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<document id="C7ED2BD0659F783527E2BBD14E24AE23" ID-CLB-Dataset="22232" ID-DOI="10.11646/zootaxa.4776.1.1" ID-GBIF-Dataset="d92dcbf9-8bc1-43a6-8be1-b553a257ca0b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3821288" ID-ZooBank="5A3743A2-9E64-45E8-920D-92C9ED9CAB5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589267646334" checkinUser="plazi" docAuthor="Serrano, Artur R. M., Capela, Rúben A., Nunes, Telmo &amp; Santos, Carmen Van-Dú- Nem Neto" docDate="2020" docId="1B3C87AA004FFFBCFF1DFB2BFB56F945" docLanguage="en" docName="zootaxa.4776.1.1.pdf.imf" docOrigin="Zootaxa 4776 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Gnathocera cruda subsp. leopoldi" docType="treatment" docVersion="7" lastPageNumber="95" masterDocId="E705FFD20011FFE2FF8AFFFAFFA4FFA6" masterDocTitle="The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes" masterLastPageNumber="130" masterPageNumber="1" pageNumber="95" updateTime="1698839254326" updateUser="ExternalLinkService">
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<mods:title id="1B7D28F129937429A58A6B981B364D5C">The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes</mods:title>
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<mods:namePart id="AC9D04FF1832D01645541CB58A8C61DE">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="4776922D7F5D0F62AA4EC27CB8C54B63">Capela, Rúben A.</mods:namePart>
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<treatment id="1B3C87AA004FFFBCFF1DFB2BFB56F945" ID-DOI="http://doi.org/10.5281/zenodo.3845545" ID-GBIF-Taxon="164123810" ID-Zenodo-Dep="3845545" LSID="urn:lsid:plazi:treatment:1B3C87AA004FFFBCFF1DFB2BFB56F945" httpUri="http://treatment.plazi.org/id/1B3C87AA004FFFBCFF1DFB2BFB56F945" lastPageNumber="95" pageId="94" pageNumber="95">
<subSubSection id="DB8F6537004FFFBCFF1DFB2BFD0FFB4D" box="[151,683,1232,1259]" pageId="94" pageNumber="95" type="nomenclature">
<paragraph id="932A36BC004FFFBCFF1DFB2BFD0FFB4D" blockId="94.[151,1437,1232,1763]" box="[151,683,1232,1259]" pageId="94" pageNumber="95">
<heading id="C86281D0004FFFBCFF1DFB2BFD0FFB4D" bold="true" box="[151,683,1232,1259]" fontSize="11" level="1" pageId="94" pageNumber="95" reason="1">
<emphasis id="A1E1EAAE004FFFBCFF1DFB2BFD0FFB4D" bold="true" box="[151,683,1232,1259]" pageId="94" pageNumber="95">
*
<taxonomicName id="54954D3F004FFFBCFF2FFB2AFD0FFB4D" authority="(Bourgoin, 1929)" baseAuthorityName="Bourgoin" baseAuthorityYear="1929" box="[165,683,1232,1259]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="cruda" subSpecies="leopoldi">
<emphasis id="A1E1EAAE004FFFBCFF2FFB2AFE7FFB4D" bold="true" box="[165,475,1232,1259]" italics="true" pageId="94" pageNumber="95">Gnathocera cruda leopoldi</emphasis>
(Bourgoin, 1929)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537004FFFBCFF1DFB0FFEA3FAB6" box="[151,263,1269,1296]" pageId="94" pageNumber="95" type="description">
<paragraph id="932A36BC004FFFBCFF1DFB0FFEA3FAB6" blockId="94.[151,1437,1232,1763]" box="[151,263,1269,1296]" pageId="94" pageNumber="95">
(
<figureCitation id="0BAE2A39004FFFBCFF2AFB0FFF5BFAB6" box="[160,255,1269,1296]" captionStart="FIGURE 22" captionStartId="97.[151,250,1876,1901]" captionTargetBox="[247,1340,229,1838]" captionTargetId="figure@97.[235,1352,181,1851]" captionTargetPageId="97" captionText="FIGURE 22. Cetoniinae species of Angola. a, Gnathocera cruda gracilis (dorsal view), male, MZUC; b, Gnathocera cruda leopoldi (dorsal view), male, MZUC; c and d, Gnathocera marginata (dorsal views), males, MZUC and Dundo, LNO, IICT, respectively; e, Gnathocera overlaeti (dorsal view), male, Dundo, LNO, IICT; f, Gnathocera trivialis (dorsal view), male, Muita, LNO, IICT; g and h, Gnathocera trivittata ardoini (dorsal views), male and female, respectively, Dundo, LNO, IICT; i, Gnathocera trivittata dorsodiscolor (dorsal view), male, MZUC." figureDoi="http://doi.org/10.5281/zenodo.3821332" httpUri="https://zenodo.org/record/3821332/files/figure.png" pageId="94" pageNumber="95">Fig. 22b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537004FFFBCFF4DFAE2FE73FADD" pageId="94" pageNumber="95" type="distribution">
<paragraph id="932A36BC004FFFBCFF4DFAE2FE50FA95" blockId="94.[151,1437,1232,1763]" box="[199,500,1304,1331]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFAE2FEFDFA95" bold="true" box="[199,345,1304,1331]" pageId="94" pageNumber="95">Distribution</emphasis>
: ANG, DOC.
</paragraph>
<paragraph id="932A36BC004FFFBCFF4DFAC6FD74FAF1" blockId="94.[151,1437,1232,1763]" box="[199,720,1340,1367]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFAC6FDCEFAF1" bold="true" box="[199,618,1340,1367]" pageId="94" pageNumber="95">
Distribution in
<collectingCountry id="EB82762C004FFFBCFEF7FAC6FE77FAF1" box="[381,467,1340,1367]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
(Provinces):
</emphasis>
1) LNO.
</paragraph>
<paragraph id="932A36BC004FFFBCFF4DFA9AFE73FADD" blockId="94.[151,1437,1232,1763]" box="[199,471,1376,1403]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFA9AFE36FADD" bold="true" box="[199,402,1376,1403]" pageId="94" pageNumber="95">Historic records:</emphasis>
none.
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537004FFFBCFF4DFA7EFBA3FA65" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="932A36BC004FFFBCFF4DFA7EFBA3FA65" blockId="94.[151,1437,1232,1763]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFA7EFE16FA39" bold="true" box="[199,434,1412,1439]" pageId="94" pageNumber="95">Material examined:</emphasis>
<materialsCitation id="23FD3CE1004FFFBCFE32FA7FFBBFFA06" ID-GBIF-Occurrence="2612165350" box="[440,1051,1413,1440]" country="Angola" location="Angola" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1">
<collectingCountry id="EB82762C004FFFBCFE32FA7FFDA9FA39" box="[440,525,1413,1439]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
,
<specimenCount id="8593FD35004FFFBCFD92FA7FFDE5FA39" box="[536,577,1413,1439]" pageId="94" pageNumber="95" type="female">1 ♀</specimenCount>
, Nº 1907 (blue label), nº 142 (white label)
</materialsCitation>
,
<specimenCount id="8593FD35004FFFBCFBAFFA7FFBF5FA39" box="[1061,1105,1413,1439]" pageId="94" pageNumber="95" type="male">2 ♂</specimenCount>
, Nº 1907 (blue labels), nº 143 (white labels),
<specimenCount id="8593FD35004FFFBCFEB4FA53FECEFA65" box="[318,362,1449,1475]" pageId="94" pageNumber="95" type="male">1 ♂</specimenCount>
, Nº 1907 (blue label), old collection, box nº IX-78, MZUC.
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537004FFFBCFF4DFA37FB56F945" pageId="94" pageNumber="95" type="discussion">
<paragraph id="932A36BC004FFFBCFF4DFA37FB56F945" blockId="94.[151,1437,1232,1763]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFA37FE9FFA41" bold="true" box="[199,315,1485,1511]" pageId="94" pageNumber="95">Remarks.</emphasis>
This subspecies of
<taxonomicName id="54954D3F004FFFBCFD9EFA37FDD2FA41" authorityName="Janson" authorityYear="1877" box="[532,630,1485,1511]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="cruda">
<emphasis id="A1E1EAAE004FFFBCFD9EFA37FDD2FA41" box="[532,630,1485,1511]" italics="true" pageId="94" pageNumber="95">G. cruda</emphasis>
</taxonomicName>
was established by Antoinne (2002a), but
<bibRefCitation id="F7044B4D004FFFBCFBC4FA37FB5BFA41" author="Krajcik, M." box="[1102,1279,1485,1511]" pageId="94" pageNumber="95" refId="ref84860" refString="Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI." type="book" year="2008">Krajcik (2008a)</bibRefCitation>
and
<bibRefCitation id="F7044B4D004FFFBCFABEFA37FE8CF9AD" author="Beinhundner, G." pageId="94" pageNumber="95" refId="ref81744" refString="Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp." type="book" year="2017">Beinhundner (2017a)</bibRefCitation>
raised it to the species level without any explanation. Its presence in
<collectingCountry id="EB82762C004FFFBCFBBCFA0BFB2CF9AD" box="[1078,1160,1521,1547]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
is suggested by
<bibRefCitation id="F7044B4D004FFFBCFAC8FA0BFF4AF989" author="Antoine, Ph." pageId="94" pageNumber="95" pagination="65 - 92" refId="ref80368" refString="Antoine, Ph. (2002 a) Contribution a la connaissance des Cetoniidae africains (Coleoptera, Scarabaeoidea). Coleopteres, 8 (4), 65 - 92." type="journal article" year="2002">Antoine (2002a)</bibRefCitation>
based on Ruters notes under the invalid name
<taxonomicName id="54954D3F004FFFBCFC87F9EFFC4CF989" box="[781,1000,1557,1583]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="cruda" subSpecies="machadoi">
<emphasis id="A1E1EAAE004FFFBCFC87F9EFFC4CF989" box="[781,1000,1557,1583]" italics="true" pageId="94" pageNumber="95">G. cruda machadoi</emphasis>
</taxonomicName>
, a form never described by Ruter (see
<bibRefCitation id="F7044B4D004FFFBCFF1DF9C3FE80F9F5" author="Allard, V." box="[151,292,1593,1619]" pageId="94" pageNumber="95" pagination="1 - 142" refId="ref79922" refString="Allard, V. (1991) Goliathini 4: Les Coleopteres du monde. Sciences Naturelles de Venette, 11, 1 - 142." type="journal article" year="1991">Allard 1991</bibRefCitation>
), which fits very well the description of
<taxonomicName id="54954D3F004FFFBCFC8CF9C3FB39F9F5" authority="(Antoine 2002 a)" baseAuthorityName="Antoine" baseAuthorityYear="2002" box="[774,1181,1593,1619]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="cruda" subSpecies="leopoldi">
<emphasis id="A1E1EAAE004FFFBCFC8CF9C3FC77F9F5" box="[774,979,1593,1619]" italics="true" pageId="94" pageNumber="95">G. cruda leopoldi</emphasis>
(
<bibRefCitation id="F7044B4D004FFFBCFC6FF9C3FB31F9F5" author="Antoine, Ph." box="[997,1173,1593,1619]" pageId="94" pageNumber="95" pagination="65 - 92" refId="ref80368" refString="Antoine, Ph. (2002 a) Contribution a la connaissance des Cetoniidae africains (Coleoptera, Scarabaeoidea). Coleopteres, 8 (4), 65 - 92." type="journal article" year="2002">Antoine 2002a</bibRefCitation>
)
</taxonomicName>
. However, this author (
<bibRefCitation id="F7044B4D004FFFBCFF15F9A7FEE3F9D1" author="Antoine, Ph." box="[159,327,1629,1655]" pageId="94" pageNumber="95" pagination="65 - 92" refId="ref80368" refString="Antoine, Ph. (2002 a) Contribution a la connaissance des Cetoniidae africains (Coleoptera, Scarabaeoidea). Coleopteres, 8 (4), 65 - 92." type="journal article" year="2002">Antoine 2002a</bibRefCitation>
) studied many specimens and segregated
<taxonomicName id="54954D3F004FFFBCFC91F9A7FC7FF9D1" baseAuthorityName="Janson" baseAuthorityYear="1877" box="[795,987,1629,1655]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="cruda" subSpecies="gracilis">
<emphasis id="A1E1EAAE004FFFBCFC91F9A7FC7FF9D1" box="[795,987,1629,1655]" italics="true" pageId="94" pageNumber="95">G. cruda gracilis</emphasis>
</taxonomicName>
and
<taxonomicName id="54954D3F004FFFBCFB9AF9A7FB72F9D1" baseAuthorityName="Antoine" baseAuthorityYear="2002" box="[1040,1238,1629,1655]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="cruda" subSpecies="leopoldi">
<emphasis id="A1E1EAAE004FFFBCFB9AF9A7FB72F9D1" box="[1040,1238,1629,1655]" italics="true" pageId="94" pageNumber="95">G. cruda leopoldi</emphasis>
</taxonomicName>
by its geographic distribution [the former in ANG, CON and DOC and the later in DOC (“Haut
<collectingRegion id="5151F85E004FFFBCFC65F97BFB8EF93D" box="[1007,1066,1665,1691]" country="Angola" name="Zaire" pageId="94" pageNumber="95">Zaire</collectingRegion>
et Shaba=Katanga)]. The MZUC specimens fit well the description of
<taxonomicName id="54954D3F004FFFBCFDBBF95FFD52F919" baseAuthorityName="Antoine" baseAuthorityYear="2002" box="[561,758,1701,1727]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="cruda" subSpecies="leopoldi">
<emphasis id="A1E1EAAE004FFFBCFDBBF95FFD52F919" box="[561,758,1701,1727]" italics="true" pageId="94" pageNumber="95">G. cruda leopoldi</emphasis>
</taxonomicName>
, but we can not precise their localization in
<collectingCountry id="EB82762C004FFFBCFB55F95FFA91F919" box="[1247,1333,1701,1727]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
. Biological or ecological data are not available for this subspecies. It is a new subspecies record for
<collectingCountry id="EB82762C004FFFBCFB12F933FB4AF945" box="[1176,1262,1737,1763]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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