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<document id="4BA6033AAFAD827BE5D981F00F6ECD28" ID-CLB-Dataset="32119" ID-DOI="10.5852/ejt.2017.293" ID-GBIF-Dataset="3099a544-54cd-48e5-8aac-8e1ca24c0e8b" ID-ISSN="2118-9773" ID-Zenodo-Dep="889857" ID-ZooBank="urn:lsid:zoobank.org:pub:608B2AAB-39A4-494C-9CE0-AA84916008BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505237457348" checkinUser="plazi" docAuthor="Anh D. Nguyen, Zoltán Korsós, Kuem-Hee Jang &amp; Ui-Wook Hwang" docDate="2017" docId="287287F2FFDD3328DF331EDBFDAB8659" docLanguage="en" docName="EJT-293_Nguyen.pdf" docOrigin="European Journal of Taxonomy 293" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Oxidus riukiaria" docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="D44BFF8AFFD73323DD631D42FF8F8422" masterDocTitle="A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida, Paradoxosomatidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="11" updateTime="1698468380887" updateUser="ExternalLinkService">
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<mods:title id="57731E79F86EE18BCA4490BC6C01AEA3">A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="42A0F06DA12185D34EBCD3D286450C64">Anh D. Nguyen</mods:namePart>
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<mods:namePart id="1A9B4D26219815ED46BD038ACC9DEBAD">Zoltán Korsós</mods:namePart>
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<mods:namePart id="142F538D202248DFFE85DF8383D2E18C">Kuem-Hee Jang</mods:namePart>
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<mods:namePart id="A9BF1AFE1CBE95EC45ECF2CD8E40F020">Ui-Wook Hwang</mods:namePart>
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<mods:title id="78F4040C53153B723739E7562BF2E116">European Journal of Taxonomy</mods:title>
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<treatment id="287287F2FFDD3328DF331EDBFDAB8659" ID-DOI="http://doi.org/10.5281/zenodo.5672740" ID-GBIF-Taxon="134034196" ID-Zenodo-Dep="5672740" LSID="urn:lsid:plazi:treatment:287287F2FFDD3328DF331EDBFDAB8659" httpUri="http://treatment.plazi.org/id/287287F2FFDD3328DF331EDBFDAB8659" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="E8C1656FFFDD3329DF331EDBFC6C8791" box="[592,995,921,948]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="A06436E4FFDD3329DF331EDBFC6C8791" blockId="10.[592,995,921,983]" box="[592,995,921,948]" pageId="10" pageNumber="11">
<heading id="FB2C8188FFDD3329DF331EDBFC6C8791" box="[592,995,921,948]" centered="true" fontSize="11" level="4" pageId="10" pageNumber="11" reason="4">
<taxonomicName id="67DB4D67FFDD3329DF331EDBFC6C8791" ID-CoL="4BBY4" authority="(Verhoeff, 1940)" baseAuthorityName="Verhoeff" baseAuthorityYear="1940" box="[592,995,921,948]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riukiaria">
<emphasis id="92AFEAF6FFDD3329DF331EDBFC958796" bold="true" box="[592,794,921,948]" italics="true" pageId="10" pageNumber="11">Oxidus riukiaria</emphasis>
(
<bibRefCitation id="C44A4B15FFDD3329DE4B1EDBFC548791" author="Verhoeff K. W." box="[808,987,921,947]" journalOrPublisher="Zoologischer Anzeiger" pageId="10" pageNumber="11" pagination="129 - 145" part="131" refString="Verhoeff K. W. 1940. Zur Kenntnis ostasiatischer Diplopoden. V (Polydesmoidea). Zoologischer Anzeiger 131 (5 / 6): 129 - 145." title="Zur Kenntnis ostasiatischer Diplopoden. V (Polydesmoidea)" type="journal article" year="1940">Verhoeff, 1940</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDD3329DF841EFEFCC387F5" box="[743,844,956,983]" pageId="10" pageNumber="11" type="description">
<paragraph id="A06436E4FFDD3329DF841EFEFCC387F5" blockId="10.[592,995,921,983]" box="[743,844,956,983]" pageId="10" pageNumber="11">
<figureCitation id="38E02A61FFDD3329DF841EFEFCC387F5" box="[743,844,956,983]" captionStart-0="Fig. 5" captionStart-1="Fig. 6" captionStart-2="Fig. 7" captionStartId-0="11.[189,232,1965,1991]" captionStartId-1="12.[189,232,1181,1207]" captionStartId-2="12.[191,234,1965,1991]" captionTargetBox-0="[310,1278,850,1931]" captionTargetBox-1="[233,1337,264,1077]" captionTargetBox-2="[190,1400,1306,1906]" captionTargetId-0="figure@11.[310,1278,850,1931]" captionTargetId-1="figure@12.[191,1401,264,1125]" captionTargetId-2="figure@12.[190,1400,1306,1906]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="12" captionText-0="Fig. 5. Oxidus riukiaria (Verhoeff, 1940) from Okinawa, Japan. A. Entire body, length ca 20 mm. B C. Segments 8 9 10. B. ♂. C. ♀. Scale bars = 1 mm. (photos by Z. Korsós)" captionText-1="Fig. 6. Left gonopod of Oxidus riukiaria (Verhoeff, 1940). A. Lateral view. B. Postfemoral region, lateral view. Redrawn from Verhoeff 1940. No scale bars." captionText-2="Fig. 7. Left gonopod of Oxidus riukiaria (Verhoeff, 1940), from sample IEBR-H 470. A. Lateroventral view. B. Ventral view. C. Postfemoral region, ventral view. Note: z = spine z, but broken." httpUri-0="https://zenodo.org/record/889867/files/figure.png" httpUri-1="https://zenodo.org/record/889869/files/figure.png" httpUri-2="https://zenodo.org/record/889871/files/figure.png" pageId="10" pageNumber="11">Figs 57</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDD3329DDDE1EBCFC948082" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="A06436E4FFDD3329DDDE1EBCFCB9803A" blockId="10.[189,822,1021,1048]" box="[189,822,1021,1048]" pageId="10" pageNumber="11">
<taxonomicName id="67DB4D67FFDD3329DDDE1EBCFD4C803A" ID-CoL="6SWNP" authority="Verhoeff, 1940: 139" authorityName="Verhoeff" authorityPageNumber="139" authorityYear="1940" box="[189,707,1022,1048]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riukiaria">
<emphasis id="92AFEAF6FFDD3329DDDE1EBCFE41803A" box="[189,462,1022,1048]" italics="true" pageId="10" pageNumber="11">Orthomorpha riukiaria</emphasis>
<bibRefCitation id="C44A4B15FFDD3329DCB61EBCFD06803A" author="Verhoeff K. W." box="[469,649,1022,1048]" journalOrPublisher="Zoologischer Anzeiger" pageId="10" pageNumber="11" pagination="129 - 145" part="131" refString="Verhoeff K. W. 1940. Zur Kenntnis ostasiatischer Diplopoden. V (Polydesmoidea). Zoologischer Anzeiger 131 (5 / 6): 129 - 145." title="Zur Kenntnis ostasiatischer Diplopoden. V (Polydesmoidea)" type="journal article" year="1940">Verhoeff, 1940</bibRefCitation>
: 139
</taxonomicName>
, fgs 56.
</paragraph>
<paragraph id="A06436E4FFDD3329DDDE197DFCBC807B" blockId="10.[189,819,1087,1184]" box="[189,819,1087,1114]" pageId="10" pageNumber="11">
<taxonomicName id="67DB4D67FFDD3329DDDE197DFE41807B" ID-CoL="6SWNP" authorityName="Verhoeff" authorityYear="1940" box="[189,462,1087,1113]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riukiaria">
<emphasis id="92AFEAF6FFDD3329DDDE197DFE41807B" box="[189,462,1087,1113]" italics="true" pageId="10" pageNumber="11">Orthomorpha riukiaria</emphasis>
</taxonomicName>
<bibRefCitation id="C44A4B15FFDD3329DC88197DFD268078" author="Takakuwa Y." box="[491,681,1087,1114]" journalOrPublisher="Japan Society for the Promotion of Science, Tokyo" pageId="10" pageNumber="11" refString="Takakuwa Y. 1954. Diplopoden aus Japan und ihn angrenzenden Gebieten. Japan Society for the Promotion of Science, Tokyo." title="Diplopoden aus Japan und ihn angrenzenden Gebieten" type="book" year="1954">Takakuwa 1954</bibRefCitation>
: 39, fg. 34.
</paragraph>
<paragraph id="A06436E4FFDD3329DDDE1920FDFA805E" blockId="10.[189,819,1087,1184]" box="[189,629,1122,1149]" pageId="10" pageNumber="11">
<taxonomicName id="67DB4D67FFDD3329DDDE1920FE04805E" ID-CoL="4BBY3" baseAuthorityName="Verhoeff" baseAuthorityYear="1940" box="[189,395,1122,1148]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riukiarius">
<emphasis id="92AFEAF6FFDD3329DDDE1920FE04805E" box="[189,395,1122,1148]" italics="true" pageId="10" pageNumber="11">Oxidus riukiarius</emphasis>
</taxonomicName>
<bibRefCitation id="C44A4B15FFDD3329DCCB1920FDCA805E" author="Jeekel C. A. W." box="[424,581,1122,1148]" journalOrPublisher="Studies on the Fauna of Suriname and other Guyanas" pageId="10" pageNumber="11" pagination="1 - 157" part="4" refString="Jeekel C. A. W. 1963 a. Diplopoda of Guiana. Studies on the Fauna of Suriname and other Guyanas 4 (11): 1 - 157." title="Diplopoda of Guiana" type="journal article" year="1963" yearSuffix="a">Jeekel 1963a</bibRefCitation>
: 20.
</paragraph>
<paragraph id="A06436E4FFDD3329DDDE19C7FC948082" blockId="10.[189,819,1087,1184]" box="[189,795,1157,1184]" pageId="10" pageNumber="11">
<taxonomicName id="67DB4D67FFDD3329DDDE19C7FEF080BD" ID-CoL="4BBY4" baseAuthorityName="Verhoeff" baseAuthorityYear="1940" box="[189,383,1157,1183]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riukiaria">
<emphasis id="92AFEAF6FFDD3329DDDE19C7FEF080BD" box="[189,383,1157,1183]" italics="true" pageId="10" pageNumber="11">Oxidus riukiaria</emphasis>
</taxonomicName>
<bibRefCitation id="C44A4B15FFDD3329DCFE19C4FD418082" author="Nguyen A. D. &amp; Sierwald P." box="[413,718,1157,1184]" journalOrPublisher="CheckList" pageId="10" pageNumber="11" pagination="1132 - 1353" part="9" refString="Nguyen A. D. &amp; Sierwald P. 2013. A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). CheckList 9: 1132 - 1353." title="A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida)" type="journal article" year="2013">Nguyen &amp; Sierwald 2013</bibRefCitation>
: 1295.
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDD3329DDDE1985FAF88249" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="A06436E4FFDD3329DDDE1985FE2580C3" blockId="10.[189,426,1223,1249]" box="[189,426,1223,1249]" pageId="10" pageNumber="11">
<heading id="FB2C8188FFDD3329DDDE1985FE2580C3" bold="true" box="[189,426,1223,1249]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="2">
<emphasis id="92AFEAF6FFDD3329DDDE1985FE2580C3" bold="true" box="[189,426,1223,1249]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A06436E4FFDD3329DDDE19B4FAF881C1" blockId="10.[189,1406,1269,1507]" pageId="10" pageNumber="11">JAPAN: 4 ♂♂, 4 ♀♀, Central Ryukyus, Okinawa-jima, Ogimi village, Nerome, above road no.58, 26°4137″ N, 128°0643″ E, 37 m a.s.l., 25 Sep. 2012, leg. Z. Korsós (IEBR-H470); 2 ♂♂, 2 ♀♀, Okinawa-jima, Yanbaru, Ogimi village, Nunha, Okuni rindo, 26°4110.1″ N, 128°0902.7″ E, 15 Sep. 2012, leg. Nakamura Y. (IEBR-H500); 2 ♂♂, Central Ryukyus, Okinawa Island, Yanbaru, Kunigami village, Cape Hedo, Ginama, broad-leaved evergreen forest, 176 m, 26°4952″ N, 128°1620″ E, 5 Apr. 2011, leg. Z. Korsós (HNHM-355); 1 ♂, Northern Ryukyus, Tokara group, Kuchino-shima Island, around Kuroshio-so guesthouse, 29°5924″ N, 129°5521″ E, 63 m, 14 Oct. 2012, leg. Z. Korsós (HNHM-477).</paragraph>
<paragraph id="A06436E4FFDD3329DD891B48FC858207" blockId="10.[189,1399,1546,1643]" box="[234,778,1546,1573]" pageId="10" pageNumber="11">
<emphasis id="92AFEAF6FFDD3329DD891B48FE158206" bold="true" box="[234,410,1546,1572]" pageId="10" pageNumber="11">Other records</emphasis>
(newly found ones underlined)
</paragraph>
<paragraph id="A06436E4FFDD3329DDDE1B6FFAF88249" blockId="10.[189,1399,1546,1643]" pageId="10" pageNumber="11">JAPAN: Northern Ryukyus, Tokara Group: Kuchino-shima Island; Central Ryukyus: Okinawa-jima, Kume-jima, Iheya-jima, Aguni-jima, Zamami-jima, Aka-jima, Geruma-jima (observations by Z. Korsós).</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDD3329DDDE1BD0FDBB83A8" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="A06436E4FFDD3329DDDE1BD0FEB9828E" blockId="10.[189,1399,1682,1930]" box="[189,310,1682,1708]" pageId="10" pageNumber="11">
<heading id="FB2C8188FFDD3329DDDE1BD0FEB9828E" bold="true" box="[189,310,1682,1708]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="6">
<emphasis id="92AFEAF6FFDD3329DDDE1BD0FEB9828E" bold="true" box="[189,310,1682,1708]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A06436E4FFDD3329DDDE1B82FDBB83A8" blockId="10.[189,1399,1682,1930]" pageId="10" pageNumber="11">
<taxonomicName id="67DB4D67FFDD3329DDDE1B82FEF182F8" baseAuthorityName="Verhoeff" baseAuthorityYear="1940" box="[189,382,1728,1754]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riukiaria">
<emphasis id="92AFEAF6FFDD3329DDDE1B82FEF182F8" box="[189,382,1728,1754]" italics="true" pageId="10" pageNumber="11">Oxidus riukiaria</emphasis>
</taxonomicName>
can be distinguished from all other species of
<taxonomicName id="67DB4D67FFDD3329DEF71B82FC6882F8" authorityName="Cook" authorityYear="1911" box="[916,999,1728,1754]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="92AFEAF6FFDD3329DEF71B82FC6882F8" box="[916,999,1728,1754]" italics="true" pageId="10" pageNumber="11">Oxidus</emphasis>
</taxonomicName>
by its smaller size (19.320.6 mm in length, width of pro- and metazonae 1.51.8 mm and 2.12.2 mm); gonopod coxae more or less stouter and longer than femorite; femorite narrow at base, and strongly expanded towards distal end; gonopod process
<emphasis id="92AFEAF6FFDD3329DCEC1A68FE118366" box="[399,414,1834,1860]" italics="true" pageId="10" pageNumber="11">h</emphasis>
hook-like, larger and getting narrower towards pointed tip, spine
<emphasis id="92AFEAF6FFDD3329D9DA1A69FB4B8366" box="[1209,1220,1835,1860]" italics="true" pageId="10" pageNumber="11">z</emphasis>
small, pointed spiniform; postfemoral lamina
<emphasis id="92AFEAF6FFDD3329DF4A1A0FFDBE8345" box="[553,561,1869,1895]" italics="true" pageId="10" pageNumber="11">l</emphasis>
triangularly rounded; mesal lobule not seperated from solenophore tip, which is distinctly emarginated.
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDC3328DDDE1C49FB878576" box="[189,1032,267,340]" pageId="11" pageNumber="12" type="description">
<paragraph id="A06436E4FFDC3328DDDE1C49FF728507" blockId="11.[189,253,267,293]" box="[189,253,267,293]" pageId="11" pageNumber="12">
<heading id="FB2C8188FFDC3328DDDE1C49FF728507" allCaps="true" bold="true" box="[189,253,267,293]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="92AFEAF6FFDC3328DDDE1C49FF728507" bold="true" box="[189,253,267,293]" pageId="11" pageNumber="12">DNA</emphasis>
</heading>
</paragraph>
<paragraph id="A06436E4FFDC3328DDDE1C78FB878576" blockId="11.[189,1032,313,340]" box="[189,1032,313,340]" pageId="11" pageNumber="12">
COI and 16S barcode data (partial) are deposited in GenBank (
<tableCitation id="ED59035FFFDC3328DEFC1C7BFC788576" box="[927,1015,313,340]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionText="Table 1. Details of specimens analysed for DNA, including GenBank numbers." pageId="11" pageNumber="12">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDC3328DDDE1C39FF798628" pageId="11" pageNumber="12" type="discussion">
<paragraph id="A06436E4FFDC3328DDDE1C39FEA085B7" blockId="11.[189,303,379,405]" box="[189,303,379,405]" pageId="11" pageNumber="12">
<heading id="FB2C8188FFDC3328DDDE1C39FEA085B7" bold="true" box="[189,303,379,405]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="2">
<emphasis id="92AFEAF6FFDC3328DDDE1C39FEA085B7" bold="true" box="[189,303,379,405]" pageId="11" pageNumber="12">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="A06436E4FFDC3328DDDE1CEBFF798628" blockId="11.[189,1398,425,522]" pageId="11" pageNumber="12">
This species was originally described by
<bibRefCitation id="C44A4B15FFDC3328DFC01CEBFCD185E1" author="Verhoeff K. W." box="[675,862,425,451]" journalOrPublisher="Zoologischer Anzeiger" pageId="11" pageNumber="12" pagination="129 - 145" part="131" refString="Verhoeff K. W. 1940. Zur Kenntnis ostasiatischer Diplopoden. V (Polydesmoidea). Zoologischer Anzeiger 131 (5 / 6): 129 - 145." title="Zur Kenntnis ostasiatischer Diplopoden. V (Polydesmoidea)" type="journal article" year="1940">Verhoeff (1940)</bibRefCitation>
in the genus
<taxonomicName id="67DB4D67FFDC3328DE9D1CEBFB1085E1" authorityName="Bollman" authorityYear="1893" box="[1022,1183,425,451]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="92AFEAF6FFDC3328DE9D1CEBFB1085E1" box="[1022,1183,425,451]" italics="true" pageId="11" pageNumber="12">Orthomorpha</emphasis>
</taxonomicName>
, but was assigned to
<taxonomicName id="67DB4D67FFDC3328DDB81C8FFEA185C5" authorityName="Cook" authorityYear="1911" box="[219,302,461,487]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="92AFEAF6FFDC3328DDB81C8FFEA185C5" box="[219,302,461,487]" italics="true" pageId="11" pageNumber="12">Oxidus</emphasis>
</taxonomicName>
by
<bibRefCitation id="C44A4B15FFDC3328DC391C8EFD8685C5" author="Jeekel C. A. W." box="[346,521,460,487]" journalOrPublisher="Studies on the Fauna of Suriname and other Guyanas" pageId="11" pageNumber="12" pagination="1 - 157" part="4" refString="Jeekel C. A. W. 1963 a. Diplopoda of Guiana. Studies on the Fauna of Suriname and other Guyanas 4 (11): 1 - 157." title="Diplopoda of Guiana" type="journal article" year="1963" yearSuffix="a">Jeekel (1963a)</bibRefCitation>
. The taxonomic position of the species is also well supported by molecular data.
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDC3328DDDE1F73FDAB8659" box="[189,548,561,635]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="A06436E4FFDC3328DDDE1F73FED98669" blockId="11.[189,342,561,587]" box="[189,342,561,587]" pageId="11" pageNumber="12">
<heading id="FB2C8188FFDC3328DDDE1F73FED98669" bold="true" box="[189,342,561,587]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="2">
<emphasis id="92AFEAF6FFDC3328DDDE1F73FED98669" bold="true" box="[189,342,561,587]" pageId="11" pageNumber="12">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="A06436E4FFDC3328DDDE1F22FDAB8659" blockId="11.[189,548,608,635]" box="[189,548,608,635]" pageId="11" pageNumber="12">South Japan (Ryukyu Islands).</paragraph>
</subSubSection>
</treatment>
</document>