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<document id="DDCD18C209370C1DB8C8344D95336EA4" ID-CLB-Dataset="270321" ID-DOI="10.5252/zoosystema2023v45a17" ID-GBIF-Dataset="dd6aa488-f2ce-4641-9392-5c428430fc96" ID-ISSN="1638-9387" ID-Zenodo-Dep="8382549" ID-ZooBank="C92EEB3F-2639-4891-958E-B8CE87EB74F1" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1695803671856" checkinUser="plazi" docAuthor="Signorotto, Fiorella, Mancini, Mariana &amp; Ferretti, Nelson" docDate="2023" docId="244487DDD712C165FF18449CFA89F803" docLanguage="en" docName="zoosystema.2023.45.17.pdf" docOrigin="Zoosystema 45 (17)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2023v45a17_0.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Acanthogonatus messii Signorotto &amp; Mancini &amp; Ferretti 2023, n. sp." docType="treatment" docUuid="F7886060-729F-4F17-A16B-565DB20641DA" docUuidSource="ZooBank" docVersion="6" lastPageNumber="507" masterDocId="D87DFFA5D714C16FFFAD414EFFC9FFE8" masterDocTitle="A new small Acanthogonatus Karsch, 1880 (Mygalomorphae, Pycnothelidae) species from Argentinean Patagonia: description of A. messii Signorotto &amp; Ferretti n. sp. and its phylogenetic placement" masterLastPageNumber="512" masterPageNumber="499" pageNumber="503" updateTime="1702002861377" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F06F235CBD92C7896BCDAA0C53A0ECA9">A new small Acanthogonatus Karsch, 1880 (Mygalomorphae, Pycnothelidae) species from Argentinean Patagonia: description of A. messii Signorotto &amp; Ferretti n. sp. and its phylogenetic placement</mods:title>
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<mods:namePart id="D219C81222D21C952311226681F41244">Signorotto, Fiorella</mods:namePart>
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<mods:namePart id="7115DC38AABC87BCA9D96DD88E851187">Mancini, Mariana</mods:namePart>
<mods:affiliation id="8C9EC5D1EFAAB30CD6CF6A66CC55AB5A">Departamento de Biología, Bioquímica y Farmacia, Universidad Nacional del Sur, San Juan 670, Bahía Blanca (8000), Buenos Aires (Argentina)</mods:affiliation>
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<mods:affiliation id="9EC810BE643038EB23C552EAA6AA489A">Departamento de Biología, Bioquímica y Farmacia, Universidad Nacional del Sur, San Juan 670, Bahía Blanca (8000), Buenos Aires (Argentina) and Centro de Recursos Naturales Renovables de la Zona Semiárida (CERZOS-UNS, CONICET), Camino La Carrindanga Km. 7, Bahía Blanca (8000), Buenos Aires (Argentina) nferretti @ conicet. gov. ar (corresponding author)</mods:affiliation>
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<mods:date id="2EC4F4BDE804E0AA7E1090E902F44A3E">2023</mods:date>
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<taxonomicName id="6BED4D48D712C169FF18449CFD5CFA05" authority="Signorotto &amp; Ferretti" authorityName="Signorotto &amp; Mancini &amp; Ferretti" authorityYear="2023" box="[181,661,1490,1517]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="503" phylum="Arthropoda" rank="species" species="messii" status="sp. nov.">
<emphasis id="9E99EAD9D712C169FF18449CFE6EFA05" bold="true" box="[181,423,1490,1517]" italics="true" pageId="6" pageNumber="503">Acanthogonatus messii</emphasis>
Signorotto &amp; Ferretti
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(
<figureCitation id="34D62A4ED712C169FE3544BDFE38F9E5" box="[408,497,1523,1549]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStartId-0="6.[133,144,1174,1192]" captionStartId-1="7.[132,143,1632,1650]" captionStartId-2="8.[132,143,1286,1304]" captionStartId-3="9.[132,143,1986,2004]" captionStartId-4="10.[202,213,1104,1122]" captionTargetBox-0="[135,1453,223,1124]" captionTargetBox-1="[132,1455,215,1576]" captionTargetBox-2="[163,1417,256,1244]" captionTargetBox-3="[134,1455,211,1944]" captionTargetBox-4="[132,1456,196,1069]" captionTargetId-0="figure-463@6.[132,1455,213,1124]" captionTargetId-1="figure-177@7.[132,1455,215,1608]" captionTargetId-2="figure-347@8.[132,1455,215,1244]" captionTargetId-3="figure-15@9.[134,1455,211,1960]" captionTargetId-4="figure-509@10.[132,1455,213,1062]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="FIG. 2. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., habitus: A, female molted in laboratory; B, female collected and photographed from field; C, male molted in laboratory; D, male collected walking in the field. Scale bars: 1 cm." captionText-1="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." captionText-2="FIG. 4. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A-D, palpal bulb. A, prolateral view; B, dorsal view (green arrow indicates the keel); C, retrolateral view (green arrow indicates the keel); D, dorsal view; E, palp, prolateral view; F, palp, retrolateral view. Abbreviations: Cy, cymbium; Ti, tibia. Scale bars: 1 mm." captionText-3="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." captionText-4="FIG. 6. — A, habitat at type locality of Acanthogonatus messii Signorotto &amp; Ferretti n. sp.; B, C, female burrows opened at night at type locality." figureDoi-0="http://doi.org/10.5281/zenodo.8382555" figureDoi-1="http://doi.org/10.5281/zenodo.8382557" figureDoi-2="http://doi.org/10.5281/zenodo.8382559" figureDoi-3="http://doi.org/10.5281/zenodo.8382561" figureDoi-4="http://doi.org/10.5281/zenodo.8382564" httpUri-0="https://zenodo.org/record/8382555/files/figure.png" httpUri-1="https://zenodo.org/record/8382557/files/figure.png" httpUri-2="https://zenodo.org/record/8382559/files/figure.png" httpUri-3="https://zenodo.org/record/8382561/files/figure.png" httpUri-4="https://zenodo.org/record/8382564/files/figure.png" pageId="6" pageNumber="503">Figs 2-6</figureCitation>
)
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urn:lsid:zoobank.org:act:
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. —
<materialsCitation id="1C853C96D712C169FEEF4729FD93F95C" ID-GBIF-Occurrence="4418721303" collectingDate="2018-11-24" collectionCode="UNS-M" collectorName="N. Ferretti" country="Argentina" latitude="-45.84833" location="near Rada Tilly" longLatPrecision="18" longitude="-67.79444" municipality="Escalante" pageId="6" pageNumber="503" specimenCode="UNS-M0658" specimenCount="1" specimenCount-male="1" stateProvince="Chubut" typeStatus="holotype">
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<collectingCountry id="D4FA765BD712C169FE014727FDC6F969" box="[428,527,1641,1665]" name="Argentina" pageId="6" pageNumber="503">Argentina</collectingCountry>
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;
<collectingRegion id="6E29F829D712C169FDFE4729FD6AF997" box="[595,675,1639,1663]" country="Argentina" name="Chubut" pageId="6" pageNumber="503">Chubut</collectingRegion>
,
<collectingMunicipality id="4C36ACB1D712C169FD014729FCCEF997" box="[684,775,1639,1663]" country="Argentina" latitude="-45.84833" longLatPrecision="18" longitude="-67.79444" name="Escalante" pageId="6" pageNumber="503" stateProvince="Chubut">Escalante</collectingMunicipality>
(
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);
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,
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;
<date id="D853100BD712C169FD8447CCFD53F972" box="[553,666,1666,1690]" pageId="6" pageNumber="503" value="2018-11-24">
<collectingDate id="C817E9E3D712C169FD8447CCFD53F972" box="[553,666,1666,1690]" pageId="6" pageNumber="503" value="2018-11-24">24.XI.2018</collectingDate>
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;
<collectorName id="0118531DD712C169FD0E47CCFCCEF972" box="[675,775,1666,1690]" pageId="6" pageNumber="503">N. Ferretti</collectorName>
leg.; found walking during night;
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.
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<typeStatus id="73568869D712C169FF2947F9FF2AF927" box="[132,227,1719,1743]" pageId="6" pageNumber="503" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<materialsCitation id="1C853C96D712C169FF4247F6FE3CF901" ID-GBIF-Occurrence="4418721304" collectingDate="2018-11-24" collectionCode="TYPE, MATERIAL" collectorName="N. Ferretti" country="Argentina" latitude="-45.84833" location="Escalante" longLatPrecision="18" longitude="-67.79444" pageId="6" pageNumber="503" specimenCode="UNS-M0657" specimenCount="3" specimenCount-female="3" stateProvince="Chubut" typeStatus="paratype">
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<collectingCountry id="D4FA765BD712C169FF4247F6FE9BF938" box="[239,338,1720,1744]" name="Argentina" pageId="6" pageNumber="503">Argentina</collectingCountry>
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<specimenCount id="BAEBFD42D712C169FEC847F9FE4FF926" box="[357,390,1719,1743]" pageId="6" pageNumber="503" type="female">3 ♀</specimenCount>
; same data as the holotype;
<date id="D853100BD712C169FD2247F9FCCAF927" box="[655,771,1719,1743]" pageId="6" pageNumber="503" value="2018-11-24">
<collectingDate id="C817E9E3D712C169FD2247F9FCCAF927" box="[655,771,1719,1743]" pageId="6" pageNumber="503" value="2018-11-24">24.XI.2018</collectingDate>
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; excavated from burrows;
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<materialsCitation id="1C853C96D712C169FDAA479FFEBDF8F6" ID-GBIF-Occurrence="4418721301" collectingDate="2018-11-24" collectingDateMax="2019-12-07" collectingDateMin="2018-11-24" collectionCode="TYPE, MATERIAL" collectorName="N. Ferretti" country="Argentina" latitude="-45.84833" location="Escalante" longLatPrecision="18" longitude="-67.79444" pageId="6" pageNumber="503" specimenCode="UNS-M0656" specimenCount="1" specimenCount-male="1" stateProvince="Chubut" typeStatus="paratype">
<specimenCount id="BAEBFD42D712C169FDAA479FFDE2F900" box="[519,555,1745,1769]" pageId="6" pageNumber="503" type="male">1 ♂</specimenCount>
; same data as the holotype;
<date id="D853100BD712C169FF1647A2FEE6F8EC" box="[187,303,1772,1796]" pageId="6" pageNumber="503" value="2018-11-24">
<collectingDate id="C817E9E3D712C169FF1647A2FEE6F8EC" box="[187,303,1772,1796]" pageId="6" pageNumber="503" value="2018-11-24">24.XI.2018</collectingDate>
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; kept alive, became adult in
<date id="D853100BD712C169FDEA47A2FD7EF8EC" box="[583,695,1772,1796]" pageId="6" pageNumber="503" value="2019-12-07">
<collectingDate id="C817E9E3D712C169FDEA47A2FD7EF8EC" box="[583,695,1772,1796]" pageId="6" pageNumber="503" value="2019-12-07">7.XII.2019</collectingDate>
</date>
, died in
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<collectingDate id="C817E9E3D712C169FF294648FF28F8F6" box="[132,225,1798,1822]" pageId="6" pageNumber="503" value="2019-12">XII.2019</collectingDate>
</date>
;
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.
</paragraph>
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<subSubSection id="E4F76540D712C169FF294676FD33F86C" pageId="6" pageNumber="503" type="etymology">
<paragraph id="AC5236CBD712C169FF294676FD33F86C" blockId="6.[131,775,1847,1925]" pageId="6" pageNumber="503">
<smallCapsWord id="AAB4A017D712C169FF294676FF34F8A6" baselines="1865,1866" box="[132,253,1848,1871]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="6" pageNumber="503">ETYMOLOGY</smallCapsWord>
. — The specific name is a patronym in honor of Lionel Andrés Messi, an
<collectingCountry id="D4FA765BD712C169FE9D461CFE59F882" box="[304,400,1874,1898]" name="Argentina" pageId="6" pageNumber="503">Argentine</collectingCountry>
professional footballer and one of the greatest players of all times, unique winner of seven Ballons dor.
</paragraph>
</subSubSection>
<subSubSection id="E4F76540D712C169FF2946D0FAB2FA1E" pageId="6" pageNumber="503" type="diagnosis">
<paragraph id="AC5236CBD712C169FF2946D0FAB2FA1E" blockId="6.[132,775,1949,2026]" lastBlockId="6.[811,1456,1264,1526]" pageId="6" pageNumber="503">
<smallCapsWord id="AAB4A017D712C169FF2946D0FF26F85D" baselines="1967,1968" box="[132,239,1950,1973]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="6" pageNumber="503">DIAGNOSIS</smallCapsWord>
. — Males can be distinguished from the known species, excepting
<taxonomicName id="6BED4D48D712C169FF4A46F6FE92F838" authorityName="Goloboff" authorityYear="1995" box="[231,347,1976,2000]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="503" phylum="Arthropoda" rank="species" species="chilechico">
<emphasis id="9E99EAD9D712C169FF4A46F6FE92F838" box="[231,347,1976,2000]" italics="true" pageId="6" pageNumber="503">A. chilechico</emphasis>
</taxonomicName>
, by the palpal bulb, with basal portion narrower and keel of embolus slightly-developed (
<figureCitation id="34D62A4ED712C169FD9E469CFDA1F802" box="[563,616,2002,2026]" captionStart="FIG" captionStartId="8.[132,143,1286,1304]" captionTargetBox="[163,1417,256,1244]" captionTargetId="figure-347@8.[132,1455,215,1244]" captionTargetPageId="8" captionText="FIG. 4. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A-D, palpal bulb. A, prolateral view; B, dorsal view (green arrow indicates the keel); C, retrolateral view (green arrow indicates the keel); D, dorsal view; E, palp, prolateral view; F, palp, retrolateral view. Abbreviations: Cy, cymbium; Ti, tibia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8382559" httpUri="https://zenodo.org/record/8382559/files/figure.png" pageId="6" pageNumber="503">Fig. 4</figureCitation>
A-D), and by the presence of strong spines on ventral metatarsus I (
<figureCitation id="34D62A4ED712C169FAAF45BEFAAAFAE0" box="[1282,1379,1264,1288]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="6" pageNumber="503">Fig. 3H, I</figureCitation>
). Males differ from those of
<taxonomicName id="6BED4D48D712C169FC5C4445FBAFFACA" authorityName="Goloboff" authorityYear="1995" box="[1009,1126,1290,1314]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="503" phylum="Arthropoda" rank="species" species="chilechico">
<emphasis id="9E99EAD9D712C169FC5C4445FBAFFACA" box="[1009,1126,1290,1314]" italics="true" pageId="6" pageNumber="503">A. chilechico</emphasis>
</taxonomicName>
by the presence of labial cuspules (
<figureCitation id="34D62A4ED712C169FC99446BFCB5FAD5" box="[820,892,1317,1341]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="6" pageNumber="503">Fig. 3E</figureCitation>
) and from
<taxonomicName id="6BED4D48D712C169FC46446BFB9DFAD5" authorityName="Goloboff" authorityYear="1995" box="[1003,1108,1317,1341]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="503" phylum="Arthropoda" rank="species" species="birabeni">
<emphasis id="9E99EAD9D712C169FC46446BFB9DFAD5" box="[1003,1108,1317,1341]" italics="true" pageId="6" pageNumber="503">A. birabeni</emphasis>
</taxonomicName>
by the presence of strong spines on ventral metatarsus I (
<figureCitation id="34D62A4ED712C169FC584471FB9FFABF" box="[1013,1110,1343,1367]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="6" pageNumber="503">Fig. 3H, I</figureCitation>
). Males resemble those of
<taxonomicName id="6BED4D48D712C169FAFC440EFA66FABF" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1940" box="[1361,1455,1344,1367]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="503" phylum="Arthropoda" rank="species" species="notatus">
<emphasis id="9E99EAD9D712C169FAFC440EFA66FABF" box="[1361,1455,1344,1367]" italics="true" pageId="6" pageNumber="503">A. notatus</emphasis>
</taxonomicName>
by the coloration with dark chevron on dorsal abdomen (
<figureCitation id="34D62A4ED712C169FAFA4414FA6BFA9A" box="[1367,1442,1370,1394]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="6" pageNumber="503">Fig. 3C</figureCitation>
), but can be distinguished by the curved embolus without small teeth on apical region (
<figureCitation id="34D62A4ED712C169FC7844C1FBD6FA4F" box="[981,1055,1423,1447]" captionStart="FIG" captionStartId="8.[132,143,1286,1304]" captionTargetBox="[163,1417,256,1244]" captionTargetId="figure-347@8.[132,1455,215,1244]" captionTargetPageId="8" captionText="FIG. 4. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A-D, palpal bulb. A, prolateral view; B, dorsal view (green arrow indicates the keel); C, retrolateral view (green arrow indicates the keel); D, dorsal view; E, palp, prolateral view; F, palp, retrolateral view. Abbreviations: Cy, cymbium; Ti, tibia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8382559" httpUri="https://zenodo.org/record/8382559/files/figure.png" pageId="6" pageNumber="503">Fig. 4C</figureCitation>
) and presence of strong spines on ventral metatarsus I (
<figureCitation id="34D62A4ED712C169FC0344E7FC38FA29" box="[942,1009,1449,1473]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="6" pageNumber="503">Fig. 3I</figureCitation>
). Females differ from the known species of the genus by the shape of the spermathecae, with short oval receptacles, and digitiform internal expansions from their base (
<figureCitation id="34D62A4ED712C169FAB24490FAA7FA1E" box="[1311,1390,1502,1526]" captionStart="FIG" captionStartId="9.[132,143,1986,2004]" captionTargetBox="[134,1455,211,1944]" captionTargetId="figure-15@9.[134,1455,211,1960]" captionTargetPageId="9" captionText="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="6" pageNumber="503">Fig. 5H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E4F76540D712C165FC80477DFC33FAA6" lastPageId="10" lastPageNumber="507" pageId="6" pageNumber="503" type="description">
<paragraph id="AC5236CBD712C169FC80477DFC74F9A3" blockId="6.[811,1458,1587,2028]" box="[813,957,1587,1613]" pageId="6" pageNumber="503">
<smallCapsWord id="AAB4A017D712C169FC80477DFC74F9A3" baselines="1606,1606" box="[813,957,1587,1613]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="6" pageNumber="503">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="AC5236CBD712C169FC80471CFC75F984" blockId="6.[811,1458,1587,2028]" box="[813,956,1618,1644]" pageId="6" pageNumber="503">
<emphasis id="9E99EAD9D712C169FC80471CFC75F984" box="[813,956,1618,1644]" italics="true" pageId="6" pageNumber="503">
Male
<typeStatus id="73568869D712C169FCC4471CFC75F984" box="[873,956,1618,1644]" pageId="6" pageNumber="503" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="AC5236CBD712C169FC80473CFB89F884" blockId="6.[811,1458,1587,2028]" pageId="6" pageNumber="503">
<emphasis id="9E99EAD9D712C169FC80473CFC02F964" bold="true" box="[813,971,1650,1676]" pageId="6" pageNumber="503">Color pattern.</emphasis>
In life, carapace light brown with dark striae and margins, legs light brown with lighter coxae and trochanter, abdomen light brown with dorsal black chevron; in ethanol, carapace brown with dark striae and margins, abdomen with dorsal black chevron and ventrally light brown (
<figureCitation id="34D62A4ED712C169FC98465CFC60F8C4" box="[821,937,1810,1836]" captionStart="FIG" captionStartId="6.[133,144,1174,1192]" captionTargetBox="[135,1453,223,1124]" captionTargetId="figure-463@6.[132,1455,213,1124]" captionTargetPageId="6" captionText="FIG. 2. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., habitus: A, female molted in laboratory; B, female collected and photographed from field; C, male molted in laboratory; D, male collected walking in the field. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8382555" httpUri="https://zenodo.org/record/8382555/files/figure.png" pageId="6" pageNumber="503">Fig. 2C, D</figureCitation>
), in ethanol, carapace brown with dark striae and margins, abdomen with dorsal black chevron and ventrally light brown (
<figureCitation id="34D62A4ED712C169FC17461FFBFBF884" box="[954,1074,1873,1900]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="6" pageNumber="503">Fig. 3C, D</figureCitation>
).
</paragraph>
<paragraph id="AC5236CBD712C168FC80463FFBFDF8A3" blockId="6.[811,1458,1587,2028]" lastBlockId="7.[811,1457,1746,2028]" lastPageId="7" lastPageNumber="504" pageId="6" pageNumber="503">
<emphasis id="9E99EAD9D712C169FC80463FFC2BF863" bold="true" box="[813,994,1905,1931]" pageId="6" pageNumber="503">Cephalothorax.</emphasis>
Total length 9.19. Carapace 4.31 long, 3.47 wide, thoracic region raised. Abdomen 4.66 long, 2.63 wide. Clypeus very narrow, 0.08. Fovea slightly procurved (
<figureCitation id="34D62A4ED712C169FC98469FFC58F803" box="[821,913,2001,2027]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="6" pageNumber="503">Fig. 3A</figureCitation>
), 0.3 long. Eye tubercle 0.59 long, 0.94 wide, slightly elevated. Anterior eye row procurved, posterior row recurved (
<figureCitation id="34D62A4ED713C168FF4347BCFEF6F8E4" box="[238,319,1778,1804]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="7" pageNumber="504">Fig. 3F</figureCitation>
). Eyes sizes and interdistances: AME 0.17, ALE 0.2, PME 0.13, PLE 0.18, AME-AME 0.2, AME-ALE 0.06, PME-PME 0.45, PME-PLE 0.03, ALE-PLE 0.13. Chelicerae 2.19 long, 1.43 wide, with 5 small teeth in prolateral row. Rastellum weak formed by long attenuate setae. Intercheliceral tumescence small with 14 thin setae (
<figureCitation id="34D62A4ED713C168FF2346FFFF20F823" box="[142,233,1969,1995]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="7" pageNumber="504">Fig. 3G</figureCitation>
). Labium 0.67 long, 0.72 wide, with 7 cuspules (
<figureCitation id="34D62A4ED713C168FF23469FFF2DF803" box="[142,228,2001,2027]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="7" pageNumber="504">Fig. 3E</figureCitation>
). Maxillae with 21/18 blunt cuspules on internal basal angle (
<figureCitation id="34D62A4ED713C168FC14479CFBD8F904" box="[953,1041,1746,1772]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="7" pageNumber="504">Fig. 3E</figureCitation>
). Sternum oval (
<figureCitation id="34D62A4ED713C168FB63479CFAEDF904" box="[1230,1316,1746,1772]" captionStart="FIG" captionStartId="7.[132,143,1632,1650]" captionTargetBox="[132,1455,215,1576]" captionTargetId="figure-177@7.[132,1455,215,1608]" captionTargetPageId="7" captionText="FIG. 3. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen, ventral view; E, labium and maxillae, ventral view; F, eyes, dorsal view; G, chelicerae, prolateral view (green arrow indicates intercheliceral tumescence); H, leg I, prolateral view; I, leg I, ventral view. Abbreviations: Mt, metatarsus, Tap, tibial apophysis, Ti, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="7" pageNumber="504">Fig. 3B</figureCitation>
), 2.15 long, 1.61 wide. Sternal sigilla: anterior and medium of same size and close to margin, posterior the largest and distant from margin by
<emphasis id="9E99EAD9D713C168FC09467CFC7CF8A3" box="[932,949,1842,1867]" italics="true" pageId="7" pageNumber="504">c.</emphasis>
1 × length.
</paragraph>
<caption id="F8926643D713C168FF29472EFA6DF948" ID-DOI="http://doi.org/10.5281/zenodo.8382557" ID-Zenodo-Dep="8382557" httpUri="https://zenodo.org/record/8382557/files/figure.png" pageId="7" pageNumber="504" startId="7.[132,143,1632,1650]" targetBox="[132,1455,215,1576]" targetPageId="7" targetType="figure">
<paragraph id="AC5236CBD713C168FF29472EFA6DF948" blockId="7.[132,1457,1632,1696]" pageId="7" pageNumber="504">
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. 3. —
<taxonomicName id="6BED4D48D713C168FF7A472EFD98F99A" authority="Signorotto &amp; Ferretti" authorityName="Signorotto &amp; Mancini &amp; Ferretti" authorityYear="2023" box="[215,593,1632,1650]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="504" phylum="Arthropoda" rank="species" species="messii" status="sp. nov.">
<emphasis id="9E99EAD9D713C168FF7A472EFE54F99A" box="[215,413,1632,1650]" italics="true" pageId="7" pageNumber="504">Acanthogonatus messii</emphasis>
Signorotto &amp; Ferretti
</taxonomicName>
<taxonomicNameLabel id="85AA57A2D713C168FDFA472EFD42F99A" box="[599,651,1632,1650]" pageId="7" pageNumber="504" rank="species">n. sp.</taxonomicNameLabel>
, holotype male:
<emphasis id="9E99EAD9D713C168FCBA472EFCEDF99A" bold="true" box="[791,804,1632,1650]" pageId="7" pageNumber="504">A</emphasis>
, carapace, dorsal view;
<emphasis id="9E99EAD9D713C168FC5A472EFBCDF99A" bold="true" box="[1015,1028,1632,1650]" pageId="7" pageNumber="504">B</emphasis>
, sternum, ventral view;
<emphasis id="9E99EAD9D713C168FB7E472EFB28F99A" bold="true" box="[1235,1249,1632,1650]" pageId="7" pageNumber="504">C</emphasis>
, abdomen, dorsal view;
<emphasis id="9E99EAD9D713C168FF294739FF5BF961" bold="true" box="[132,146,1655,1673]" pageId="7" pageNumber="504">D</emphasis>
, abdomen, ventral view;
<emphasis id="9E99EAD9D713C168FEC44739FEBCF961" bold="true" box="[361,373,1655,1673]" pageId="7" pageNumber="504">E</emphasis>
, labium and maxillae, ventral view;
<emphasis id="9E99EAD9D713C168FD094739FD66F961" bold="true" box="[676,687,1655,1673]" pageId="7" pageNumber="504">F</emphasis>
, eyes, dorsal view;
<emphasis id="9E99EAD9D713C168FCF54739FCAFF961" bold="true" box="[856,870,1655,1673]" pageId="7" pageNumber="504">G</emphasis>
, chelicerae, prolateral view (
<emphasis id="9E99EAD9D713C168FBF44739FB0FF961" bold="true" box="[1113,1222,1655,1673]" pageId="7" pageNumber="504">green arrow</emphasis>
indicates intercheliceral tumescence);
<emphasis id="9E99EAD9D713C168FF4447C0FF3EF948" bold="true" box="[233,247,1678,1696]" pageId="7" pageNumber="504">H</emphasis>
, leg I, prolateral view;
<emphasis id="9E99EAD9D713C168FE1F47C0FE71F948" bold="true" box="[434,440,1678,1696]" pageId="7" pageNumber="504">I</emphasis>
, leg I, ventral view. Abbreviations:
<emphasis id="9E99EAD9D713C168FD7547C0FD39F948" bold="true" box="[728,752,1678,1696]" pageId="7" pageNumber="504">Mt</emphasis>
, metatarsus,
<emphasis id="9E99EAD9D713C168FCCC47C0FC49F948" bold="true" box="[865,896,1678,1696]" pageId="7" pageNumber="504">Tap</emphasis>
, tibial apophysis,
<emphasis id="9E99EAD9D713C168FBBA47C0FBEEF948" bold="true" box="[1047,1063,1678,1696]" pageId="7" pageNumber="504">Ti</emphasis>
, tibia. Scale bars: A-D, F-I, 1 mm, E, 0.5 mm.
</paragraph>
</caption>
<paragraph id="AC5236CBD713C168FC80461CFABAF803" blockId="7.[811,1457,1746,2028]" pageId="7" pageNumber="504">
<emphasis id="9E99EAD9D713C168FC80461CFC1DF884" bold="true" box="[813,980,1874,1900]" pageId="7" pageNumber="504">Measurements.</emphasis>
Palp: femur 1.95/ patella 0.93/ tibia 1.36/ cymbium 0.88/ total 4.49; legs: I: femur 3.49/ patella 2.06/ tibia 2.26/ metatarsus 3.32/ tarsus 2.19/ total 13.32; II: 3.62/ 1.75/ 2.58/ 3.23/ 2.14/ 13.32; III: 3.65/ 1.86/ 2.6/ 4.23/ 2.45/ 14.79; IV: 3.87/ 1.98/ 3.57/ 4.84/ 3.12/ 17.38.
</paragraph>
<caption id="F8926643D71CC167FF294448FED4FAAE" ID-DOI="http://doi.org/10.5281/zenodo.8382559" ID-Zenodo-Dep="8382559" httpUri="https://zenodo.org/record/8382559/files/figure.png" pageId="8" pageNumber="505" startId="8.[132,143,1286,1304]" targetBox="[163,1417,256,1244]" targetPageId="8" targetType="figure">
<paragraph id="AC5236CBD71CC167FF294448FED4FAAE" blockId="8.[132,1457,1286,1350]" pageId="8" pageNumber="505">
<smallCapsWord id="AAB4A017D71CC167FF294448FF54FAFF" baselines="1299,1300" box="[132,157,1286,1304]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="8" pageNumber="505">FIG</smallCapsWord>
. 4. —
<taxonomicName id="6BED4D48D71CC167FF7E4448FD89FAF0" authority="Signorotto &amp; Ferretti" authorityName="Signorotto &amp; Mancini &amp; Ferretti" authorityYear="2023" box="[211,576,1286,1304]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="505" phylum="Arthropoda" rank="species" species="messii" status="sp. nov.">
<emphasis id="9E99EAD9D71CC167FF7E4448FE5DFAF0" box="[211,404,1286,1304]" italics="true" pageId="8" pageNumber="505">Acanthogonatus messii</emphasis>
Signorotto &amp; Ferretti
</taxonomicName>
<taxonomicNameLabel id="85AA57A2D71CC167FDE84448FDBEFAF0" box="[581,631,1286,1304]" pageId="8" pageNumber="505" rank="species">n. sp.</taxonomicNameLabel>
, holotype male:
<emphasis id="9E99EAD9D71CC167FD514448FCD7FAF0" bold="true" box="[764,798,1286,1304]" pageId="8" pageNumber="505">A -D</emphasis>
, palpal bulb.
<emphasis id="9E99EAD9D71CC167FC204448FC53FAF0" bold="true" box="[909,922,1286,1304]" pageId="8" pageNumber="505">A</emphasis>
, prolateral view;
<emphasis id="9E99EAD9D71CC167FB8B4448FBFAFAF0" bold="true" box="[1062,1075,1286,1304]" pageId="8" pageNumber="505">B</emphasis>
, dorsal view (
<emphasis id="9E99EAD9D71CC167FB084448FAD9FAF0" bold="true" box="[1189,1296,1286,1304]" pageId="8" pageNumber="505">green arrow</emphasis>
indicates the keel);
<emphasis id="9E99EAD9D71CC167FF294453FF5BFAC7" bold="true" box="[132,146,1309,1327]" pageId="8" pageNumber="505">C</emphasis>
, retrolateral view (
<emphasis id="9E99EAD9D71CC167FE834453FE55FAC7" bold="true" box="[302,412,1309,1327]" pageId="8" pageNumber="505">green arrow</emphasis>
indicates the keel);
<emphasis id="9E99EAD9D71CC167FDEA4453FD9CFAC7" bold="true" box="[583,597,1309,1327]" pageId="8" pageNumber="505">D</emphasis>
, dorsal view;
<emphasis id="9E99EAD9D71CC167FD614453FD11FAC7" bold="true" box="[716,728,1309,1327]" pageId="8" pageNumber="505">E</emphasis>
, palp, prolateral view;
<emphasis id="9E99EAD9D71CC167FC374453FC6CFAC7" bold="true" box="[922,933,1309,1327]" pageId="8" pageNumber="505">F</emphasis>
, palp, retrolateral view. Abbreviations:
<emphasis id="9E99EAD9D71CC167FB424453FACEFAC7" bold="true" box="[1263,1287,1309,1327]" pageId="8" pageNumber="505">Cy</emphasis>
, cymbium;
<emphasis id="9E99EAD9D71CC167FAC64453FAB2FAC7" bold="true" box="[1387,1403,1309,1327]" pageId="8" pageNumber="505">Ti</emphasis>
, tibia. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="AC5236CBD71CC167FF2944DDFE7DF803" blockId="8.[130,778,1427,2027]" pageId="8" pageNumber="505">
<emphasis id="9E99EAD9D71CC167FF2944DDFF30FA45" bold="true" box="[132,249,1427,1453]" pageId="8" pageNumber="505">Spination.</emphasis>
Palp: tibia: d1-1, p1-1, v2-3. Tarsi of legs I-IV, 0. I: femur: d1-1-1-1-1-1-1-1-1-1; patella: p1-1-1; tibia: p1-1-2-2ap (on common base), r1-1, v2-1-1-1-1-1; metatarsus: v1-2-1. II: femur: d1-1-1-1-1-1-1-1, p1; patella: p1-1-1; tibia: p1-1-1, r1-1, v1-1-1-1-1-1-2ap, metatarsus: d1, p1-1-1, r1-1, v1-1-1-1-1. III: femur: d1-1-1-1-1-1-1, p1-1-1; patella: p1-1-1, r1; tibia: d1, p1-1, r1-1, v2-1-1-3ap; metatarsus: d1-1-1-1-1, p1-1-1-1, r1-1-1ap, v1-1-2ap. IV: femur: d1-1-1-1-1, p1-1-1, r1-1; patella: p1-1, r1; tibia: d1-1-1, p1-1, r1, v1-1-1-1-3ap; metatarsus: d2-1-1-1-1, p1-1-1, r1-1, v2-1-1-3ap. Tarsi I-IV flexible. Scopulae on tarsi I-IV light; I, II entire, III divided by two rows of thin setae, IV divided by three rows of thin setae. Scopulae of metatarsus I entire on 1/2 of length; II entire on 1/3 of length; III, IV absent. Metatarsal preening combs absent from legs I-IV. STC with double row of teeth: I: 12 13 12 12; II: 14 14 12 13; III: 14 13 13 13; IV: 14 14 14 12. Four spinnerets: PMS 0.5 long; PLS: basal segment 1.08 long, median 0.38 long, apical 0.43 long.
</paragraph>
<paragraph id="AC5236CBD71CC167FC8044DDFB0BF9A5" blockId="8.[812,1457,1427,1613]" pageId="8" pageNumber="505">
<emphasis id="9E99EAD9D71CC167FC8044DDFCAEFA45" bold="true" box="[813,871,1427,1453]" pageId="8" pageNumber="505">Palp.</emphasis>
Cymbium with elongate dense setae, denser at tip; tibia dorsally with two elongated setae (
<figureCitation id="34D62A4ED71CC167FB7E44FDFA88FA25" box="[1235,1345,1459,1485]" captionStart="FIG" captionStartId="8.[132,143,1286,1304]" captionTargetBox="[163,1417,256,1244]" captionTargetId="figure-347@8.[132,1455,215,1244]" captionTargetPageId="8" captionText="FIG. 4. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A-D, palpal bulb. A, prolateral view; B, dorsal view (green arrow indicates the keel); C, retrolateral view (green arrow indicates the keel); D, dorsal view; E, palp, prolateral view; F, palp, retrolateral view. Abbreviations: Cy, cymbium; Ti, tibia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8382559" httpUri="https://zenodo.org/record/8382559/files/figure.png" pageId="8" pageNumber="505">Fig. 4E, F</figureCitation>
), ventrally with a deep excavation on apical third; bulb with tegulum globose, embolus thin, slightly curved to retrolateral face on apical half, one well-developed keel,
<emphasis id="9E99EAD9D71CC167FB1E475DFB0DF9C4" box="[1203,1220,1555,1580]" italics="true" pageId="8" pageNumber="505">c.</emphasis>
1/3 of embolus length, sperm duct very sinuous (
<figureCitation id="34D62A4ED71CC167FBEF477CFB48F9A5" box="[1090,1153,1586,1613]" captionStart="FIG" captionStartId="8.[132,143,1286,1304]" captionTargetBox="[163,1417,256,1244]" captionTargetId="figure-347@8.[132,1455,215,1244]" captionTargetPageId="8" captionText="FIG. 4. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., holotype male: A-D, palpal bulb. A, prolateral view; B, dorsal view (green arrow indicates the keel); C, retrolateral view (green arrow indicates the keel); D, dorsal view; E, palp, prolateral view; F, palp, retrolateral view. Abbreviations: Cy, cymbium; Ti, tibia. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8382559" httpUri="https://zenodo.org/record/8382559/files/figure.png" pageId="8" pageNumber="505">Fig. 4</figureCitation>
A-D).
</paragraph>
<paragraph id="AC5236CBD71CC167FC80473CFBB5F964" blockId="8.[811,1457,1650,2028]" box="[813,1148,1650,1677]" pageId="8" pageNumber="505">
<emphasis id="9E99EAD9D71CC167FC80473CFBB5F964" box="[813,1148,1650,1677]" italics="true" pageId="8" pageNumber="505">
Female (
<typeStatus id="73568869D71CC167FC29473DFC14F965" box="[900,989,1651,1677]" pageId="8" pageNumber="505" type="paratype">paratype</typeStatus>
UNS-M0657)
</emphasis>
</paragraph>
<paragraph id="AC5236CBD71CC167FC8047DCFBF0F8C4" blockId="8.[811,1457,1650,2028]" pageId="8" pageNumber="505">
<emphasis id="9E99EAD9D71CC167FC8047DCFC0BF944" bold="true" box="[813,962,1682,1708]" pageId="8" pageNumber="505">Color pattern.</emphasis>
In life, carapace brown with dark striae and margins, abdomen light brown with dorsal black chevron (
<figureCitation id="34D62A4ED71CC167FAF147FCFCF7F904" captionStart="FIG" captionStartId="6.[133,144,1174,1192]" captionTargetBox="[135,1453,223,1124]" captionTargetId="figure-463@6.[132,1455,213,1124]" captionTargetPageId="6" captionText="FIG. 2. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., habitus: A, female molted in laboratory; B, female collected and photographed from field; C, male molted in laboratory; D, male collected walking in the field. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.8382555" httpUri="https://zenodo.org/record/8382555/files/figure.png" pageId="8" pageNumber="505">Fig. 2A, B</figureCitation>
); in ethanol, carapace brown with dark striae and margins (
<figureCitation id="34D62A4ED71CC167FC9B47BCFC40F8E5" box="[822,905,1778,1805]" captionStart="FIG" captionStartId="9.[132,143,1986,2004]" captionTargetBox="[134,1455,211,1944]" captionTargetId="figure-15@9.[134,1455,211,1960]" captionTargetPageId="9" captionText="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="8" pageNumber="505">Fig. 5A</figureCitation>
), abdomen with dorsal black chevron and ventrally light brown (
<figureCitation id="34D62A4ED71CC167FC1A465CFBE4F8C4" box="[951,1069,1809,1836]" captionStart="FIG" captionStartId="9.[132,143,1986,2004]" captionTargetBox="[134,1455,211,1944]" captionTargetId="figure-15@9.[134,1455,211,1960]" captionTargetPageId="9" captionText="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="8" pageNumber="505">Fig. 5C, D</figureCitation>
).
</paragraph>
<paragraph id="AC5236CBD71CC165FC80467FFE4EFA25" blockId="8.[811,1457,1650,2028]" lastBlockId="10.[130,777,1204,2028]" lastPageId="10" lastPageNumber="507" pageId="8" pageNumber="505">
<emphasis id="9E99EAD9D71CC167FC80467FFC1CF8A3" bold="true" box="[813,981,1841,1867]" pageId="8" pageNumber="505">Cephalothorax.</emphasis>
Total length 11.93. Carapace 6.43 long, 5.03 wide, thoracic region raised. Abdomen 5.51 long, 3.28 wide. Clypeus very narrow, 0.13. Fovea slightly procurved, 0.7 long. Eye tubercle 0.71 long, 1.3 wide, slightly elevated. Anterior eye row procurved, posterior row recurved (
<figureCitation id="34D62A4ED71CC167FAB846FFFAA3F824" box="[1301,1386,1969,1996]" captionStart="FIG" captionStartId="9.[132,143,1986,2004]" captionTargetBox="[134,1455,211,1944]" captionTargetId="figure-15@9.[134,1455,211,1960]" captionTargetPageId="9" captionText="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="8" pageNumber="505">Fig. 5F</figureCitation>
). Eyes sizes and interdistances: AME 0.2, ALE 0.34, PME 0.18, PLE 0.25, AME-AME 0.26, AME-ALE 0.11, PME-PME 0.58, PME-PLE 0.05, ALE-PLE 0.12. Chelicerae 2.85 long, 2.14 wide, with 5 small teeth in prolateral row. Rastellum weak formed by long attenuate setae. Labium 0.85 long, 0.9 wide, with 1 cuspule (
<figureCitation id="34D62A4ED71EC165FE81447DFEB4FAA6" box="[300,381,1331,1358]" captionStart="FIG" captionStartId="9.[132,143,1986,2004]" captionTargetBox="[134,1455,211,1944]" captionTargetId="figure-15@9.[134,1455,211,1960]" captionTargetPageId="9" captionText="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="10" pageNumber="507">Fig. 5E</figureCitation>
). Maxillae with 36/34 blunt cuspules on internal basal angle. Sternum oval (
<figureCitation id="34D62A4ED71EC165FD81441DFD48FA86" box="[556,641,1363,1390]" captionStart="FIG" captionStartId="9.[132,143,1986,2004]" captionTargetBox="[134,1455,211,1944]" captionTargetId="figure-15@9.[134,1455,211,1960]" captionTargetPageId="9" captionText="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="10" pageNumber="507">Fig. 5B</figureCitation>
), 3.08 long, 2.5 wide. Sternal sigilla: anterior slightly smaller than medium and close to margin, posterior the largest and distant from margin by
<emphasis id="9E99EAD9D71EC165FF5444FAFEC3FA25" box="[249,266,1460,1485]" italics="true" pageId="10" pageNumber="507">c.</emphasis>
1 × length.
</paragraph>
<caption id="F8926643D71DC166FF29468CFA78F803" ID-DOI="http://doi.org/10.5281/zenodo.8382561" ID-Zenodo-Dep="8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="9" pageNumber="506" startId="9.[132,143,1986,2004]" targetBox="[134,1455,211,1944]" targetPageId="9" targetType="figure">
<paragraph id="AC5236CBD71DC166FF29468CFA78F803" blockId="9.[132,1457,1986,2027]" pageId="9" pageNumber="506">
<smallCapsWord id="AAB4A017D71DC166FF29468CFF55F83B" baselines="1999,2000" box="[132,156,1986,2004]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="9" pageNumber="506">FIG</smallCapsWord>
. 5. —
<taxonomicName id="6BED4D48D71DC166FF7B468CFD82F83C" authority="Signorotto &amp; Ferretti" authorityName="Signorotto &amp; Mancini &amp; Ferretti" authorityYear="2023" box="[214,587,1986,2004]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="506" phylum="Arthropoda" rank="species" species="messii" status="sp. nov.">
<emphasis id="9E99EAD9D71DC166FF7B468CFE50F83C" box="[214,409,1986,2004]" italics="true" pageId="9" pageNumber="506">Acanthogonatus messii</emphasis>
Signorotto &amp; Ferretti
</taxonomicName>
<taxonomicNameLabel id="85AA57A2D71DC166FDFC468CFD4CF83C" box="[593,645,1986,2004]" pageId="9" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
, paratype female:
<emphasis id="9E99EAD9D71DC166FC8D468CFCE4F83C" bold="true" box="[800,813,1986,2004]" pageId="9" pageNumber="506">A</emphasis>
, carapace, dorsal view;
<emphasis id="9E99EAD9D71DC166FC50468CFBC3F83C" bold="true" box="[1021,1034,1986,2004]" pageId="9" pageNumber="506">B</emphasis>
, sternum, ventral view;
<emphasis id="9E99EAD9D71DC166FB7B468CFB2DF83C" bold="true" box="[1238,1252,1986,2004]" pageId="9" pageNumber="506">C</emphasis>
, abdomen, dorsal view;
<emphasis id="9E99EAD9D71DC166FF294697FF5BF803" bold="true" box="[132,146,2009,2027]" pageId="9" pageNumber="506">D</emphasis>
, abdomen,ventral view;
<emphasis id="9E99EAD9D71DC166FECD4697FEA5F803" bold="true" box="[352,364,2009,2027]" pageId="9" pageNumber="506">E</emphasis>
, labium and maxillae, ventral view;
<emphasis id="9E99EAD9D71DC166FD3C4697FD55F803" bold="true" box="[657,668,2009,2027]" pageId="9" pageNumber="506">F</emphasis>
, eyes, dorsal view;
<emphasis id="9E99EAD9D71DC166FC934697FC85F803" bold="true" box="[830,844,2009,2027]" pageId="9" pageNumber="506">G</emphasis>
, leg IV, ventral view;
<emphasis id="9E99EAD9D71DC166FC5A4697FBCCF803" bold="true" box="[1015,1029,2009,2027]" pageId="9" pageNumber="506">H</emphasis>
, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm.
</paragraph>
</caption>
<caption id="F8926643D71EC165FF67451EFAA5FB8A" ID-DOI="http://doi.org/10.5281/zenodo.8382564" ID-Zenodo-Dep="8382564" box="[202,1388,1104,1122]" httpUri="https://zenodo.org/record/8382564/files/figure.png" pageId="10" pageNumber="507" startId="10.[202,213,1104,1122]" targetBox="[132,1456,196,1069]" targetPageId="10" targetType="figure">
<paragraph id="AC5236CBD71EC165FF67451EFAA5FB8A" blockId="10.[202,1388,1104,1122]" box="[202,1388,1104,1122]" pageId="10" pageNumber="507">
<smallCapsWord id="AAB4A017D71EC165FF67451EFF2BFB89" baselines="1117,1118" box="[202,226,1104,1122]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="10" pageNumber="507">FIG</smallCapsWord>
. 6. —
<emphasis id="9E99EAD9D71EC165FEB7451EFEEEFB8A" bold="true" box="[282,295,1104,1122]" pageId="10" pageNumber="507">A</emphasis>
, habitat at type locality of
<taxonomicName id="6BED4D48D71EC165FDAE451EFCBBFB8A" authority="Signorotto &amp; Ferretti" authorityName="Signorotto &amp; Mancini &amp; Ferretti" authorityYear="2023" box="[515,882,1104,1122]" class="Arachnida" family="Pycnothelidae" genus="Acanthogonatus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="507" phylum="Arthropoda" rank="species" species="messii" status="sp. nov.">
<emphasis id="9E99EAD9D71EC165FDAE451EFD0DFB8A" box="[515,708,1104,1122]" italics="true" pageId="10" pageNumber="507">Acanthogonatus messii</emphasis>
Signorotto &amp; Ferretti
</taxonomicName>
<taxonomicNameLabel id="85AA57A2D71EC165FCDA451EFC60FB8A" box="[887,937,1104,1122]" pageId="10" pageNumber="507" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="9E99EAD9D71EC165FC1D451EFC74FB8A" bold="true" box="[944,957,1104,1122]" pageId="10" pageNumber="507">B</emphasis>
,
<emphasis id="9E99EAD9D71EC165FC6A451EFC1CFB8A" bold="true" box="[967,981,1104,1122]" pageId="10" pageNumber="507">C</emphasis>
, female burrows opened at night at type locality.
</paragraph>
</caption>
<paragraph id="AC5236CBD71EC165FF29449DFF36F964" blockId="10.[130,777,1204,2028]" pageId="10" pageNumber="507">
<emphasis id="9E99EAD9D71EC165FF29449DFEE4FA05" bold="true" box="[132,301,1491,1517]" pageId="10" pageNumber="507">Measurements.</emphasis>
Palp: femur 3.22/ patella 1.82/ tibia 2.04/ cymbium 1.92/ total 9; legs: measurements: I: femur 4.58/ patella 3.04/ tibia 2.87/ metatarsus 3.07/ tarsus 1.64/ total 15.2; II: 4.29/ 2.75/ 2.29/ 2.82/ 1.96/ 14.11; III: 3.81/ 2.29/ 2.08/ 3.64/ 2.29/ 14.11; IV: 4.67/ 2.93/ 3.46/ 4.31/ 2.5/ 17.87.
</paragraph>
<paragraph id="AC5236CBD71EC165FF2947DCFAD5FB06" blockId="10.[130,777,1204,2028]" lastBlockId="10.[813,1456,1204,1358]" pageId="10" pageNumber="507">
<emphasis id="9E99EAD9D71EC165FF2947DCFF30F944" bold="true" box="[132,249,1682,1708]" pageId="10" pageNumber="507">Spination.</emphasis>
Palp: tibia: p1ap, v4ap. Tarsi of legs I-IV, 0. I: femur:d1-1-1-1-1-1-1, p1; patella: 0; tibia: p1, v1ap; metatarsus: 0. II: femur: d1-1-1-1; patella: 0; tibia: v1-1; metatarsus: v1-1. III: femur: d1; patella: p1-1; tibia: r1, v1-1ap; metatarsus: d1, p1-1, r1-1, v1-1ap. IV: femur: 0; patella: 0; tibia: r1, v1ap; metatarsus: p1, r1. Scopulae on tarsi I-IV light; I, II entire, III divided by three rows of thin setae, IV divided by four rows of thin setae. Scopulae of metatarsus I, II entire and on full length; III on 1/5 of length; IV absent. Metatarsal preening combs absent from legs I-IV (
<figureCitation id="34D62A4ED71EC165FD8246FFFD4EF824" box="[559,647,1969,1996]" captionStart="FIG" captionStartId="9.[132,143,1986,2004]" captionTargetBox="[134,1455,211,1944]" captionTargetId="figure-15@9.[134,1455,211,1960]" captionTargetPageId="9" captionText="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="10" pageNumber="507">Fig. 5G</figureCitation>
). STC with double row of teeth: I: 8 8 8 7; II: 7 8 8 7; III: 9 8 8 8; IV: 8 8 8 6. Four spinnerets: PMS 0.83 long; PLS: basal segment 1.31 long, median 0.40 long, apical 0.55 long.
</paragraph>
<paragraph id="AC5236CBD71EC165FC8045BDFC33FAA6" blockId="10.[813,1456,1204,1358]" pageId="10" pageNumber="507">
<emphasis id="9E99EAD9D71EC165FC8045BDFC05FAE5" bold="true" box="[813,972,1267,1293]" pageId="10" pageNumber="507">Spermathecae.</emphasis>
Each with one globose oval dome with very short duct from which arises an elongated digitiform receptaculum (
<figureCitation id="34D62A4ED71EC165FC39447DFC24FAA6" box="[916,1005,1331,1358]" captionStart="FIG" captionStartId="9.[132,143,1986,2004]" captionTargetBox="[134,1455,211,1944]" captionTargetId="figure-15@9.[134,1455,211,1960]" captionTargetPageId="9" captionText="FIG. 5. — Acanthogonatus messii Signorotto &amp; Ferretti n. sp., paratype female: A, carapace, dorsal view; B, sternum, ventral view; C, abdomen, dorsal view; D, abdomen,ventral view;E, labium and maxillae, ventral view; F, eyes, dorsal view; G, leg IV, ventral view; H, spermathecae. Scale bars:A-G, 1 mm, H, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8382561" httpUri="https://zenodo.org/record/8382561/files/figure.png" pageId="10" pageNumber="507">Fig. 5H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E4F76540D71EC165FC80443AFC09F964" pageId="10" pageNumber="507" type="distribution">
<paragraph id="AC5236CBD71EC165FC80443AFB0EFA64" blockId="10.[813,1457,1396,1676]" box="[813,1223,1396,1422]" pageId="10" pageNumber="507">
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<smallCapsWord id="AAB4A017D71EC165FC7D4437FC37FA64" baselines="1415" box="[976,1022,1401,1420]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="10" pageNumber="507">AND</smallCapsWord>
<smallCapsWord id="AAB4A017D71EC165FBA84437FBAFFA64" baselines="1415" box="[1029,1126,1401,1420]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="natural" pageId="10" pageNumber="507">NATURAL</smallCapsWord>
<smallCapsWord id="AAB4A017D71EC165FBC04437FB0EFA64" baselines="1415" box="[1133,1223,1401,1420]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="history" pageId="10" pageNumber="507">HISTORY</smallCapsWord>
</paragraph>
<paragraph id="AC5236CBD71EC165FC8044DAFC09F964" blockId="10.[813,1457,1396,1676]" pageId="10" pageNumber="507">
Known only from the
<typeStatus id="73568869D71EC165FBB044DAFB82FA46" box="[1053,1099,1428,1454]" pageId="10" pageNumber="507">type</typeStatus>
locality (
<figureCitation id="34D62A4ED71EC165FB0344DDFACAFA45" box="[1198,1283,1427,1453]" captionStart="FIG" captionStartId="10.[202,213,1104,1122]" captionTargetBox="[132,1456,196,1069]" captionTargetId="figure-509@10.[132,1455,213,1062]" captionTargetPageId="10" captionText="FIG. 6. — A, habitat at type locality of Acanthogonatus messii Signorotto &amp; Ferretti n. sp.; B, C, female burrows opened at night at type locality." figureDoi="http://doi.org/10.5281/zenodo.8382564" httpUri="https://zenodo.org/record/8382564/files/figure.png" pageId="10" pageNumber="507">Fig. 6A</figureCitation>
). The open burrows from females (
<figureCitation id="34D62A4ED71EC165FC5344FDFBA6FA25" box="[1022,1135,1459,1485]" captionStart="FIG" captionStartId="10.[202,213,1104,1122]" captionTargetBox="[132,1456,196,1069]" captionTargetId="figure-509@10.[132,1455,213,1062]" captionTargetPageId="10" captionText="FIG. 6. — A, habitat at type locality of Acanthogonatus messii Signorotto &amp; Ferretti n. sp.; B, C, female burrows opened at night at type locality." figureDoi="http://doi.org/10.5281/zenodo.8382564" httpUri="https://zenodo.org/record/8382564/files/figure.png" pageId="10" pageNumber="507">Fig. 6B, C</figureCitation>
) were found at night at about 21:00 h in a sandy area with small shrubs. Burrow entrances had a diameter of approximately
<quantity id="6B159B2ED71EC165FB3344BCFB26F9E5" box="[1182,1263,1522,1549]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="507" unit="mm" value="15.0">15 mm</quantity>
and spiders were actively feeding. One adult male was found walking in the area at 21:50 h in a sandy area close to the female burrows, which is indicative of the reproductive period of the species in November.
</paragraph>
</subSubSection>
<subSubSection id="E4F76540D71EC165FC8047FCFA89F803" pageId="10" pageNumber="507" type="discussion">
<paragraph id="AC5236CBD71EC165FC8047FCFBCAF923" blockId="10.[811,1455,1714,2028]" box="[813,1027,1714,1740]" pageId="10" pageNumber="507">
<smallCapsWord id="AAB4A017D71EC165FC8047FCFC57F924" baselines="1733,1734" box="[813,926,1714,1740]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Cladistic" pageId="10" pageNumber="507">CLADISTIC</smallCapsWord>
<smallCapsWord id="AAB4A017D71EC165FC0847F6FBCAF923" baselines="1734" box="[933,1027,1720,1739]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="analysis" pageId="10" pageNumber="507">ANALYSIS</smallCapsWord>
</paragraph>
<paragraph id="AC5236CBD71EC165FC80479CFA89F803" blockId="10.[811,1455,1714,2028]" pageId="10" pageNumber="507">
Search with implied weighting (IW) (k-values = 3.122, 3.665, 4.124, 4.877, 5.972 and 7.237) resulted in 3 equally parsimonious trees with 111 steps. The remaining k-values (8.253, 12.832, 16.117 and 25.376) resulted in 8 equally parsimonious trees with 113 steps. The strict consensus of IW resulted in a tree with 113 steps, CI = 42 and RI = 66. The tree in
<figureCitation id="34D62A4ED71EC165FC8046DFFC4CF844" box="[813,901,1937,1964]" captionStart="FIG" captionStartId="14.[132,143,1552,1570]" captionTargetBox="[189,1455,202,1510]" captionTargetId="figure-54@14.[189,1313,205,1504]" captionTargetPageId="14" captionText="FIG. 7. — Most parsimonious cladogram using IW. Symbols: black squares: exclusive synapomorphies; white squares: homoplasies. Numbers above the circles: characters; below: states. Species group notations follows Goloboff (1995)." figureDoi="http://doi.org/10.5281/zenodo.8398922" httpUri="https://zenodo.org/record/8398922/files/figure.png" pageId="10" pageNumber="507">Figure 7</figureCitation>
is the most consistent topology from the sensitivity analysis (script iw.run) and correspond to the results of the IW analyses using K-value (concavity) of 16.117.
</paragraph>
</subSubSection>
</treatment>
</document>