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<document id="C1F43F9ADD43FA8E808F431AFC14364D" ID-CLB-Dataset="22232" ID-DOI="10.11646/zootaxa.4776.1.1" ID-GBIF-Dataset="d92dcbf9-8bc1-43a6-8be1-b553a257ca0b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3821288" ID-ZooBank="5A3743A2-9E64-45E8-920D-92C9ED9CAB5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589267646334" checkinUser="plazi" docAuthor="Serrano, Artur R. M., Capela, Rúben A., Nunes, Telmo &amp; Santos, Carmen Van-Dú- Nem Neto" docDate="2020" docId="1B3C87AA000FFFFCFF1DFF62FB54FD55" docLanguage="en" docName="zootaxa.4776.1.1.pdf.imf" docOrigin="Zootaxa 4776 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pachnoda orphanula subsp. nachtigali" docType="treatment" docVersion="7" lastPageNumber="31" masterDocId="E705FFD20011FFE2FF8AFFFAFFA4FFA6" masterDocTitle="The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes" masterLastPageNumber="130" masterPageNumber="1" pageNumber="31" updateTime="1698839254326" updateUser="ExternalLinkService">
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<mods:title id="35B4823A2B5F706ED14D0589E6BA7036">The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes</mods:title>
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<mods:namePart id="B2CEE58619FE63C673FF402AD45D4859">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="249A66410C4EF3DFAE301319A7F36D75">Capela, Rúben A.</mods:namePart>
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<mods:namePart id="19E4832B01BA402023C18F433FA754A9">Nunes, Telmo</mods:namePart>
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<treatment id="1B3C87AA000FFFFCFF1DFF62FB54FD55" ID-DOI="http://doi.org/10.5281/zenodo.3845229" ID-GBIF-Taxon="164123893" ID-Zenodo-Dep="3845229" LSID="urn:lsid:plazi:treatment:1B3C87AA000FFFFCFF1DFF62FB54FD55" httpUri="http://treatment.plazi.org/id/1B3C87AA000FFFFCFF1DFF62FB54FD55" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="DB8F6537000FFFFCFF1DFF62FD18FF15" box="[151,700,152,179]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="932A36BC000FFFFCFF1DFF62FD18FF15" blockId="30.[151,1437,152,755]" box="[151,700,152,179]" pageId="30" pageNumber="31">
<heading id="C86281D0000FFFFCFF1DFF62FD18FF15" bold="true" box="[151,700,152,179]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<taxonomicName id="54954D3F000FFFFCFF1DFF62FD18FF15" authority="(Kraatz, 1885)" baseAuthorityName="Kraatz" baseAuthorityYear="1885" box="[151,700,152,179]" class="Insecta" family="Cetoniidae" genus="Pachnoda" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="orphanula" subSpecies="nachtigali">
<emphasis id="A1E1EAAE000FFFFCFF1DFF62FD18FF15" bold="true" box="[151,700,152,179]" pageId="30" pageNumber="31">
<emphasis id="A1E1EAAE000FFFFCFF1DFF62FDACFF15" bold="true" box="[151,520,152,179]" italics="true" pageId="30" pageNumber="31">Pachnoda orphanula nachtigali</emphasis>
(Kraatz, 1885)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537000FFFFCFF1DFF47FEE7FF71" box="[151,323,188,215]" pageId="30" pageNumber="31" type="description">
<paragraph id="932A36BC000FFFFCFF1DFF47FEE7FF71" blockId="30.[151,1437,152,755]" box="[151,323,188,215]" pageId="30" pageNumber="31">
(
<figureCitation id="0BAE2A39000FFFFCFF15FF47FEB6FF71" box="[159,274,189,215]" captionStart="FIGURE 4" captionStartId="29.[151,250,1876,1901]" captionTargetBox="[253,1345,201,1840]" captionTargetId="figure@29.[234,1353,181,1850]" captionTargetPageId="29" captionText="FIGURE 4. Cetoniinae species of Angola. a and b, Pachnoda discolor kolbei, male (dorsal view) and parameres (frontal view), respectively, Satchijamba, BIE, ASC; c, Pachnoda frontalis (dorsal view), female, MZUC; d, Pachnoda katangensis (dorsal view), female, Luso (=Luena), MOX, IICT; e, Pachnoda leopoldiana (dorsal view), female, MZUC; f, Pachnoda leopoldiana (dorsal view), Dundo, LNO, IICT; g, Pachnoda marginata marginata (dorsal view), male, MZUC; h and i, Pachnoda orphanula nachtigali, male (dorsal view) and parameres (frontal view), respectively, Dundo, LNO, IICT." figureDoi="http://doi.org/10.5281/zenodo.3821296" httpUri="https://zenodo.org/record/3821296/files/figure.png" pageId="30" pageNumber="31">Figs. 4h, i</figureCitation>
,
<figureCitation id="0BAE2A39000FFFFCFE95FF46FE9EFF71" box="[287,314,188,215]" captionStart="FIGURE 5" captionStartId="32.[151,250,1876,1901]" captionTargetBox="[252,1353,216,1843]" captionTargetId="figure@32.[232,1356,181,1851]" captionTargetPageId="32" captionText="FIGURE 5. Cetoniinae species of Angola. a, Pachnoda orphanula nachtigali (dorsal view), male, MZUC; b, Pachnoda picturata (dorsal view), female, MZUC; c, Pachnoda poggei, parameres (frontal view), Dundo, LNO, IICT; d and e, Pachnoda rubriventris rubrocinctoides, male (dorsal view) and parameres (lateral view), respectively, Carima, MAL, ASC; f and g, Pachnoda rubrocincta (dorsal views), male and female, respectively, MZUC; h and i, Pachnoda rufovirens, male (dorsal view) and parameres (frontal view), respectively, Kakande, BIE, ASC." figureDoi="http://doi.org/10.5281/zenodo.3821298" httpUri="https://zenodo.org/record/3821298/files/figure.png" pageId="30" pageNumber="31">5a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537000FFFFCFF4DFF1AFF7AFE2D" pageId="30" pageNumber="31" type="distribution">
<paragraph id="932A36BC000FFFFCFF4DFF1AFD99FF5D" blockId="30.[151,1437,152,755]" box="[199,573,224,251]" pageId="30" pageNumber="31">
<emphasis id="A1E1EAAE000FFFFCFF4DFF1AFEFDFF5D" bold="true" box="[199,345,224,251]" pageId="30" pageNumber="31">Distribution</emphasis>
: ANG, CON, DOC.
</paragraph>
<paragraph id="932A36BC000FFFFCFF4DFEFEFD74FEB9" blockId="30.[151,1437,152,755]" box="[199,720,260,287]" pageId="30" pageNumber="31">
<emphasis id="A1E1EAAE000FFFFCFF4DFEFEFDCEFEB9" bold="true" box="[199,618,260,287]" pageId="30" pageNumber="31">
Distribution in
<collectingCountry id="EB82762C000FFFFCFEF7FEFEFE77FEB9" box="[381,467,260,287]" name="Angola" pageId="30" pageNumber="31">Angola</collectingCountry>
(Provinces):
</emphasis>
1) LNO.
</paragraph>
<paragraph id="932A36BC000FFFFCFF4DFED2FE36FEE5" blockId="30.[151,1437,152,755]" box="[199,402,296,323]" pageId="30" pageNumber="31">
<emphasis id="A1E1EAAE000FFFFCFF4DFED2FE36FEE5" bold="true" box="[199,402,296,323]" pageId="30" pageNumber="31">Historic records:</emphasis>
</paragraph>
<paragraph id="932A36BC000FFFFCFF4DFEB7FF7AFE2D" blockId="30.[151,1437,152,755]" pageId="30" pageNumber="31">
—Dundo (LNO) (
<bibRefCitation id="F7044B4D000FFFFCFE1CFEB7FDC9FEC1" author="Gomes Alves, M. G." box="[406,621,332,359]" pageId="30" pageNumber="31" pagination="63 - 88" refId="ref83304" refString="Gomes Alves, M. G. (1959) Cetonideos da Africa portuguesa. Memorias da Junta de Investigacao do Ultramar, 8, 63 - 88, pls. 1 - 8." type="journal article" year="1959">Gomes Alves 1959</bibRefCitation>
and
<bibRefCitation id="F7044B4D000FFFFCFD2DFEB7FCE4FEC1" author="Ferreira, M. C." box="[679,832,332,359]" pageId="30" pageNumber="31" pagination="415 - 1317" refId="ref83177" refString="Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317." type="journal article" year="1965">Ferreira 1965</bibRefCitation>
sub
<taxonomicName id="54954D3F000FFFFCFCFDFEB7FF76FE2D" authority="Burgeon" authorityName="Burgeon" authorityYear="1932" class="Insecta" family="Cetoniidae" genus="Pachnoda" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="variety" species="nachtigali" variety="pseudomarginata">
<emphasis id="A1E1EAAE000FFFFCFCFDFEB7FBC7FEC1" box="[887,1123,333,359]" italics="true" pageId="30" pageNumber="31">Pachnoda nachtigali</emphasis>
var.
<emphasis id="A1E1EAAE000FFFFCFB14FEB7FAC5FEC1" box="[1182,1377,333,359]" italics="true" pageId="30" pageNumber="31">pseudomarginata</emphasis>
Burgeon
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537000FFFFCFF4DFE6EFD1EFE51" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="932A36BC000FFFFCFF4DFE6EFD1EFE51" blockId="30.[151,1437,152,755]" pageId="30" pageNumber="31">
<emphasis id="A1E1EAAE000FFFFCFF4DFE6EFE16FE09" bold="true" box="[199,434,404,431]" pageId="30" pageNumber="31">Material examined:</emphasis>
<materialsCitation id="23FD3CE1000FFFFCFE3DFE6FFBB8FE75" ID-GBIF-Occurrence="2612165308" collectingDate="1948-10" collectionCode="LNO" collectorName="Monteiro &amp; Dundo" country="Angola" elevation="1470" latitude="-7.366667" location="Angola" longLatPrecision="1303" longitude="20.833334" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1">
<collectingCountry id="EB82762C000FFFFCFE3DFE6FFDA8FE09" box="[439,524,405,431]" name="Angola" pageId="30" pageNumber="31">Angola</collectingCountry>
,
<specimenCount id="8593FD35000FFFFCFD9DFE6FFDE7FE09" box="[535,579,405,431]" pageId="30" pageNumber="31" type="male">1 ♂</specimenCount>
, Nº 1907 (blue label), nº 122 (white label),
<collectorName id="3E60536A000FFFFCFBADFE6FFB2BFE09" box="[1063,1167,405,431]" pageId="30" pageNumber="31">Monteiro</collectorName>
leg., old collection, box nº XIV-83, MZUC;
<collectorName id="3E60536A000FFFFCFEFDFE43FE67FE75" box="[375,451,441,467]" pageId="30" pageNumber="31">Dundo</collectorName>
(
<geoCoordinate id="F6A1507B000FFFFCFE58FE43FD8CFE75" box="[466,552,441,467]" degrees="7" direction="south" minutes="22" orientation="latitude" pageId="30" pageNumber="31" precision="925" value="-7.366667">7º 22´S</geoCoordinate>
,
<geoCoordinate id="F6A1507B000FFFFCFDB9FE43FD3CFE75" box="[563,664,440,467]" degrees="20" direction="east" minutes="50" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="20.833334">20º 50´E</geoCoordinate>
,
<quantity id="546D9B59000FFFFCFD29FE43FD5CFE75" box="[675,760,441,467]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="30" pageNumber="31" unit="m" value="1470.0">
<elevation id="18B8D18F000FFFFCFD29FE43FD5CFE75" box="[675,760,441,467]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="30" pageNumber="31" unit="m" value="1470.0">1470 m</elevation>
</quantity>
alt., 51) (
<collectionCode id="F584AE79000FFFFCFCEEFE43FC07FE75" box="[868,931,441,467]" pageId="30" pageNumber="31">LNO</collectionCode>
), -.
<date id="E72B107C000FFFFCFC48FE43FBB8FE75" box="[962,1052,441,467]" pageId="30" pageNumber="31" value="1948-10">
<collectingDate id="F76FE994000FFFFCFC48FE43FBB8FE75" box="[962,1052,441,467]" pageId="30" pageNumber="31" value="1948-10">X.1948</collectingDate>
</date>
</materialsCitation>
,
<specimenCount id="8593FD35000FFFFCFBA2FE43FBF7FE75" box="[1064,1107,441,467]" pageId="30" pageNumber="31" type="male">1 ♂</specimenCount>
, Nº 2130, Junta de Investiga- ções Coloniais (Entomologia), IICT, MUHNAC.
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537000FFFFCFF4DFDFBFB54FD55" pageId="30" pageNumber="31" type="discussion">
<paragraph id="932A36BC000FFFFCFF4DFDFBFB54FD55" blockId="30.[151,1437,152,755]" pageId="30" pageNumber="31">
<emphasis id="A1E1EAAE000FFFFCFF4DFDFBFE9FFDBD" bold="true" box="[199,315,513,539]" pageId="30" pageNumber="31">Remarks.</emphasis>
The species is polytypic, but the number of valid subspecies remains controversial.
<bibRefCitation id="F7044B4D000FFFFCFB61FDFBFF40FD99" author="Marais, E. &amp; Holm, E." pageId="30" pageNumber="31" pagination="1 - 125" refId="ref85857" refString="Marais, E. &amp; Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125." type="journal article" year="1992">Marais &amp; Holm (1992)</bibRefCitation>
,
<bibRefCitation id="F7044B4D000FFFFCFF78FDDFFE03FD99" author="Krajcik, M." box="[242,423,549,575]" pageId="30" pageNumber="31" refId="ref84860" refString="Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI." type="book" year="2008">Krajcik (2008a)</bibRefCitation>
and
<bibRefCitation id="F7044B4D000FFFFCFE69FDDFFD43FD99" author="Schoolmeesters, P." box="[483,743,549,575]" pageId="30" pageNumber="31" pagination="2017" refId="ref88502" refString="Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. &amp; Penev, L. (Eds.), Species 2000 &amp; ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)" type="book chapter" year="2018">Schoolmeesters (2018)</bibRefCitation>
recognized five subspecies in Africa, including the nominal one. However,
<bibRefCitation id="F7044B4D000FFFFCFECFFDB3FD92FDC5" author="Beinhundner, G." box="[325,566,585,611]" pageId="30" pageNumber="31" refId="ref81744" refString="Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp." type="book" year="2017">Beinhundner (2017a)</bibRefCitation>
recognized also
<taxonomicName id="54954D3F000FFFFCFD70FDB3FAC2FDC5" authority="Burgeon, 1932" authorityName="Burgeon" authorityYear="1932" box="[762,1382,585,611]" class="Insecta" family="Cetoniidae" genus="Pachnoda" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="orphanula" subSpecies="pseudomarginata">
<emphasis id="A1E1EAAE000FFFFCFD70FDB3FB11FDC5" box="[762,1205,585,611]" italics="true" pageId="30" pageNumber="31">Pachnoda orphanula pseudomarginata</emphasis>
<bibRefCitation id="F7044B4D000FFFFCFB34FDB3FAC2FDC5" author="Burgeon, L." box="[1214,1382,585,611]" pageId="30" pageNumber="31" pagination="47 - 128" refId="ref82218" refString="Burgeon, L. (1932) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres-Scarabeides- Cetoniines. Annales du Musee Royal du Congo Belge, Sciences Zoologiques, Series 3, 2 (2), 47 - 128." type="journal article" year="1932">Burgeon, 1932</bibRefCitation>
</taxonomicName>
as a good subspecies, which is considered a synonym of
<taxonomicName id="54954D3F000FFFFCFD68FD94FC4FFD21" baseAuthorityName="Kraatz" baseAuthorityYear="1885" box="[738,1003,621,647]" class="Insecta" family="Cetoniidae" genus="Pachnoda" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="orphanula" subSpecies="nachtigali">
<emphasis id="A1E1EAAE000FFFFCFD68FD94FC4FFD21" box="[738,1003,621,647]" italics="true" pageId="30" pageNumber="31">P. orphanula nachtigali</emphasis>
</taxonomicName>
by the former authors. The subspecies was recorded for northeastern
<collectingCountry id="EB82762C000FFFFCFE6DFD6BFD9DFD0D" box="[487,569,657,683]" name="Angola" pageId="30" pageNumber="31">Angola</collectingCountry>
by
<bibRefCitation id="F7044B4D000FFFFCFDE8FD6BFCEFFD0D" author="Gomes Alves, M. G." box="[610,843,656,683]" pageId="30" pageNumber="31" pagination="63 - 88" refId="ref83304" refString="Gomes Alves, M. G. (1959) Cetonideos da Africa portuguesa. Memorias da Junta de Investigacao do Ultramar, 8, 63 - 88, pls. 1 - 8." type="journal article" year="1959">Gomes Alves (1959)</bibRefCitation>
. The adult male specimens of MZUC and MUHNAC fit well this species description, including the parameres conformation (
<figureCitation id="0BAE2A39000FFFFCFC4FFD4FFB9EFD69" box="[965,1082,693,719]" captionStart="FIGURE 4" captionStartId="29.[151,250,1876,1901]" captionTargetBox="[253,1345,201,1840]" captionTargetId="figure@29.[234,1353,181,1850]" captionTargetPageId="29" captionText="FIGURE 4. Cetoniinae species of Angola. a and b, Pachnoda discolor kolbei, male (dorsal view) and parameres (frontal view), respectively, Satchijamba, BIE, ASC; c, Pachnoda frontalis (dorsal view), female, MZUC; d, Pachnoda katangensis (dorsal view), female, Luso (=Luena), MOX, IICT; e, Pachnoda leopoldiana (dorsal view), female, MZUC; f, Pachnoda leopoldiana (dorsal view), Dundo, LNO, IICT; g, Pachnoda marginata marginata (dorsal view), male, MZUC; h and i, Pachnoda orphanula nachtigali, male (dorsal view) and parameres (frontal view), respectively, Dundo, LNO, IICT." figureDoi="http://doi.org/10.5281/zenodo.3821296" httpUri="https://zenodo.org/record/3821296/files/figure.png" pageId="30" pageNumber="31">Figs. 4h, i</figureCitation>
,
<figureCitation id="0BAE2A39000FFFFCFBC2FD4EFBC1FD69" box="[1096,1125,692,719]" captionStart="FIGURE 5" captionStartId="32.[151,250,1876,1901]" captionTargetBox="[252,1353,216,1843]" captionTargetId="figure@32.[232,1356,181,1851]" captionTargetPageId="32" captionText="FIGURE 5. Cetoniinae species of Angola. a, Pachnoda orphanula nachtigali (dorsal view), male, MZUC; b, Pachnoda picturata (dorsal view), female, MZUC; c, Pachnoda poggei, parameres (frontal view), Dundo, LNO, IICT; d and e, Pachnoda rubriventris rubrocinctoides, male (dorsal view) and parameres (lateral view), respectively, Carima, MAL, ASC; f and g, Pachnoda rubrocincta (dorsal views), male and female, respectively, MZUC; h and i, Pachnoda rufovirens, male (dorsal view) and parameres (frontal view), respectively, Kakande, BIE, ASC." figureDoi="http://doi.org/10.5281/zenodo.3821298" httpUri="https://zenodo.org/record/3821298/files/figure.png" pageId="30" pageNumber="31">5a</figureCitation>
). The MUHNAC specimen was collected by light trapping. Biological and ecological data are not available for this subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>