179 lines
18 KiB
XML
179 lines
18 KiB
XML
<document id="A3DDAAF0A547A008367E391A0FF0319F" ID-DOI="10.5281/zenodo.276598" ID-GBIF-Dataset="db749b2d-e601-4558-beb0-4d7414b7350a" ID-ISSN="1175-5326" ID-Zenodo-Dep="276598" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460224332844" checkinUser="plazi" docAuthor="Shattuck, Steven O." docDate="2011" docId="286F87FD26109F2B528CFD1EFA164235" docLanguage="en" docName="zt02743p026.pdf" docOrigin="Zootaxa 2743" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Calyptomyrmex ryderae Shattuck, 2011, sp. n." docType="treatment" docVersion="11" lastPageNumber="21" masterDocId="D456FF8526039F3F521BFFC9FF8A466D" masterDocTitle="Revision of the ant genus Calyptomyrmex (Hymenoptera: Formicidae) in South-east Asia and Oceania" masterLastPageNumber="26" masterPageNumber="1" pageNumber="20" updateTime="1698238071186" updateUser="plazi">
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<mods:titleInfo id="6BF261F425C496F0F0528850868D05C1">
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<mods:title id="64BF77CFFF0E814EF0B048020D8AE2E1">Revision of the ant genus Calyptomyrmex (Hymenoptera: Formicidae) in South-east Asia and Oceania</mods:title>
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</mods:titleInfo>
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<mods:name id="977F5B7121C7AC5FEA225AA1F44489BB" type="personal">
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<mods:roleTerm id="7CAFECBF5ED3AFA11F14292CB686D352">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F59A21257085B42F1F400CACD16F74EE">Shattuck, Steven O.</mods:namePart>
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<mods:typeOfResource id="09DA1F35E101E6486B7DCE56B374ADD3">text</mods:typeOfResource>
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<mods:title id="0D6D58E0886BD325989B61269867DB32">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CADA3CEB4B6E51AFECA233B267F0AB92">
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<mods:date id="93DA6838AF529ADB4B913A78F0E3D6FF">2011</mods:date>
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<mods:detail id="67D9DB66EE07E5759F349C822587A5A0" type="volume">
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<mods:number id="232DDF89C302EA5D72126099E0915D8C">2743</mods:number>
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<mods:start id="5267D147E93F17F53331BA815B121063">1</mods:start>
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<mods:classification id="4F62EED11642E1A12119A8FF4E2F7E53">journal article</mods:classification>
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<mods:identifier id="F8534B0D5F946B66E871D72EAFB3DB71" type="DOI">10.5281/zenodo.276598</mods:identifier>
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<mods:identifier id="D783D3710A2F4AC3D824B5557D6ED4E0" type="GBIF-Dataset">db749b2d-e601-4558-beb0-4d7414b7350a</mods:identifier>
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<mods:identifier id="6D7452091809DF6367A00145BD487E13" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="02C15526DAFF7F31C17A5DD3BD6C0955" type="Zenodo-Dep">276598</mods:identifier>
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</mods:mods>
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<treatment id="286F87FD26109F2B528CFD1EFA164235" ID-DOI="http://doi.org/10.5281/zenodo.6188495" ID-GBIF-Taxon="119385931" ID-Zenodo-Dep="6188495" LSID="urn:lsid:plazi:treatment:286F87FD26109F2B528CFD1EFA164235" httpUri="http://treatment.plazi.org/id/286F87FD26109F2B528CFD1EFA164235" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
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<subSubSection id="E8DC656026109F2C528CFD1EFEAD457F" pageId="19" pageNumber="20" type="nomenclature">
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||
<paragraph id="A07936EB26109F2C528CFD1EFD8B449C" blockId="19.[151,513,727,786]" box="[151,513,727,753]" pageId="19" pageNumber="20">
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||
<heading id="FB31818726109F2C528CFD1EFD8B449C" bold="true" box="[151,513,727,753]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
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<emphasis id="92B2EAF926109F2C528CFD1EFD8B449C" bold="true" box="[151,513,727,753]" pageId="19" pageNumber="20">
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<taxonomicName id="67C64D6826109F2C528CFD1EFE32449C" box="[151,440,727,753]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ryderae" status="sp. nov.">
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<emphasis id="92B2EAF926109F2C528CFD1EFE32449C" bold="true" box="[151,440,727,753]" italics="true" pageId="19" pageNumber="20">Calyptomyrmex ryderae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8981578226109F2C53A4FD1EFD8B449C" box="[447,513,727,753]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="A07936EB26109F2C528CFD33FEAD457F" blockId="19.[151,513,727,786]" box="[151,295,762,786]" pageId="19" pageNumber="20">
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(
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<figureCitation id="38FD2A6E26109F2C5284FD33FF7F457F" box="[159,245,762,786]" captionStart="FIGURE 14" captionStartId="19.[151,250,1963,1986]" captionTargetBox="[151,1436,842,1941]" captionTargetId="figure@19.[151,1436,842,1942]" captionTargetPageId="19" captionText="FIGURE 14. Calyptomyrmex ryderae (holotype): A. Front of head; B. Side of body; C. Top of body; D. Distribution of material examined." httpUri="https://zenodo.org/record/276612/files/figure.png" pageId="19" pageNumber="20">Figs 14</figureCitation>
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,
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<figureCitation id="38FD2A6E26109F2C531AFD33FE94457F" box="[257,286,762,786]" captionStart="FIGURE 18" captionStartId="24.[151,250,1605,1628]" captionTargetBox="[175,1421,749,1574]" captionTargetId="figure@24.[151,1436,735,1583]" captionTargetPageId="24" captionText="FIGURE 18. Head length versus head width for C. asper, C. beccarii, C. danum, C. fritillus, C. loweryi, C. ryderae, C. sparsus and C. taylori." httpUri="https://zenodo.org/record/276616/files/figure.png" pageId="19" pageNumber="20">18</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<caption id="F4B9666326109F2C528CF862FEA34188" httpUri="https://zenodo.org/record/276612/files/figure.png" pageId="19" pageNumber="20" targetBox="[151,1436,842,1941]" targetPageId="19">
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<paragraph id="A07936EB26109F2C528CF862FEA34188" blockId="19.[151,1436,1963,2021]" pageId="19" pageNumber="20">
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<emphasis id="92B2EAF926109F2C528CF862FEAA41AC" bold="true" box="[151,288,1963,1986]" pageId="19" pageNumber="20">FIGURE 14.</emphasis>
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<taxonomicName id="67C64D6826109F2C533DF865FD9141AF" box="[294,539,1964,1986]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ryderae">
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<emphasis id="92B2EAF926109F2C533DF865FD9141AF" box="[294,539,1964,1986]" italics="true" pageId="19" pageNumber="20">Calyptomyrmex ryderae</emphasis>
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</taxonomicName>
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(holotype): A. Front of head; B. Side of body; C. Top of body; D. Distribution of material examined.
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</paragraph>
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</caption>
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<subSubSection id="E8DC656026179F2B528CFF51FD154709" pageId="20" pageNumber="21" type="description">
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<paragraph id="A07936EB26179F2B528CFF51FD154709" blockId="20.[151,1437,151,1112]" pageId="20" pageNumber="21">
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<emphasis id="92B2EAF926179F2B528CFF51FF6B46DD" bold="true" box="[151,225,152,176]" pageId="20" pageNumber="21">
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<typeStatus id="7F7D884926179F2B528CFF51FF5046DD" box="[151,218,152,176]" pageId="20" pageNumber="21">Types</typeStatus>
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.
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</emphasis>
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<typeStatus id="7F7D884926179F2B52F7FF5EFEDE46C2" box="[236,340,151,175]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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worker from G. Mulu National Park, RGS Expedition, Long
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<collectingCountry id="D8D1767B26179F2B5615FF5EFBCB46C2" box="[1038,1089,151,175]" name="Palau" pageId="20" pageNumber="21">Pala</collectingCountry>
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, 4th Dist.,
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<collectingCountry id="D8D1767B26179F2B56A4FF5EFAA846C2" box="[1215,1314,151,175]" name="Malaysia" pageId="20" pageNumber="21">Sarawak</collectingCountry>
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,
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<collectingCountry id="D8D1767B26179F2B5735FF5EFA1D46C2" box="[1326,1431,151,175]" name="Malaysia" pageId="20" pageNumber="21">Malaysia</collectingCountry>
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,
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<date id="D478102B26179F2B528CFF75FEDD46B9" box="[151,343,188,212]" pageId="20" pageNumber="21" value="1977-10-21">21 October 1977</date>
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, B. Bolton, lowland rainforest, leaf litter (BMNH).
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<typeStatus id="7F7D884926179F2B5187FF74FB8746B8" box="[924,1037,189,213]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="B6C0FD6226179F2B5603FF75FB1346B9" box="[1048,1177,188,212]" pageId="20" pageNumber="21" type="worker">21 workers</specimenCount>
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, same data as
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<typeStatus id="7F7D884926179F2B5721FF75FA1646B9" box="[1338,1436,188,212]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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except
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<specimenCount id="B6C0FD6226179F2B52FCFF16FEDD469A" box="[231,343,223,247]" pageId="20" pageNumber="21" type="worker">5 workers</specimenCount>
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collected
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<date id="D478102B26179F2B53D7FF16FDC1469A" box="[460,587,223,247]" pageId="20" pageNumber="21" value="1921-10">21 October</date>
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,
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<specimenCount id="B6C0FD6226179F2B504CFF16FD59469A" box="[599,723,223,247]" pageId="20" pageNumber="21" type="worker">11 workers</specimenCount>
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collected
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<date id="D478102B26179F2B5153FF16FC4D469A" box="[840,967,223,247]" pageId="20" pageNumber="21" value="1916-10">16 October</date>
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,
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<specimenCount id="B6C0FD6226179F2B51C8FF16FBB2469A" box="[979,1080,223,247]" pageId="20" pageNumber="21" type="worker">1 worker</specimenCount>
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collected
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<date id="D478102B26179F2B56B7FF16FAA1469A" box="[1196,1323,223,247]" pageId="20" pageNumber="21" value="1918-10">18 October</date>
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,
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<specimenCount id="B6C0FD6226179F2B572CFF16FA17469A" box="[1335,1437,223,247]" pageId="20" pageNumber="21" type="worker">1 worker</specimenCount>
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collected
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<date id="D478102B26179F2B5319FECDFE144771" box="[258,414,260,284]" pageId="20" pageNumber="21" value="1928-09">28 September</date>
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,
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<specimenCount id="B6C0FD6226179F2B53B3FECDFD814771" box="[424,523,260,284]" pageId="20" pageNumber="21" type="worker">1 worker</specimenCount>
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collected
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<date id="D478102B26179F2B5066FECDFC924771" box="[637,792,260,284]" pageId="20" pageNumber="21" value="1920-09">20 September</date>
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(soil on rock),
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<specimenCount id="B6C0FD6226179F2B51DAFECDFBAE4771" box="[961,1060,260,284]" pageId="20" pageNumber="21" type="worker">1 worker</specimenCount>
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collected
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<date id="D478102B26179F2B568DFECDFA8E4771" box="[1174,1284,260,284]" pageId="20" pageNumber="21">1 October</date>
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(leaf litter on log);
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<specimenCount id="B6C0FD6226179F2B52C9FEEEFECB4752" box="[210,321,295,319]" pageId="20" pageNumber="21" type="worker">6 workers</specimenCount>
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from Gn. Mulu National Park, 4th Dist.,
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<collectingCountry id="D8D1767B26179F2B5115FEEEFCF84752" box="[782,882,295,319]" name="Malaysia" pageId="20" pageNumber="21">Sarawak</collectingCountry>
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,
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<collectingCountry id="D8D1767B26179F2B5167FEEEFC6C4752" box="[892,998,295,319]" name="Malaysia" pageId="20" pageNumber="21">Malaysia</collectingCountry>
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,
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<date id="D478102B26179F2B51EAFEE1FB4F4752" box="[1009,1221,295,320]" pageId="20" pageNumber="21" value="1978-06" valueMax="1978-08-31" valueMin="1978-06-00">June–August 1978</date>
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, P.M. Hammond & J.E.
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<collectingCountry id="D8D1767B26179F2B52D3FE85FEA44709" box="[200,302,332,356]" name="Marshall Islands" pageId="20" pageNumber="21">Marshall</collectingCountry>
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, B.M. 1978-49 (ANIC, BMNH).
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</paragraph>
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</subSubSection>
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<subSubSection id="E8DC656026179F2B52DCFEA6FA9144E9" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="A07936EB26179F2B52DCFEA6FA9144E9" blockId="20.[151,1437,151,1112]" pageId="20" pageNumber="21">
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<emphasis id="92B2EAF926179F2B52DCFEA6FECB47E5" bold="true" box="[199,321,367,392]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
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Hairs on head and body thin (essentially the same width along entire length and with blunt tips), gaster weakly punctate, propodeum lacking or essentially lacking spines or angles, petiolar node in dorsal view noticeably wider than long (PetI> 130) and body smaller (HW <
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<quantity id="673E9B0E26179F2B519DFE7EFC6D47A2" box="[902,999,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="21" unit="mm" value="1.05">1.05mm</quantity>
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). This species is most similar to
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<taxonomicName id="67C64D6826179F2B5745FE70FA1647BD" box="[1374,1436,441,464]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="asper">
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<emphasis id="92B2EAF926179F2B5745FE70FA1647BD" box="[1374,1436,441,464]" italics="true" pageId="20" pageNumber="21">asper</emphasis>
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</taxonomicName>
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but differs in having the lateral surfaces of mesosoma and the dorsal surface of head with more numerous, narrower and more closely spaced rugae (the rugae are less numerous, broader and more widely spaced in
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<taxonomicName id="67C64D6826179F2B56FFFDC8FAA84475" box="[1252,1314,513,536]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="asper">
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<emphasis id="92B2EAF926179F2B56FFFDC8FAA84475" box="[1252,1314,513,536]" italics="true" pageId="20" pageNumber="21">asper</emphasis>
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</taxonomicName>
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), the hairs on the mesonotum are longer and thinner (they are shorter and tending to be weakly spatulate in
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<taxonomicName id="67C64D6826179F2B56D3FDECFA8C4451" box="[1224,1286,549,572]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="asper">
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<emphasis id="92B2EAF926179F2B56D3FDECFA8C4451" box="[1224,1286,549,572]" italics="true" pageId="20" pageNumber="21">asper</emphasis>
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</taxonomicName>
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), the petiolar node in dorsal view noticeably wider than long (PetI> 130) while it is only slightly wider than long (PetI approx. 115) in
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<taxonomicName id="67C64D6826179F2B52F6FDA4FEA044E9" box="[237,298,621,644]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="asper">
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<emphasis id="92B2EAF926179F2B52F6FDA4FEA044E9" box="[237,298,621,644]" italics="true" pageId="20" pageNumber="21">asper</emphasis>
|
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</taxonomicName>
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, and the body is smaller (HW <
|
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<quantity id="673E9B0E26179F2B5083FDA5FD7D44E9" box="[664,759,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="21" unit="mm" value="1.05">1.05mm</quantity>
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) in
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<taxonomicName id="67C64D6826179F2B5139FDA5FCF344E9" box="[802,889,620,644]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ryderae">
|
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<emphasis id="92B2EAF926179F2B5139FDA5FCF344E9" box="[802,889,620,644]" italics="true" pageId="20" pageNumber="21">ryderae</emphasis>
|
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</taxonomicName>
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compared to
|
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<taxonomicName id="67C64D6826179F2B5609FDA4FBDA44E9" box="[1042,1104,621,644]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="asper">
|
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<emphasis id="92B2EAF926179F2B5609FDA4FBDA44E9" box="[1042,1104,621,644]" italics="true" pageId="20" pageNumber="21">asper</emphasis>
|
||
</taxonomicName>
|
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(HW>
|
||
<quantity id="673E9B0E26179F2B56B6FDA5FA8444E9" box="[1197,1294,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="21" unit="mm" value="1.05">1.05mm</quantity>
|
||
).
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</paragraph>
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</subSubSection>
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<subSubSection id="E8DC656026179F2B52DCFD59FAEE4262" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="A07936EB26179F2B52DCFD59FD8545AA" blockId="20.[151,1437,151,1112]" pageId="20" pageNumber="21">
|
||
<emphasis id="92B2EAF926179F2B52DCFD59FE3144C5" bold="true" box="[199,443,655,680]" pageId="20" pageNumber="21">Worker description.</emphasis>
|
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Mandibles striate along entire length. Eyes with 5–6 ommatidia in greatest diameter. Propodeum in lateral view with spines reduced to broad angles and essentially absent. Propodeal lobes uniform in thickness. Node of petiole in profile higher than and approximately the same size as postpetiole. In dorsal view the petiolar node slightly narrower than the postpetiolar node. Head indistinctly rugose, the rugae forming an indistinct network, with the underlying surface finely but distinctly punctate. Mesosoma and petiolar and postpetiolar nodes with irregular, wavy rugae superimposed over an indistinctly punctate background. Gaster finely and indistinctly sculptured and with a matte appearance. Hairs on head and body erect, uniform in diameter and with sharp or blunt tips. Clypeal fork with thin hairs only, the apical pair slightly thinner than the lateral pair. Colour red-brown, the antennae and legs slightly lighter.
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</paragraph>
|
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<paragraph id="A07936EB26179F2B52DCFC1DFAEE4262" blockId="20.[151,1437,151,1112]" pageId="20" pageNumber="21">
|
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<emphasis id="92B2EAF926179F2B52DCFC1DFEF74581" bold="true" box="[199,381,980,1004]" pageId="20" pageNumber="21">Measurements.</emphasis>
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Worker (n = 5) - CFW 0.14–0.15; CI 112–115: HL 0.79–0.89; HW 0.89–1.01; ML 0.87–1.00; MTL 0.38–0.45; PetI 134–155; PetL 0.19–0.23; PetW 0.30–0.34; PronW 0.56–0.63; SI 50–52; SL 0.45–0.51.
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</paragraph>
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</subSubSection>
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<subSubSection id="E8DC656026179F2B52DCFBD2FA164235" pageId="20" pageNumber="21" type="discussion">
|
||
<paragraph id="A07936EB26179F2B52DCFBD2FA164235" blockId="20.[151,1437,151,1112]" pageId="20" pageNumber="21">
|
||
<emphasis id="92B2EAF926179F2B52DCFBD2FEC54259" bold="true" box="[199,335,1051,1076]" pageId="20" pageNumber="21">Comments.</emphasis>
|
||
<taxonomicName id="67C64D6826179F2B534CFBD5FDED4259" box="[343,615,1052,1076]" class="Insecta" family="Formicidae" genus="Calyptomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ryderae">
|
||
<emphasis id="92B2EAF926179F2B534CFBD5FDED4259" box="[343,615,1052,1076]" italics="true" pageId="20" pageNumber="21">Calyptomyrmex ryderae</emphasis>
|
||
</taxonomicName>
|
||
is known from a handful of collections from a single lowland rainforest area of
|
||
<collectingCountry id="D8D1767B26179F2B52F0FB89FEC54235" box="[235,335,1088,1112]" name="Malaysia" pageId="20" pageNumber="21">Sarawak</collectingCountry>
|
||
. Most collections have been from leaf litter samples while one worker was found in soil on a rock.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |