of American Indians.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="84A26503FFF2B03EFF26FC2821F9FB64" pageId="5" pageNumber="214" type="discussion">
|
||
<paragraph id="CC073688FFF2B03EFF26FC2824A3FC29" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="214">
|
||
<emphasis id="FECCEA9AFFF2B03EFF26FC282148FC61" bold="true" box="[199,315,908,933]" pageId="5" pageNumber="214">Remarks.</emphasis>
|
||
The specimen (
|
||
<typeStatus id="1303882AFFF2B03EFE0FFC282221FC60" box="[494,594,908,932]" pageId="5" pageNumber="214" type="paratype">paratype</typeStatus>
|
||
) with green label “Am. bor.” in the collection of V.I. Motschulsky (ZMM) were collected by him between
|
||
<date id="B8061048FFF2B03EFE1AFC0B22C2FC0C" box="[507,689,943,968]" pageId="5" pageNumber="214" value="1853-11">November 1853</date>
|
||
and
|
||
<date id="B8061048FFF2B03EFD09FC0B2301FC0C" box="[744,882,943,968]" pageId="5" pageNumber="214" value="1854-03">March 1854</date>
|
||
, when he visited the
|
||
<collectingCountry id="B4AF7618FFF2B03EFBBEFC1424E4FC0C" box="[1119,1175,944,968]" name="United States of America" pageId="5" pageNumber="214">USA</collectingCountry>
|
||
(Krivokhatskiy, 2013). The specimen was noted by him as a “
|
||
<taxonomicName id="0BB84D0BFFF2B03EFDA4FC702331FC29" authority="Motch." authorityName="Motch." box="[581,834,980,1005]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="breviuscula">
|
||
<emphasis id="FECCEA9AFFF2B03EFDA4FC702295FC28" box="[581,742,980,1004]" italics="true" pageId="5" pageNumber="214">G. breviuscula</emphasis>
|
||
Motch.
|
||
</taxonomicName>
|
||
”, but no description was published.
|
||
</paragraph>
|
||
<paragraph id="CC073688FFF2B03EFF26FC5321F9FB64" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="214">
|
||
The
|
||
<taxonomicName id="0BB84D0BFFF2B03EFF1FFC5C2126FBD4" baseAuthorityName="Gyllenhal" baseAuthorityYear="1810" box="[254,341,1016,1040]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="214" phylum="Arthropoda" rank="species" species="nitidula">
|
||
<emphasis id="FECCEA9AFFF2B03EFF1FFC5C2126FBD4" box="[254,341,1016,1040]" italics="true" pageId="5" pageNumber="214">nitidula</emphasis>
|
||
</taxonomicName>
|
||
species group is erected here for this new and two above-mentioned species. This group is characterized by similar shape of the male tergite VIII and aedeagus (see the key below). Additionally, species of this group have long antennae with elongated antennomere V and subquadrate antennomeres VI–X; surface of head and pronotum moderately glossy, with well pronounced microsculpture; elytra with coarse and widely distributed punctures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |