170 lines
20 KiB
XML
170 lines
20 KiB
XML
<document id="FDF06880127B6A690F3325F00EC5923C" ID-CLB-Dataset="32119" ID-DOI="10.5852/ejt.2017.293" ID-GBIF-Dataset="3099a544-54cd-48e5-8aac-8e1ca24c0e8b" ID-ISSN="2118-9773" ID-Zenodo-Dep="889857" ID-ZooBank="urn:lsid:zoobank.org:pub:608B2AAB-39A4-494C-9CE0-AA84916008BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505237457348" checkinUser="plazi" docAuthor="Anh D. Nguyen, Zoltán Korsós, Kuem-Hee Jang & Ui-Wook Hwang" docDate="2017" docId="287287F2FFD9332DDF2D1C49FED78126" docLanguage="en" docName="EJT-293_Nguyen.pdf" docOrigin="European Journal of Taxonomy 293" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Oxidus obtusus" docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="D44BFF8AFFD73323DD631D42FF8F8422" masterDocTitle="A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida, Paradoxosomatidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="15" updateTime="1698468380887" updateUser="ExternalLinkService">
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<mods:title id="CBF50C9DAB5EEBAFFF7582884DFFE379">A revision and phylogenetic analysis of the millipede genus Oxidus Cook, 1911 (Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="6E32266D04407D9D888A6E1BC0F25F2B">Anh D. Nguyen</mods:namePart>
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<mods:namePart id="EA2D2C9D6C4B027D7B56042174585561">Zoltán Korsós</mods:namePart>
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<mods:namePart id="4973F6CD428F5EC649BB813983B609A0">Kuem-Hee Jang</mods:namePart>
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<mods:namePart id="3F7C2B2A1E9F4F833034351E8DDF259E">Ui-Wook Hwang</mods:namePart>
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<mods:title id="2298A97E5FD2B56D8BE3544EB6535BFC">European Journal of Taxonomy</mods:title>
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<mods:date id="FC6385FAD64DC5D67ED5796CD3A42575">2017</mods:date>
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<mods:number id="45E1433685887503A1A8E6B8FFA4EFFE">2017-03-07</mods:number>
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<mods:number id="8543991572830F71FA013745318E628F">293</mods:number>
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<treatment id="287287F2FFD9332DDF2D1C49FED78126" ID-DOI="http://doi.org/10.5281/zenodo.5672744" ID-GBIF-Taxon="134034198" ID-Zenodo-Dep="5672744" LSID="urn:lsid:plazi:treatment:287287F2FFD9332DDF2D1C49FED78126" httpUri="http://treatment.plazi.org/id/287287F2FFD9332DDF2D1C49FED78126" lastPageNumber="15" pageId="14" pageNumber="15">
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<subSubSection id="E8C1656FFFD9332DDF2D1C49FC6A8507" box="[590,997,267,294]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="A06436E4FFD9332DDF2D1C49FC6A8507" blockId="14.[590,997,267,329]" box="[590,997,267,294]" pageId="14" pageNumber="15">
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<heading id="FB2C8188FFD9332DDF2D1C49FC6A8507" box="[590,997,267,294]" centered="true" fontSize="11" level="4" pageId="14" pageNumber="15" reason="4">
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<taxonomicName id="67DB4D67FFD9332DDF2D1C49FC6A8507" ID-CoL="4BBXY" authority="(Takakuwa, 1942)" baseAuthorityName="Takakuwa" baseAuthorityYear="1942" box="[590,997,267,294]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obtusus">
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<emphasis id="92AFEAF6FFD9332DDF2D1C49FC868504" bold="true" box="[590,777,267,294]" italics="true" pageId="14" pageNumber="15">Oxidus obtusus</emphasis>
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(
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<bibRefCitation id="C44A4B15FFD9332DDE7B1C49FC528507" author="Takakuwa Y." box="[792,989,267,293]" journalOrPublisher="Transactions of the Natural History Society of Formosa" pageId="14" pageNumber="15" pagination="359 - 367" part="32" refString="Takakuwa Y. 1942 a. Die Myriopoden aus Formosa, Philippinien u. s. w. Transactions of the Natural History Society of Formosa 32 (231): 359 - 367." title="Die Myriopoden aus Formosa, Philippinien u. s. w." type="journal article" year="1942">Takakuwa, 1942</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E8C1656FFFD9332DDF8D1C6CFCCA856B" box="[750,837,302,329]" pageId="14" pageNumber="15" type="description">
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||
<paragraph id="A06436E4FFD9332DDF8D1C6CFCCA856B" blockId="14.[590,997,267,329]" box="[750,837,302,329]" pageId="14" pageNumber="15">
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<figureCitation id="38E02A61FFD9332DDF8D1C6CFCCA856B" box="[750,837,302,329]" captionStart="Fig. 8" captionStartId="13.[189,232,1930,1956]" captionTargetBox="[331,1205,1027,1871]" captionTargetId="figure@13.[189,1399,1003,1915]" captionTargetPageId="13" captionText="Fig. 8. A. Left gonopod of Oxidus avia (Verhoeff, 1937), lateral view. B. Right gonopod of Oxidus obtusus (Takakuwa, 1942), mesal view. Redrawn from Verhoeff 1937 and Takakuwa 1942 a, respectively. No scale bars." httpUri="https://zenodo.org/record/889873/files/figure.png" pageId="14" pageNumber="15">Fig. 8B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E8C1656FFFD9332DDDDE1C32FDB485CD" pageId="14" pageNumber="15" type="reference_group">
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<paragraph id="A06436E4FFD9332DDDDE1C32FCDD85A8" blockId="14.[189,850,368,394]" box="[189,850,368,394]" pageId="14" pageNumber="15">
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<taxonomicName id="67DB4D67FFD9332DDDDE1C32FD4785A8" authority="Takakuwa, 1942 a: 363" authorityName="Takakuwa" authorityPageNumber="363" authorityYear="1942" box="[189,712,368,394]" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obtusa">
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<emphasis id="92AFEAF6FFD9332DDDDE1C32FE3B85A8" box="[189,436,368,394]" italics="true" pageId="14" pageNumber="15">Orthomorpha obtusa</emphasis>
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<bibRefCitation id="C44A4B15FFD9332DDCD81C32FD0185A8" author="Takakuwa Y." box="[443,654,368,394]" journalOrPublisher="Transactions of the Natural History Society of Formosa" pageId="14" pageNumber="15" pagination="359 - 367" part="32" refString="Takakuwa Y. 1942 a. Die Myriopoden aus Formosa, Philippinien u. s. w. Transactions of the Natural History Society of Formosa 32 (231): 359 - 367." title="Die Myriopoden aus Formosa, Philippinien u. s. w." type="journal article" year="1942" yearSuffix="a">Takakuwa, 1942a</bibRefCitation>
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: 363
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</taxonomicName>
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, 367, fg. 7.
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</paragraph>
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<paragraph id="A06436E4FFD9332DDDDE1CF3FDB485CD" blockId="14.[189,1398,433,495]" pageId="14" pageNumber="15">
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<taxonomicName id="67DB4D67FFD9332DDDDE1CF3FEF885E9" ID-CoL="4BBXY" baseAuthorityName="Takakuwa" baseAuthorityYear="1942" box="[189,375,433,459]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obtusus">
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<emphasis id="92AFEAF6FFD9332DDDDE1CF3FEF885E9" box="[189,375,433,459]" italics="true" pageId="14" pageNumber="15">Oxidus obtusus</emphasis>
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</taxonomicName>
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–
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<bibRefCitation id="C44A4B15FFD9332DDCFC1CF3FD6785EE" author="Chamberlin R. & Wang Y. M." box="[415,744,433,460]" journalOrPublisher="American Museum Novitates" pageId="14" pageNumber="15" pagination="1 - 13" part="1621" refString="Chamberlin R. V. & Wang Y. M. 1953. Records of millipeds (Diplopoda) from Japan and other Oriental areas, with descriptions of new genera and species. American Museum Novitates 1621: 1 - 13." title="Records of millipeds (Diplopoda) from Japan and other Oriental areas, with descriptions of new genera and species" type="journal article" year="1953">Chamberlin & Wang 1953</bibRefCitation>
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: 6. —
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<bibRefCitation id="C44A4B15FFD9332DDE261CF3FB8685EE" author="Takakuwa Y." box="[837,1033,433,460]" journalOrPublisher="Japan Society for the Promotion of Science, Tokyo" pageId="14" pageNumber="15" refString="Takakuwa Y. 1954. Diplopoden aus Japan und ihn angrenzenden Gebieten. Japan Society for the Promotion of Science, Tokyo." title="Diplopoden aus Japan und ihn angrenzenden Gebieten" type="book" year="1954">Takakuwa 1954</bibRefCitation>
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: 37. —
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<bibRefCitation id="C44A4B15FFD9332DD9161CF3FA9885E9" author="Jeekel C. A. W." box="[1141,1303,433,459]" journalOrPublisher="Studies on the Fauna of Suriname and other Guyanas" pageId="14" pageNumber="15" pagination="1 - 157" part="4" refString="Jeekel C. A. W. 1963 a. Diplopoda of Guiana. Studies on the Fauna of Suriname and other Guyanas 4 (11): 1 - 157." title="Diplopoda of Guiana" type="journal article" year="1963" yearSuffix="a">Jeekel 1963a</bibRefCitation>
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: 20. —
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<bibRefCitation id="C44A4B15FFD9332DDDDE1C97FE6185CD" author="Nguyen A. D. & Sierwald P." box="[189,494,469,495]" journalOrPublisher="CheckList" pageId="14" pageNumber="15" pagination="1132 - 1353" part="9" refString="Nguyen A. D. & Sierwald P. 2013. A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). CheckList 9: 1132 - 1353." title="A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida)" type="journal article" year="2013">Nguyen & Sierwald 2013</bibRefCitation>
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: 1295.
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</paragraph>
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</subSubSection>
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<subSubSection id="E8C1656FFFD9332DDDDE1F54FAF88687" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="A06436E4FFD9332DDDDE1F54FEB98612" blockId="14.[189,1399,534,678]" box="[189,310,534,560]" pageId="14" pageNumber="15">
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<heading id="FB2C8188FFD9332DDDDE1F54FEB98612" bold="true" box="[189,310,534,560]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="6">
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<emphasis id="92AFEAF6FFD9332DDDDE1F54FEB98612" bold="true" box="[189,310,534,560]" pageId="14" pageNumber="15">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="A06436E4FFD9332DDDDE1F06FAF88687" blockId="14.[189,1399,534,678]" pageId="14" pageNumber="15">
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This species can be distinguished by its gonopod femorite being short, very broad gently distad; process
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<emphasis id="92AFEAF6FFD9332DD80B1F07FAF8867D" box="[1384,1399,581,607]" italics="true" pageId="14" pageNumber="15">h</emphasis>
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bent upward at tip, like a lamellar sickle; spine
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<emphasis id="92AFEAF6FFD9332DDF981F2BFC8986A0" box="[763,774,617,642]" italics="true" pageId="14" pageNumber="15">z</emphasis>
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small, obtuse; solenophore large, broad and simple, especially wide at its base; mesal lobule not separated from rounded solenophore tip (
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<bibRefCitation id="C44A4B15FFD9332DD9FE1FC9FAE58687" author="Takakuwa Y." box="[1181,1386,651,677]" journalOrPublisher="Transactions of the Natural History Society of Formosa" pageId="14" pageNumber="15" pagination="359 - 367" part="32" refString="Takakuwa Y. 1942 a. Die Myriopoden aus Formosa, Philippinien u. s. w. Transactions of the Natural History Society of Formosa 32 (231): 359 - 367." title="Die Myriopoden aus Formosa, Philippinien u. s. w." type="journal article" year="1942" yearSuffix="a">Takakuwa 1942a</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="E8C1656FFFD9332DDDDE1F8FFDAF80B1" pageId="14" pageNumber="15" type="discussion">
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<paragraph id="A06436E4FFD9332DDDDE1F8FFEA086C5" blockId="14.[189,303,717,743]" box="[189,303,717,743]" pageId="14" pageNumber="15">
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<heading id="FB2C8188FFD9332DDDDE1F8FFEA086C5" bold="true" box="[189,303,717,743]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="2">
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<emphasis id="92AFEAF6FFD9332DDDDE1F8FFEA086C5" bold="true" box="[189,303,717,743]" pageId="14" pageNumber="15">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="A06436E4FFD9332DDDDE1FB9FD6F8780" blockId="14.[189,1399,763,930]" pageId="14" pageNumber="15">
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This species was described from South Korea (
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<bibRefCitation id="C44A4B15FFD9332DDF901FB9FC4B8737" author="Takakuwa Y." box="[755,964,763,789]" journalOrPublisher="Transactions of the Natural History Society of Formosa" pageId="14" pageNumber="15" pagination="359 - 367" part="32" refString="Takakuwa Y. 1942 a. Die Myriopoden aus Formosa, Philippinien u. s. w. Transactions of the Natural History Society of Formosa 32 (231): 359 - 367." title="Die Myriopoden aus Formosa, Philippinien u. s. w." type="journal article" year="1942" yearSuffix="a">Takakuwa 1942a</bibRefCitation>
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). Later,
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<bibRefCitation id="C44A4B15FFD9332DD9441FB9FAF88734" author="Chamberlin R. & Wang Y. M." box="[1063,1399,763,790]" journalOrPublisher="American Museum Novitates" pageId="14" pageNumber="15" pagination="1 - 13" part="1621" refString="Chamberlin R. V. & Wang Y. M. 1953. Records of millipeds (Diplopoda) from Japan and other Oriental areas, with descriptions of new genera and species. American Museum Novitates 1621: 1 - 13." title="Records of millipeds (Diplopoda) from Japan and other Oriental areas, with descriptions of new genera and species" type="journal article" year="1953">Chamberlin & Wang (1953)</bibRefCitation>
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reported two ♀♀ in the collection of the American Natural History Museum (New York) from Japan. It is possible that the locality was mislabelled because they stated that those specimens were collected and deposited by Dr. Takakuwa Y. Thus, it is believed that these two ♀♀ were collected from the same locality as the species holotype (South Korea).
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</paragraph>
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<paragraph id="A06436E4FFD9332DDDDE1E8BFDAF80B1" blockId="14.[189,1399,969,1171]" pageId="14" pageNumber="15">
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<bibRefCitation id="C44A4B15FFD9332DDDDE1E8BFEC987C1" author="Lim K. Y." box="[189,326,969,995]" journalOrPublisher="PhD dissertation, Chonbuk National University, Jeonju, Korea" pageId="14" pageNumber="15" refString="Lim K. Y. 2001. Taxonomy of Millipedes (Arthropoda: Diplopoda) in Korea. PhD dissertation, Chonbuk National University, Jeonju, Korea." title="Taxonomy of Millipedes (Arthropoda: Diplopoda) in Korea" type="book" year="2001">Lim (2001)</bibRefCitation>
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synonymized
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<taxonomicName id="67DB4D67FFD9332DDC9B1E88FDF687C1" baseAuthorityName="Takakuwa" baseAuthorityYear="1942" box="[504,633,969,996]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obtusus">
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<emphasis id="92AFEAF6FFD9332DDC9B1E88FDF687C1" box="[504,633,969,996]" italics="true" pageId="14" pageNumber="15">O. obtusus</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="67DB4D67FFD9332DDFA11E88FCCC87C1" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[706,835,969,996]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis id="92AFEAF6FFD9332DDFA11E88FCCC87C1" box="[706,835,969,996]" italics="true" pageId="14" pageNumber="15">O. gracilis</emphasis>
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</taxonomicName>
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. He argued that the two species differ only in the number of postfemoral branches, three in
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<taxonomicName id="67DB4D67FFD9332DDF811EAFFCED8024" baseAuthorityName="Takakuwa" baseAuthorityYear="1942" box="[738,866,1004,1031]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obtusus">
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<emphasis id="92AFEAF6FFD9332DDF811EAFFCED8024" box="[738,866,1004,1031]" italics="true" pageId="14" pageNumber="15">O. obtusus</emphasis>
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</taxonomicName>
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and four in
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<taxonomicName id="67DB4D67FFD9332DDE9A1EAFFBF68024" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[1017,1145,1004,1031]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis id="92AFEAF6FFD9332DDE9A1EAFFBF68024" box="[1017,1145,1004,1031]" italics="true" pageId="14" pageNumber="15">O. gracilis</emphasis>
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</taxonomicName>
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; in his opinion, such a difference is minor and cannot be used to separate two species. With the two species
|
||
<taxonomicName id="67DB4D67FFD9332DD9A61952FACB8008" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[1221,1348,1040,1066]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="92AFEAF6FFD9332DD9A61952FACB8008" box="[1221,1348,1040,1066]" italics="true" pageId="14" pageNumber="15">O. gracilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="67DB4D67FFD9332DDDDE1971FEAA806F" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[189,293,1075,1101]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="92AFEAF6FFD9332DDDDE1971FEAA806F" box="[189,293,1075,1101]" italics="true" pageId="14" pageNumber="15">O. gigas</emphasis>
|
||
</taxonomicName>
|
||
, those characters are important for species delimitation. It is recommended, therefore, that
|
||
<taxonomicName id="67DB4D67FFD9332DDDDE1914FEFE8052" baseAuthorityName="Takakuwa" baseAuthorityYear="1942" box="[189,369,1110,1136]" class="Diplopoda" family="Paradoxosomatidae" genus="Oxidus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obtusus">
|
||
<emphasis id="92AFEAF6FFD9332DDDDE1914FEFE8052" box="[189,369,1110,1136]" italics="true" pageId="14" pageNumber="15">Oxidus obtusus</emphasis>
|
||
</taxonomicName>
|
||
should be considered a valid species. Examination of further fresh material is needed to confrm that recommendation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E8C1656FFFD9332DDDDE19F8FED78126" box="[189,344,1210,1284]" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="A06436E4FFD9332DDDDE19F8FED980F6" blockId="14.[189,342,1210,1236]" box="[189,342,1210,1236]" pageId="14" pageNumber="15">
|
||
<heading id="FB2C8188FFD9332DDDDE19F8FED980F6" bold="true" box="[189,342,1210,1236]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="2">
|
||
<emphasis id="92AFEAF6FFD9332DDDDE19F8FED980F6" bold="true" box="[189,342,1210,1236]" pageId="14" pageNumber="15">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A06436E4FFD9332DDDDE19ABFED78126" blockId="14.[189,344,1257,1284]" box="[189,344,1257,1284]" pageId="14" pageNumber="15">South Korea.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |