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<document id="B2479C09890C1EFB9B4FF50A23824E35" ID-CLB-Dataset="32119" ID-DOI="10.5852/ejt.2017.293" ID-GBIF-Dataset="3099a544-54cd-48e5-8aac-8e1ca24c0e8b" ID-ISSN="2118-9773" ID-Zenodo-Dep="889857" ID-ZooBank="urn:lsid:zoobank.org:pub:608B2AAB-39A4-494C-9CE0-AA84916008BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505237457348" checkinUser="plazi" docAuthor="Anh D. Nguyen, Zoltán Korsós, Kuem-Hee Jang &amp; Ui-Wook Hwang" docDate="2017" docId="287287F2FFDC332EDF0F1FE3FC7E8707" docLanguage="en" docName="EJT-293_Nguyen.pdf" docOrigin="European Journal of Taxonomy 293" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Oxidus avia" docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="D44BFF8AFFD73323DD631D42FF8F8422" masterDocTitle="A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida, Paradoxosomatidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="12" updateTime="1698468380887" updateUser="ExternalLinkService">
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<mods:title id="720C2ED5EBE981727B39B12EFAE23D35">A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="254E81475B26B9796143D0DD004C6178">Anh D. Nguyen</mods:namePart>
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<mods:namePart id="466F456D09A5ED3A5C0D87A392C75D6B">Zoltán Korsós</mods:namePart>
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<mods:namePart id="3D81D1011ED3E052997212C3A6D40E46">Kuem-Hee Jang</mods:namePart>
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<mods:namePart id="B802B1391564461BDDEFD200B389191B">Ui-Wook Hwang</mods:namePart>
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<mods:date id="AB474D1884AC3B3A50D263B14F04448E">2017</mods:date>
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<treatment id="287287F2FFDC332EDF0F1FE3FC7E8707" ID-DOI="http://doi.org/10.5281/zenodo.5672742" ID-GBIF-Taxon="134034197" ID-Zenodo-Dep="5672742" LSID="urn:lsid:plazi:treatment:287287F2FFDC332EDF0F1FE3FC7E8707" httpUri="http://treatment.plazi.org/id/287287F2FFDC332EDF0F1FE3FC7E8707" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection id="E8C1656FFFDC3328DF0F1FE3FC488699" box="[620,967,673,700]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="A06436E4FFDC3328DF0F1FE3FC488699" blockId="11.[620,967,673,734]" box="[620,967,673,700]" pageId="11" pageNumber="12">
<heading id="FB2C8188FFDC3328DF0F1FE3FC488699" box="[620,967,673,700]" centered="true" fontSize="11" level="4" pageId="11" pageNumber="12" reason="4">
<taxonomicName id="67DB4D67FFDC3328DF0F1FE3FC488699" ID-CoL="4BBXL" authority="(Verhoeff, 1937)" baseAuthorityName="Verhoeff" baseAuthorityYear="1937" box="[620,967,673,700]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="avia">
<emphasis id="92AFEAF6FFDC3328DF0F1FE3FD728699" bold="true" box="[620,765,673,700]" italics="true" pageId="11" pageNumber="12">Oxidus avia</emphasis>
(
<bibRefCitation id="C44A4B15FFDC3328DE6F1FE3FC308699" author="Verhoeff K. W." box="[780,959,673,699]" journalOrPublisher="Zoologischer Anzeiger" pageId="11" pageNumber="12" pagination="33 - 40" part="119" refString="Verhoeff K. W. 1937. Zur Kenntnis ostasiatischer Diplopoden. II. Zoologischer Anzeiger 119 (1 / 2): 33 - 40." title="Zur Kenntnis ostasiatischer Diplopoden. II" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDC3328DF8D1F86FCC886FC" box="[750,839,708,734]" pageId="11" pageNumber="12" type="description">
<paragraph id="A06436E4FFDC3328DF8D1F86FCC886FC" blockId="11.[620,967,673,734]" box="[750,839,708,734]" pageId="11" pageNumber="12">
<figureCitation id="38E02A61FFDC3328DF8D1F86FCC886FC" box="[750,839,708,734]" captionStart="Fig. 8" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[331,1205,1027,1871]" captionTargetId="figure@13.[189,1399,1003,1915]" captionTargetPageId="13" captionText="Fig. 8. A. Left gonopod of Oxidus avia (Verhoeff, 1937), lateral view. B. Right gonopod of Oxidus obtusus (Takakuwa, 1942), mesal view. Redrawn from Verhoeff 1937 and Takakuwa 1942 a, respectively. No scale bars." httpUri="https://zenodo.org/record/889873/files/figure.png" pageId="11" pageNumber="12">Fig. 8A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDC332EDDDE1E44FADE8504" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="A06436E4FFDC3328DDDE1E44FD418702" blockId="11.[189,718,774,800]" box="[189,718,774,800]" pageId="11" pageNumber="12">
<taxonomicName id="67DB4D67FFDC3328DDDE1E44FDF18702" ID-CoL="758NY" authority="Verhoeff, 1937: 33" authorityName="Verhoeff" authorityPageNumber="33" authorityYear="1937" box="[189,638,774,800]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="avia">
<emphasis id="92AFEAF6FFDC3328DDDE1E44FE168702" box="[189,409,774,800]" italics="true" pageId="11" pageNumber="12">Orthomorpha avia</emphasis>
<bibRefCitation id="C44A4B15FFDC3328DCFC1E44FDDC8702" author="Verhoeff K. W." box="[415,595,774,800]" journalOrPublisher="Zoologischer Anzeiger" pageId="11" pageNumber="12" pagination="33 - 40" part="119" refString="Verhoeff K. W. 1937. Zur Kenntnis ostasiatischer Diplopoden. II. Zoologischer Anzeiger 119 (1 / 2): 33 - 40." title="Zur Kenntnis ostasiatischer Diplopoden. II" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
: 33
</taxonomicName>
, fg. 1.
</paragraph>
<caption id="F4A4666CFFDC3328DDDE1AEFFB9883C9" httpUri="https://zenodo.org/record/889867/files/figure.png" pageId="11" pageNumber="12" targetBox="[310,1278,850,1931]" targetPageId="11">
<paragraph id="A06436E4FFDC3328DDDE1AEFFB9883C9" blockId="11.[189,1398,1965,2027]" pageId="11" pageNumber="12">
<emphasis id="92AFEAF6FFDC3328DDDE1AEFFE8583EA" bold="true" box="[189,266,1965,1992]" pageId="11" pageNumber="12">Fig. 5.</emphasis>
<taxonomicName id="67DB4D67FFDC3328DC721AEFFD1283EA" ID-CoL="4BBY4" authority="(Verhoeff, 1940)" authorityName="Verhoeff" authorityYear="1940" baseAuthorityName="Verhoeff" baseAuthorityYear="1940" box="[273,669,1965,1992]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="riukiaria">
<emphasis id="92AFEAF6FFDC3328DC721AEFFE5C83E5" box="[273,467,1965,1991]" italics="true" pageId="11" pageNumber="12">Oxidus riukiaria</emphasis>
(Verhoeff, 1940)
</taxonomicName>
from Okinawa, Japan.
<emphasis id="92AFEAF6FFDC3328DECE1AEFFC4D83E5" bold="true" box="[941,962,1965,1991]" pageId="11" pageNumber="12">A</emphasis>
. Entire body, length ca 20 mm.
<emphasis id="92AFEAF6FFDC3328D85B1AECFAE083E5" bold="true" box="[1336,1391,1965,1992]" pageId="11" pageNumber="12">BC</emphasis>
. Segments 8910.
<emphasis id="92AFEAF6FFDC3328DCFE1A93FE3E83C9" bold="true" box="[413,433,2001,2027]" pageId="11" pageNumber="12">B</emphasis>
. ♂.
<emphasis id="92AFEAF6FFDC3328DC871A92FE7683C8" bold="true" box="[484,505,2000,2026]" pageId="11" pageNumber="12">C</emphasis>
. ♀. Scale bars = 1 mm. (photos by Z. Korsós)
</paragraph>
</caption>
<caption id="F4A4666CFFDB332FDDDE19DFFCEE80F9" httpUri="https://zenodo.org/record/889869/files/figure.png" pageId="12" pageNumber="13" targetBox="[233,1337,264,1077]" targetPageId="12">
<paragraph id="A06436E4FFDB332FDDDE19DFFCEE80F9" blockId="12.[189,1398,1181,1243]" pageId="12" pageNumber="13">
<emphasis id="92AFEAF6FFDB332FDDDE19DFFE81809A" bold="true" box="[189,270,1181,1208]" pageId="12" pageNumber="13">Fig. 6.</emphasis>
Left gonopod of
<taxonomicName id="67DB4D67FFDB332FDC8819DCFC08809A" ID-CoL="4BBY4" authority="(Verhoeff, 1940)" authorityName="Verhoeff" authorityYear="1940" baseAuthorityName="Verhoeff" baseAuthorityYear="1940" box="[491,903,1182,1208]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="riukiaria">
<emphasis id="92AFEAF6FFDB332FDC8819DCFD3E809A" box="[491,689,1182,1208]" italics="true" pageId="12" pageNumber="13">Oxidus riukiaria</emphasis>
(Verhoeff, 1940)
</taxonomicName>
.
<emphasis id="92AFEAF6FFDB332FDEF519DFFC248095" bold="true" box="[918,939,1181,1207]" pageId="12" pageNumber="13">A</emphasis>
. Lateral view.
<emphasis id="92AFEAF6FFDB332FD90619DCFBF6809A" bold="true" box="[1125,1145,1182,1208]" pageId="12" pageNumber="13">B</emphasis>
. Postfemoral region, lateral view. Redrawn from Verhoeff 1940. No scale bars.
</paragraph>
</caption>
<caption id="F4A4666CFFDB332FDDDC1AEFFB5C83C9" httpUri="https://zenodo.org/record/889871/files/figure.png" pageId="12" pageNumber="13" targetBox="[190,1400,1306,1906]" targetPageId="12">
<paragraph id="A06436E4FFDB332FDDDC1AEFFB5C83C9" blockId="12.[191,1400,1965,2027]" pageId="12" pageNumber="13">
<emphasis id="92AFEAF6FFDB332FDDDC1AEFFE8283EA" bold="true" box="[191,269,1965,1992]" pageId="12" pageNumber="13">Fig. 7.</emphasis>
Left gonopod of
<taxonomicName id="67DB4D67FFDB332FDCBE1AEFFCFF83EA" ID-CoL="4BBY4" authority="(Verhoeff, 1940)" authorityName="Verhoeff" authorityYear="1940" baseAuthorityName="Verhoeff" baseAuthorityYear="1940" box="[477,880,1965,1992]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="riukiaria">
<emphasis id="92AFEAF6FFDB332FDCBE1AEFFD2F83E5" box="[477,672,1965,1991]" italics="true" pageId="12" pageNumber="13">Oxidus riukiaria</emphasis>
(Verhoeff, 1940)
</taxonomicName>
, from sample IEBR-H470.
<emphasis id="92AFEAF6FFDB332FD9D51AEFFB4483E5" bold="true" box="[1206,1227,1965,1991]" pageId="12" pageNumber="13">A</emphasis>
. Lateroventral view.
<emphasis id="92AFEAF6FFDB332FDC671A93FE9783C9" bold="true" box="[260,280,2001,2027]" pageId="12" pageNumber="13">B</emphasis>
. Ventral view.
<emphasis id="92AFEAF6FFDB332FDCAB1A92FE5283C8" bold="true" box="[456,477,2000,2026]" pageId="12" pageNumber="13">C</emphasis>
. Postfemoral region, ventral view. Note: z = spine
<emphasis id="92AFEAF6FFDB332FD9501A90FBB183C9" box="[1075,1086,2002,2027]" italics="true" pageId="12" pageNumber="13">z</emphasis>
, but broken.
</paragraph>
</caption>
<paragraph id="A06436E4FFDA332EDDDE1C49FADE8504" blockId="13.[189,1361,267,294]" box="[189,1361,267,294]" pageId="13" pageNumber="14">
<taxonomicName id="67DB4D67FFDA332EDDDE1C49FEC58507" ID-CoL="4BBXL" baseAuthorityName="Verhoeff" baseAuthorityYear="1937" box="[189,330,267,293]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="avia">
<emphasis id="92AFEAF6FFDA332EDDDE1C49FEC58507" box="[189,330,267,293]" italics="true" pageId="13" pageNumber="14">Oxidus avia</emphasis>
</taxonomicName>
<bibRefCitation id="C44A4B15FFDA332EDC041C49FDAA8504" author="Takakuwa Y." box="[359,549,267,294]" journalOrPublisher="Japan Society for the Promotion of Science, Tokyo" pageId="13" pageNumber="14" refString="Takakuwa Y. 1954. Diplopoden aus Japan und ihn angrenzenden Gebieten. Japan Society for the Promotion of Science, Tokyo." title="Diplopoden aus Japan und ihn angrenzenden Gebieten" type="book" year="1954">Takakuwa 1954</bibRefCitation>
: 38, fg. 33. —
<bibRefCitation id="C44A4B15FFDA332EDFB81C49FCF88507" author="Jeekel C. A. W." box="[731,887,267,293]" journalOrPublisher="Studies on the Fauna of Suriname and other Guyanas" pageId="13" pageNumber="14" pagination="1 - 157" part="4" refString="Jeekel C. A. W. 1963 a. Diplopoda of Guiana. Studies on the Fauna of Suriname and other Guyanas 4 (11): 1 - 157." title="Diplopoda of Guiana" type="journal article" year="1963" yearSuffix="a">Jeekel 1963a</bibRefCitation>
: 20. —
<bibRefCitation id="C44A4B15FFDA332EDEB01C4EFA8B8507" author="Nguyen A. D. &amp; Sierwald P." box="[979,1284,267,294]" journalOrPublisher="CheckList" pageId="13" pageNumber="14" pagination="1132 - 1353" part="9" refString="Nguyen A. D. &amp; Sierwald P. 2013. A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). CheckList 9: 1132 - 1353." title="A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida)" type="journal article" year="2013">Nguyen &amp; Sierwald 2013</bibRefCitation>
: 1295.
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDA332EDDDE1C0FFE6D8600" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="A06436E4FFDA332EDDDE1C0FFEB98545" blockId="13.[189,1398,333,546]" box="[189,310,333,359]" pageId="13" pageNumber="14">
<heading id="FB2C8188FFDA332EDDDE1C0FFEB98545" bold="true" box="[189,310,333,359]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="6">
<emphasis id="92AFEAF6FFDA332EDDDE1C0FFEB98545" bold="true" box="[189,310,333,359]" pageId="13" pageNumber="14">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A06436E4FFDA332EDDDE1C39FE6D8600" blockId="13.[189,1398,333,546]" pageId="13" pageNumber="14">
This species can be separated from all other species of
<taxonomicName id="67DB4D67FFDA332EDE551C39FC0685B7" authorityName="Cook" authorityYear="1911" box="[822,905,379,405]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="92AFEAF6FFDA332EDE551C39FC0685B7" box="[822,905,379,405]" italics="true" pageId="13" pageNumber="14">Oxidus</emphasis>
</taxonomicName>
by its much darker, almost uniformly dark brown body, slender base of gonopod solenophore, more clearly and obtusely emarginated posterior edge of telson; smaller postfemoral lamina
<emphasis id="92AFEAF6FFDA332EDFDA1C80FD4E85FE" box="[697,705,450,476]" italics="true" pageId="13" pageNumber="14">l</emphasis>
; stouter spine
<emphasis id="92AFEAF6FFDA332EDE0E1C81FCF785FE" box="[877,888,451,476]" italics="true" pageId="13" pageNumber="14">z</emphasis>
; process
<emphasis id="92AFEAF6FFDA332EDE841C80FC7985FE" box="[999,1014,450,476]" italics="true" pageId="13" pageNumber="14">h</emphasis>
more erect, slender and pointed; the base of solenophore less protruded, more strongly rounded; mesal lobule distinctly separated from rounded solenophore tip.
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDA332EDDDE1F08FC9D8694" pageId="13" pageNumber="14" type="discussion">
<paragraph id="A06436E4FFDA332EDDDE1F08FEA08646" blockId="13.[189,303,586,612]" box="[189,303,586,612]" pageId="13" pageNumber="14">
<heading id="FB2C8188FFDA332EDDDE1F08FEA08646" bold="true" box="[189,303,586,612]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="2">
<emphasis id="92AFEAF6FFDA332EDDDE1F08FEA08646" bold="true" box="[189,303,586,612]" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="A06436E4FFDA332EDDDE1F3AFC9D8694" blockId="13.[189,1399,632,694]" pageId="13" pageNumber="14">
The species was originally described by
<bibRefCitation id="C44A4B15FFDA332EDFD41F3AFCF786B0" author="Verhoeff K. W." box="[695,888,632,658]" journalOrPublisher="Zoologischer Anzeiger" pageId="13" pageNumber="14" pagination="33 - 40" part="119" refString="Verhoeff K. W. 1937. Zur Kenntnis ostasiatischer Diplopoden. II. Zoologischer Anzeiger 119 (1 / 2): 33 - 40." title="Zur Kenntnis ostasiatischer Diplopoden. II" type="journal article" year="1937">Verhoeff (1937)</bibRefCitation>
under the genus
<taxonomicName id="67DB4D67FFDA332ED9361F3AFB7986B0" authorityName="Bollman" authorityYear="1893" box="[1109,1270,632,658]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="92AFEAF6FFDA332ED9361F3AFB7986B0" box="[1109,1270,632,658]" italics="true" pageId="13" pageNumber="14">Orthomorpha</emphasis>
</taxonomicName>
, then was assigned to the genus
<taxonomicName id="67DB4D67FFDA332EDCDD1FD9FD9E8697" authorityName="Cook" authorityYear="1911" box="[446,529,667,693]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="92AFEAF6FFDA332EDCDD1FD9FD9E8697" box="[446,529,667,693]" italics="true" pageId="13" pageNumber="14">Oxidus</emphasis>
</taxonomicName>
by
<bibRefCitation id="C44A4B15FFDA332EDF5E1FD9FC818694" author="Takakuwa Y." box="[573,782,667,694]" journalOrPublisher="Japan Society for the Promotion of Science, Tokyo" pageId="13" pageNumber="14" refString="Takakuwa Y. 1954. Diplopoden aus Japan und ihn angrenzenden Gebieten. Japan Society for the Promotion of Science, Tokyo." title="Diplopoden aus Japan und ihn angrenzenden Gebieten" type="book" year="1954">Takakuwa (1954)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E8C1656FFFDA332EDDDE1F9FFC7E8707" box="[189,1009,733,806]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="A06436E4FFDA332EDDDE1F9FFED986D5" blockId="13.[189,342,733,759]" box="[189,342,733,759]" pageId="13" pageNumber="14">
<heading id="FB2C8188FFDA332EDDDE1F9FFED986D5" bold="true" box="[189,342,733,759]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="2">
<emphasis id="92AFEAF6FFDA332EDDDE1F9FFED986D5" bold="true" box="[189,342,733,759]" pageId="13" pageNumber="14">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="A06436E4FFDA332EDDDE1E49FC7E8707" blockId="13.[189,1009,779,806]" box="[189,1009,779,806]" pageId="13" pageNumber="14">South Japan (Ryukyu Islands): Ishigaki-jima, Yonaguni-jima; Taiwan.</paragraph>
</subSubSection>
</treatment>
</document>