treatments-xml/data/A0/2F/87/A02F87F5FF8BF97A1730FCC3FAA4FB3E.xml
2024-06-21 12:45:58 +02:00

275 lines
37 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="C76C53FCFA901D105C0CFD4D11B72A29" ID-DOI="10.11646/zootaxa.1182.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5064071" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625278111192" checkinUser="felipe" docAuthor="Mermudes, José Ricardo M. &amp; Napp, Dilma Solange" docDate="2006" docId="A02F87F5FF8BF97A1730FCC3FAA4FB3E" docLanguage="en" docName="zt01182p130.pdf" docOrigin="Zootaxa 1182 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1182.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Ptychoderes crustatus Frieser 1988" docType="treatment" docVersion="3" lastPageNumber="26" masterDocId="5C16FF8DFF9CF9631638FFB7FF8DFFC1" masterDocTitle="Revision and cladistic analysis of the genus: Ptychoderes Schoenherr, 1823 (Coleoptera, Anthribidae, Anthribinae, Ptychoderini)" masterLastPageNumber="130" masterPageNumber="1" pageNumber="24" updateTime="1698984982767" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="9E0871F66DB7D60A6B6F12B96C635CDA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3C3DB635D17F5D0BE8EA2C4448AEAAC7">
<mods:title id="D10D0419A3A62B993CF2D8CC63F927CA">Revision and cladistic analysis of the genus: Ptychoderes Schoenherr, 1823 (Coleoptera, Anthribidae, Anthribinae, Ptychoderini)</mods:title>
</mods:titleInfo>
<mods:name id="7A46C9DF1212AB18C3A2CDB721357BC8" type="personal">
<mods:role id="F4AF814E13802DD70B7064E7F678FCE5">
<mods:roleTerm id="A2DEE77B79E679E9BDF4ABE9F48916D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C7F6A99DE6E1195874E31902BE6D64A">Mermudes, José Ricardo M.</mods:namePart>
</mods:name>
<mods:name id="A70E6B11A81A4D74B3AC8158E8049BBA" type="personal">
<mods:role id="9057937CB76E0C87C604063756AE6CCF">
<mods:roleTerm id="E242B92272C136E1FE07E0BC36D31855">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2EBA6D1908058FAD9ED7169697356487">Napp, Dilma Solange</mods:namePart>
</mods:name>
<mods:typeOfResource id="01ADA43CBA255BB0CB0C63F682AFCBEF">text</mods:typeOfResource>
<mods:relatedItem id="BE7DA278F9533184F610E160B50DE149" type="host">
<mods:titleInfo id="CD62A43AD6E1E1323DDBA2D318B94F29">
<mods:title id="348F96F71E76C0F57BA16F66DAAC8C3A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A81E901BFFCBD5347CECB2C05760C96C">
<mods:date id="B30997D5BD3B3A0EA241D41000C55360">2006</mods:date>
<mods:detail id="DACCCB8A44BBED001345305BA2629CDB" type="pubDate">
<mods:number id="8661D1ECF230957FD3FC104B16306C31">2006-04-21</mods:number>
</mods:detail>
<mods:detail id="00B71900FB19339649A513EA91DA31C5" type="volume">
<mods:number id="25DC5C842D00ABD42936B8CC3164622F">1182</mods:number>
</mods:detail>
<mods:detail id="2972818FB94800295C9DD1B8496BCA9F" type="issue">
<mods:number id="6C2FEA51C224A577272200BF58873A80">1</mods:number>
</mods:detail>
<mods:extent id="857703523DE6CDEBB4A8875224748985" unit="page">
<mods:start id="7D57F1958EEDB07C4AAC09D6AD11F592">1</mods:start>
<mods:end id="B272F77DDFD9CE10852A36AA2461103A">130</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0E9F5C5FECDAF52864DB9AC10809F7B9">
<mods:url id="FC4B611364344D1D776D5022CE0F63AA">https://biotaxa.org/Zootaxa/article/view/zootaxa.1182.1.1</mods:url>
</mods:location>
<mods:classification id="05E511FA714E023F0DA39D0853C921DA">journal article</mods:classification>
<mods:identifier id="27DD954456F34476468C0C70633BFBF9" type="DOI">10.11646/zootaxa.1182.1.1</mods:identifier>
<mods:identifier id="A506EA73368D2A19B22D03423062A316" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="9C5EEE624384B13671575772A5152A0F" type="Zenodo-Dep">5064071</mods:identifier>
<mods:identifier id="B687DD4BAB148976038EF8BDAF91CC7A" type="ZooBank">56758062-FF9C-448F-98AE-92C3EC71022C</mods:identifier>
</mods:mods>
<treatment id="A02F87F5FF8BF97A1730FCC3FAA4FB3E" LSID="urn:lsid:plazi:treatment:A02F87F5FF8BF97A1730FCC3FAA4FB3E" httpUri="http://treatment.plazi.org/id/A02F87F5FF8BF97A1730FCC3FAA4FB3E" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="609C6568FF8BF9741730FCC3FD3AFC4F" box="[264,695,884,910]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="283936E3FF8BF9741730FCC3FD3AFC4F" blockId="23.[264,695,884,945]" box="[264,695,884,910]" pageId="23" pageNumber="24">
<heading id="7371818FFF8BF9741730FCC3FD3AFC4F" bold="true" box="[264,695,884,910]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<taxonomicName id="EF864D60FF8BF9741730FCC3FD3AFC4F" ID-CoL="8K9LP" authority="Frieser, 1988" authorityName="Frieser" authorityYear="1988" box="[264,695,884,910]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="crustatus">
<emphasis id="1AF2EAF1FF8BF9741730FCC3FD3AFC4F" bold="true" box="[264,695,884,910]" pageId="23" pageNumber="24">
<emphasis id="1AF2EAF1FF8BF9741730FCC3FD9DFC4F" bold="true" box="[264,528,884,910]" italics="true" pageId="23" pageNumber="24">Ptychoderes crustatus</emphasis>
<bibRefCitation id="4C174B12FF8BF9741420FCC3FD3AFC4F" author="Frieser, R." box="[536,695,884,910]" pageId="23" pageNumber="24" pagination="233 - 234" refId="ref63164" refString="Frieser, R. (1988) Eine neue Ptychoderes- spezies aus Guyane Francaise (Coleoptera, Anthribidae). Revue francaise d'Entomologie, (N. S.), 10 (3), 233 - 234." type="journal article" year="1988">Frieser, 1988</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="609C6568FF8BF9741730FC20FE67FC70" box="[264,490,919,945]" pageId="23" pageNumber="24" type="description">
<paragraph id="283936E3FF8BF9741730FC20FE67FC70" blockId="23.[264,695,884,945]" box="[264,490,919,945]" pageId="23" pageNumber="24">
<figureCitation id="B0BD2A66FF8BF9741730FC20FEFAFC70" box="[264,375,919,945]" captionStart="FIGURES 208214" captionStartId="117.[264,382,1545,1569]" captionTargetBox="[293,1285,343,1521]" captionTargetId="figure-64@117.[292,1294,326,1521]" captionTargetPageId="117" captionText="FIGURES 208214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes &amp; Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male." figureDoi="http://doi.org/10.5281/zenodo.5064125" httpUri="https://zenodo.org/record/5064125/files/figure.png" pageId="23" pageNumber="24">Figs. 208</figureCitation>
,
<figureCitation id="B0BD2A66FF8BF97417BCFC20FE3FFC70" box="[388,434,919,945]" captionStart="FIGURES 239250" captionStartId="121.[264,383,1756,1780]" captionTargetBox="[309,1279,326,1725]" captionTargetId="figure-77@121.[309,1279,326,1725]" captionTargetPageId="121" captionText="FIGURES 239250. Pronotum. 239, Tribotropis prasinata; 240, Unanthribus grandis; 241, Ptychoderes crustatus; 242, P. brevis, female holotype; 243, P. longicollis, holotype; 244, P. jordani, male holotype; 245, P. magnus, male holotype; 246, P. depressus; 247, P. virgatus; 248, P. bivittatus, male syntype; 249, P.rugicollis; 250, P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5064133" httpUri="https://zenodo.org/record/5064133/files/figure.png" pageId="23" pageNumber="24">241</figureCitation>
,
<figureCitation id="B0BD2A66FF8BF9741786FC20FE67FC70" box="[446,490,919,945]" captionStart="FIGURES 263264. 263" captionStartId="123.[264,383,1767,1791]" captionTargetBox="[383,1204,328,1733]" captionTargetId="figure-62@123.[383,1205,318,1763]" captionTargetPageId="123" captionText="FIGURES 263264. 263. Geographical distribution of P. longicollis (black circles), P. crustatus (white triangles), P. brevis (black squares), P. jekeli (black triangles). 264. Geographical distribution of P. jordani (black squares), P.depressus (black triangles), P. virgatus (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5064137" httpUri="https://zenodo.org/record/5064137/files/figure.png" pageId="23" pageNumber="24">263</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="609C6568FF8BF9741730FC50FE25FA70" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="283936E3FF8BF9741730FC50FB8EFC3C" blockId="23.[264,1027,998,1022]" box="[264,1027,998,1022]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="089611CDFF8BF9741730FC50FB8EFC3C" box="[264,1027,998,1022]" pageId="23" pageNumber="24">
<treatmentCitation id="A92710F2FF8BF9741730FC50FD33FC3C" author="Frieser, R." box="[264,702,998,1022]" page="233" pageId="23" pageNumber="24" year="1988">
<taxonomicName id="EF864D60FF8BF9741730FC50FD33FC3C" ID-CoL="8K9LP" authority="Frieser, 1988: 233" authorityName="Frieser" authorityPageNumber="233" authorityYear="1988" box="[264,702,998,1022]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="crustatus">
<emphasis id="1AF2EAF1FF8BF9741730FC50FE79FC3F" box="[264,500,999,1022]" italics="true" pageId="23" pageNumber="24">Ptychoderes crustatus</emphasis>
<bibRefCitation id="4C174B12FF8BF97417C2FC51FD33FC3C" author="Frieser, R." box="[506,702,998,1021]" pageId="23" pageNumber="24" pagination="233 - 234" refId="ref63164" refString="Frieser, R. (1988) Eine neue Ptychoderes- spezies aus Guyane Francaise (Coleoptera, Anthribidae). Revue francaise d'Entomologie, (N. S.), 10 (3), 233 - 234." type="journal article" year="1988">Frieser, 1988: 233</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="A92710F2FF8BF97414F1FC51FC33FC3C" author="Rheinheimer, J." box="[713,958,998,1021]" page="10" pageId="23" pageNumber="24" year="2004">
<bibRefCitation id="4C174B12FF8BF97414F1FC51FC33FC3C" author="Rheinheimer, J." box="[713,958,998,1021]" pageId="23" pageNumber="24" pagination="3 - 242" refId="ref64865" refString="Rheinheimer, J. (2004) Illustrater Katalog und Bibliographie der Anthribidae der Welt (Insecta: Coleoptera). Mitteilungen des Entomologischen Vereins Stuttgart, 39 (1 / 2), 3 - 242." type="journal article" year="2004">Rheinheimer, 2004: 10</bibRefCitation>
</treatmentCitation>
(cat.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="283936E3FF8BF9741730FB98FE25FA70" blockId="23.[264,1323,1071,1897]" pageId="23" pageNumber="24">
Male. Vestiture (
<figureCitation id="B0BD2A66FF8BF97417EDFB98FDCFFB88" box="[469,578,1071,1097]" captionStart="FIGURES 208214" captionStartId="117.[264,382,1545,1569]" captionTargetBox="[293,1285,343,1521]" captionTargetId="figure-64@117.[292,1294,326,1521]" captionTargetPageId="117" captionText="FIGURES 208214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes &amp; Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male." figureDoi="http://doi.org/10.5281/zenodo.5064125" httpUri="https://zenodo.org/record/5064125/files/figure.png" pageId="23" pageNumber="24">Fig. 208</figureCitation>
): pale dorsal vitta with dense whitish to clay­grayish scales, intermingled with light brown scales, decumbent and denser on rostrum, pronotum and elytral interstriae 12 and 4. Pygidium with predominance of brownish scales. Ventral body surface and legs with appressed, dense yellowish­white scales, longer, thicker, conspicuously denser on disk of ventrites IIIV. Sides of prosternum, below lateral carina, with a vitta of scales concolorous with those of pronotum. Sides of mesosternum with a narrow, glabrous vitta. Mesepimera, episterna and sides of metasternum with deep setigerous punctures, each provided with a brownish scale; these punctures shallow at sides of the ventrites. Outer surface of meso­ and metafemora with two spots of blackishbrown scales.
</paragraph>
</subSubSection>
<subSubSection id="609C6568FF8BF97A1700FA08FE6BFB6F" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="description">
<paragraph id="283936E3FF8BF9741700FA08FB2DF960" blockId="23.[264,1323,1071,1897]" pageId="23" pageNumber="24">Rostrum 1.5 times as long as wide across base. Distance between scrobe and eyes about half maximum width of eye. Sides looking tricarinate due the careniform upper margin of scrobe; carinae short, oblique, lower carina slightly longer. Scrobes careniform at dorsal and basal margins; extension deep, narrow with a deep distal pit. Mentum sparsely, moderately finely punctate; strongly impressed at sides and shallowly grooved at apical margin. Gula with sparse, fine punctures. Frons with prominent carinae.</paragraph>
<paragraph id="283936E3FF8BF9741700F918FEE9F8A8" blockId="23.[264,1323,1071,1897]" pageId="23" pageNumber="24">
Antennae short (
<figureCitation id="B0BD2A66FF8BF974143FF918FDF8F908" box="[519,629,1711,1737]" captionStart="FIGURES 208214" captionStartId="117.[264,382,1545,1569]" captionTargetBox="[293,1285,343,1521]" captionTargetId="figure-64@117.[292,1294,326,1521]" captionTargetPageId="117" captionText="FIGURES 208214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes &amp; Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male." figureDoi="http://doi.org/10.5281/zenodo.5064125" httpUri="https://zenodo.org/record/5064125/files/figure.png" pageId="23" pageNumber="24">Fig. 208</figureCitation>
), exceeding antebasal carinae of prothorax. Segment II thickened, slightly longer than III; IIIIV subequal in length; VVII subequal, shorter than IIIIV; VIII half as long as IX; X slightly shorter than IX and half as long as XI. (II = 0.40; III = 0.30; IV = 0.30; V = 0.20; VI = 0.20; VII = 0.20; VIII = 0.15; IX = 0.30; X = 0.25; XI = 0.40).
</paragraph>
<paragraph id="283936E3FF84F97B1700FEABFCC7FBC7" blockId="24.[264,1324,284,1790]" pageId="24" pageNumber="25">
Prothorax (
<figureCitation id="B0BD2A66FF84F97B1785FEABFDBCFEF7" box="[445,561,284,310]" captionStart="FIGURES 208214" captionStartId="117.[264,382,1545,1569]" captionTargetBox="[293,1285,343,1521]" captionTargetId="figure-64@117.[292,1294,326,1521]" captionTargetPageId="117" captionText="FIGURES 208214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes &amp; Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male." figureDoi="http://doi.org/10.5281/zenodo.5064125" httpUri="https://zenodo.org/record/5064125/files/figure.png" pageId="24" pageNumber="25">Figs. 208</figureCitation>
,
<figureCitation id="B0BD2A66FF84F97B1406FEABFDE6FEF7" box="[574,619,284,310]" captionStart="FIGURES 239250" captionStartId="121.[264,383,1756,1780]" captionTargetBox="[309,1279,326,1725]" captionTargetId="figure-77@121.[309,1279,326,1725]" captionTargetPageId="121" captionText="FIGURES 239250. Pronotum. 239, Tribotropis prasinata; 240, Unanthribus grandis; 241, Ptychoderes crustatus; 242, P. brevis, female holotype; 243, P. longicollis, holotype; 244, P. jordani, male holotype; 245, P. magnus, male holotype; 246, P. depressus; 247, P. virgatus; 248, P. bivittatus, male syntype; 249, P.rugicollis; 250, P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5064133" httpUri="https://zenodo.org/record/5064133/files/figure.png" pageId="24" pageNumber="25">241</figureCitation>
) slightly wider than long, almost rounded at sides, weakly converging anteriorly from antebasal carina, anterior margin conspicuously narrower. Pronotal depression deep, divided by a prominent, more or less transverse tubercle reaching lateral margins of depression. Rugae more prominent and elongate on median tubercle, less evident at sides near lateral carina, and poorly conspicuous or even missing inside depression. Antebasal carina interrupted at middle (at least twice width of carina), almost to barely oblique towards lateral carina. Lateral carina almost straight (
<figureCitation id="B0BD2A66FF84F97B128BFDBBFAAAFDE7" box="[1203,1319,524,550]" captionStart="FIGURES 208214" captionStartId="117.[264,382,1545,1569]" captionTargetBox="[293,1285,343,1521]" captionTargetId="figure-64@117.[292,1294,326,1521]" captionTargetPageId="117" captionText="FIGURES 208214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes &amp; Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male." figureDoi="http://doi.org/10.5281/zenodo.5064125" httpUri="https://zenodo.org/record/5064125/files/figure.png" pageId="24" pageNumber="25">Figs. 208</figureCitation>
,
<figureCitation id="B0BD2A66FF84F97B1730FD83FEB5FD8F" box="[264,312,564,590]" captionStart="FIGURES 239250" captionStartId="121.[264,383,1756,1780]" captionTargetBox="[309,1279,326,1725]" captionTargetId="figure-77@121.[309,1279,326,1725]" captionTargetPageId="121" captionText="FIGURES 239250. Pronotum. 239, Tribotropis prasinata; 240, Unanthribus grandis; 241, Ptychoderes crustatus; 242, P. brevis, female holotype; 243, P. longicollis, holotype; 244, P. jordani, male holotype; 245, P. magnus, male holotype; 246, P. depressus; 247, P. virgatus; 248, P. bivittatus, male syntype; 249, P.rugicollis; 250, P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5064133" httpUri="https://zenodo.org/record/5064133/files/figure.png" pageId="24" pageNumber="25">241</figureCitation>
). Secondary carina almost straight, widely interrupted at middle, not reaching lateral carina. Laterobasal carina reaching posterior margin of prothorax. Prosternum almost flat, with dense, fine punctures intermingles with scarce, coarse, shallow punctures; antecoxal furrow shallow, sinuous, with a very small prosternal projection at middle. Mesosternum with moderately coarse, shallow, irregularly sparse punctures, and an irregular row of coarse punctures near the intercoxal process. Intercoxal process of mesosternum impressed apically, not swollen at lateral and apical margins, with evident lateroapical projections. Elytra 1.8 times longer than wide across humeri, subconvex, transversely depressed before middle; basal gibbosities large and prominent; surface rugose from base to behind middle, more so at base and near suture. Sides subparallel at basal half, slightly wider than humeri behind middle. Apex almost rounded without anteapical tubercles, interstriae not swollen. Wings without radial cell.
</paragraph>
<paragraph id="283936E3FF84F97B1700FBA3FCCBFB67" blockId="24.[264,1324,284,1790]" pageId="24" pageNumber="25">Abdomen. Ventrites short; ventrite I convex, sex patch absent; groove of the intercoxal process line shaped; disk of ventrites IIIV strongly depressed, clothed with dense, long, thickened pilosity. Ventrite V emarginate at apex. Pygidium longer than wide, depressed, with a short groove to fit elytra, rounded at apex.</paragraph>
<paragraph id="283936E3FF84F97B1700FB03FCFFF947" blockId="24.[264,1324,284,1790]" pageId="24" pageNumber="25">Female. Rostrum shorter, almost as long as wide across base. Antennae shorter, exceeding the middle of prothorax; segment II longest; IIIIV subequal; VVIII decreasing in length; IX twice as long as VIII; XI one third longer than X. (II = 0.4; III = 0.3; IV = 0.3; V = 0.3; VI = 0.2; VII = 0.2; VIII = 0.2; IX = 0.4; X = 0.2; XI = 0.4). Ventrites weakly convex; II and III subequal in length; IV slightly shorter than III, both without long, thick pilosity; ventrite V truncate at apex. Pygidium wider than long, somewhat convex on disk, rounded­truncate at apex. Terminalia: tergite VIII as long as wide, feebly convergent apically, truncate at apex; sternite VIII feebly shorter than apodeme, truncate at apex. Ovipositor with four teeth: two distal, conchoidal, developed and similarly sized; one proximal, dorsolateral, moderately developed, and one proximal, ventrolateral, minute; median rods thickened, rounded at proximal margin; median ventral lobe bilobated; bursa copulatrix lacking ventral lobe.</paragraph>
<paragraph id="283936E3FF84F97A1700F923FE24FEF4" blockId="24.[264,1324,284,1790]" lastBlockId="25.[264,1323,284,510]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
Measurements, in mm, male/female, respectively. TL = 7.4/10.511.3; RL = 1.4/ 1.41.5; RAW = 1.4/1.31.7; RBW = 0.9/1.31.4; HW = 1.2/1.82.0; DSO = 0.4/0.4; MLO = 0.8/1.0; PL = 2.2/2.73.2; PW = 2.4/2.73.2; EL = 4.6/6.86.9; EW = 2.5/3.33.5.
<emphasis id="1AF2EAF1FF85F97A1730FEABFE24FEF4" box="[264,425,284,309]" italics="true" pageId="25" pageNumber="26">
<typeStatus id="F73D8841FF85F97A1730FEABFEB2FEF4" box="[264,319,284,309]" pageId="25" pageNumber="26">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="283936E3FF85F97A1700FEF3FDA9FE3F" blockId="25.[264,1323,284,510]" pageId="25" pageNumber="26">
<materialsCitation id="98EE3CBEFF85F97A1700FEF3FC56FE47" collectionCode="ZSMA, R" collectorName="Female" country="French Guiana" location="Frieser" pageId="25" pageNumber="26" specimenCount="2" typeStatus="paratype">
<collectorName id="85735335FF85F97A1700FEF3FE03FE9F" box="[312,398,324,350]" pageId="25" pageNumber="26">Female</collectorName>
<typeStatus id="F73D8841FF85F97A17ADFEF3FE74FE9F" box="[405,505,324,350]" pageId="25" pageNumber="26" type="paratype">paratype</typeStatus>
(
<collectionCode id="4E97AE26FF85F97A1431FEF3FDEDFE9F" box="[521,608,324,350]" pageId="25" pageNumber="26">ZSMA</collectionCode>
), examined, labelled as follows: 1)
<taxonomicName id="EF864D60FF85F97A15C7FEF3FA8CFE9C" authority="Mermudes &amp; Napp, 2006" authorityName="Mermudes &amp; Napp" authorityYear="2006" box="[1023,1281,324,349]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crustatus" status="sp. nov.">
<emphasis id="1AF2EAF1FF85F97A15C7FEF3FA8CFE9C" box="[1023,1281,324,349]" italics="true" pageId="25" pageNumber="26">Ptychoderes crustatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="01C1578AFF85F97A1331FEF3FE92FE47" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
,
<typeStatus id="F73D8841FF85F97A1716FEDBFE1AFE47" box="[302,407,364,390]" pageId="25" pageNumber="26" type="paratype">Paratype</typeStatus>
, [red]; 2) det.
<collectionCode id="4E97AE26FF85F97A1479FEDBFDD5FE47" box="[577,600,364,390]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="26">R</collectionCode>
.
<location id="2D596038FF85F97A145DFEDBFD35FE47" LSID="urn:lsid:plazi:treatment:A02F87F5FF8BF97A1730FCC3FAA4FB3E:2D596038FF85F97A145DFEDBFD35FE47" box="[613,696,364,390]" country="French Guiana" name="Frieser" pageId="25" pageNumber="26">Frieser</location>
, 1987 [pink]; 3)
<collectingCountry id="50917673FF85F97A15B8FEDBFC56FE47" box="[896,987,364,390]" name="French Guiana" pageId="25" pageNumber="26">Guyane</collectingCountry>
</materialsCitation>
<materialsCitation id="98EE3CBEFF85F97A15DCFEDBFDADFE3F" collectionCode="MNHN" collectorName="Francaise &amp; Fevrier &amp; G. Tavakilian &amp; Route de Kaw, PK &amp; Zool. Staatsslg. &amp; Munchen &amp; Male" country="French Guiana" location="Route de Kaw" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<collectorName id="85735335FF85F97A15DCFEDBFBDAFE47" box="[996,1111,364,390]" pageId="25" pageNumber="26">Française</collectorName>
,
<collectorName id="85735335FF85F97A125CFEDBFB37FE47" box="[1124,1210,364,390]" pageId="25" pageNumber="26">Février</collectorName>
, 1983,
<collectorName id="85735335FF85F97A1329FEDBFE0EFE6F" pageId="25" pageNumber="26">G. Tavakilian</collectorName>
leg. [white]; 4)
<collectorName id="85735335FF85F97A1476FE23FCBFFE6F" box="[590,818,404,430]" pageId="25" pageNumber="26">
<location id="2D596038FF85F97A1476FE23FD70FE6F" LSID="urn:lsid:plazi:treatment:A02F87F5FF8BF97A1730FCC3FAA4FB3E:2D596038FF85F97A1476FE23FD70FE6F" box="[590,765,404,430]" country="French Guiana" name="Route de Kaw" pageId="25" pageNumber="26">Route de Kaw</location>
, PK
</collectorName>
33 [white]; 5)
<collectorName id="85735335FF85F97A15C8FE23FB26FE6F" box="[1008,1195,404,430]" pageId="25" pageNumber="26">Zool. Staatsslg.</collectorName>
,
<collectorName id="85735335FF85F97A1282FE23FAA4FE6F" box="[1210,1321,404,430]" pageId="25" pageNumber="26">München</collectorName>
[blue].
<collectorName id="85735335FF85F97A175CFE0BFE2DFE17" box="[356,416,444,470]" pageId="25" pageNumber="26">Male</collectorName>
<typeStatus id="F73D8841FF85F97A1797FE0BFD95FE17" box="[431,536,444,470]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, not examined, deposited in the
<collectionCode id="4E97AE26FF85F97A1585FE0BFB9AFE17" box="[957,1047,444,470]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="26">MNHN</collectionCode>
(
<bibRefCitation id="4C174B12FF85F97A1215FE0BFB59FE17" author="Frieser, R." box="[1069,1236,444,470]" pageId="25" pageNumber="26" pagination="233 - 234" refId="ref63164" refString="Frieser, R. (1988) Eine neue Ptychoderes- spezies aus Guyane Francaise (Coleoptera, Anthribidae). Revue francaise d'Entomologie, (N. S.), 10 (3), 233 - 234." type="journal article" year="1988">Frieser, 1988</bibRefCitation>
). Type locality:
<collectingCountry id="50917673FF85F97A1757FE53FDADFE3F" box="[367,544,484,510]" name="French Guiana" pageId="25" pageNumber="26">French Guiana</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="283936E3FF85F97A1730FD83FEE3FD8C" blockId="25.[264,1323,564,999]" box="[264,366,564,589]" pageId="25" pageNumber="26">
<emphasis id="1AF2EAF1FF85F97A1730FD83FEE3FD8C" box="[264,366,564,589]" italics="true" pageId="25" pageNumber="26">Remarks</emphasis>
</paragraph>
<paragraph id="283936E3FF85F97A1700FDEBFEE9FC26" blockId="25.[264,1323,564,999]" pageId="25" pageNumber="26">
<taxonomicName id="EF864D60FF85F97A1700FDEBFE4DFDB4" authorityName="Mermudes &amp; Napp" authorityYear="2006" box="[312,448,604,629]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crustatus">
<emphasis id="1AF2EAF1FF85F97A1700FDEBFE4DFDB4" box="[312,448,604,629]" italics="true" pageId="25" pageNumber="26">P. crustatus</emphasis>
</taxonomicName>
is the sister group of the clade
<taxonomicName id="EF864D60FF85F97A150EFDEBFC17FDB4" authorityName="Jordan" authorityYear="1894" box="[822,922,604,629]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF85F97A150EFDEBFC17FDB4" box="[822,922,604,629]" italics="true" pageId="25" pageNumber="26">P. brevis</emphasis>
</taxonomicName>
+
<taxonomicName id="EF864D60FF85F97A1593FDEBFB85FDB4" authorityName="Mermudes &amp; Napp" authorityYear="2004" box="[939,1032,604,629]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="jekeli">
<emphasis id="1AF2EAF1FF85F97A1593FDEBFB85FDB4" box="[939,1032,604,629]" italics="true" pageId="25" pageNumber="26">P. jekeli</emphasis>
</taxonomicName>
by the synapomorphies: sides of prosternum with a vitta of scales concolorous with those of pronotum (4
<subScript id="B40234A6FF85F97A12FDFD23FB40FD63" attach="right" box="[1221,1229,660,674]" fontSize="6" pageId="25" pageNumber="26">3</subScript>
); elytra transversely impressed before middle (29
<subScript id="B40234A6FF85F97A1538FD08FC85FD0C" attach="none" box="[768,776,703,717]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
); and pygidium elongate, depressed, weakly convergent at sides (32
<subScript id="B40234A6FF85F97A1423FD5DFDAEFD39" attach="none" box="[539,547,746,760]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
).
<taxonomicName id="EF864D60FF85F97A1404FD6DFD4AFD32" authorityName="Mermudes &amp; Napp" authorityYear="2006" box="[572,711,730,755]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crustatus">
<emphasis id="1AF2EAF1FF85F97A1404FD6DFD4AFD32" box="[572,711,730,755]" italics="true" pageId="25" pageNumber="26">P. crustatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="EF864D60FF85F97A155DFD6DFC40FD32" authorityName="Jordan" authorityYear="1894" box="[869,973,730,755]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF85F97A155DFD6DFC40FD32" box="[869,973,730,755]" italics="true" pageId="25" pageNumber="26">P. brevis</emphasis>
</taxonomicName>
and
<taxonomicName id="EF864D60FF85F97A123DFD6DFBEEFD32" authorityName="Mermudes &amp; Napp" authorityYear="2004" box="[1029,1123,730,755]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="jekeli">
<emphasis id="1AF2EAF1FF85F97A123DFD6DFBEEFD32" box="[1029,1123,730,755]" italics="true" pageId="25" pageNumber="26">P. jekeli</emphasis>
</taxonomicName>
, by the vestiture with whitish and light­brown scales, almost rounded prothorax, and prosternal furrow shallow and sinuous. In
<taxonomicName id="EF864D60FF85F97A141BFC9AFD07FC87" authorityName="Jordan" authorityYear="1894" box="[547,650,813,838]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF85F97A141BFC9AFD07FC87" box="[547,650,813,838]" italics="true" pageId="25" pageNumber="26">P. brevis</emphasis>
</taxonomicName>
and
<taxonomicName id="EF864D60FF85F97A1487FC9AFC97FC87" authorityName="Mermudes &amp; Napp" authorityYear="2004" box="[703,794,813,838]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="jekeli">
<emphasis id="1AF2EAF1FF85F97A1487FC9AFC97FC87" box="[703,794,813,838]" italics="true" pageId="25" pageNumber="26">P. jekeli</emphasis>
</taxonomicName>
, the vestiture is formed by green and whitish scales, the prothorax weakly convergent apically from the antebasal carina, and the prosternal furrow deep and straight. In addition, the males of
<taxonomicName id="EF864D60FF85F97A15E5FCCAFBEBFC57" authorityName="Mermudes &amp; Napp" authorityYear="2006" box="[989,1126,893,918]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crustatus">
<emphasis id="1AF2EAF1FF85F97A15E5FCCAFBEBFC57" box="[989,1126,893,918]" italics="true" pageId="25" pageNumber="26">P. crustatus</emphasis>
</taxonomicName>
have the disk of ventrites IIIV strongly depressed, with long, dense, thick hairs, autapomorphy of this species.
</paragraph>
<paragraph id="283936E3FF85F97A1730FBAAFC09FB9E" blockId="25.[264,900,1053,1119]" pageId="25" pageNumber="26">
<emphasis id="1AF2EAF1FF85F97A1730FBAAFD98FBF7" box="[264,533,1053,1078]" italics="true" pageId="25" pageNumber="26">
Distribution (
<figureCitation id="B0BD2A66FF85F97A1790FBAAFD80FBF7" box="[424,525,1053,1078]" captionStart="FIGURES 263264. 263" captionStartId="123.[264,383,1767,1791]" captionTargetBox="[383,1204,328,1733]" captionTargetId="figure-62@123.[383,1205,318,1763]" captionTargetPageId="123" captionText="FIGURES 263264. 263. Geographical distribution of P. longicollis (black circles), P. crustatus (white triangles), P. brevis (black squares), P. jekeli (black triangles). 264. Geographical distribution of P. jordani (black squares), P.depressus (black triangles), P. virgatus (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5064137" httpUri="https://zenodo.org/record/5064137/files/figure.png" pageId="25" pageNumber="26">Fig. 263</figureCitation>
)
</emphasis>
<collectingCountry id="50917673FF85F97A1700FBF2FDAAFB9E" box="[312,551,1093,1119]" name="French Guiana" pageId="25" pageNumber="26">FRENCH GUIANA</collectingCountry>
and
<collectingCountry id="50917673FF85F97A1467FBF2FD44FB9E" box="[607,713,1093,1119]" name="Brazil" pageId="25" pageNumber="26">BRAZIL</collectingCountry>
(
<collectingRegion id="EA42F801FF85F97A14E0FBF2FCF5FB9E" box="[728,888,1093,1119]" country="Brazil" name="Mato Grosso" pageId="25" pageNumber="26">Mato Grosso</collectingRegion>
).
</paragraph>
<paragraph id="283936E3FF85F97A1730FB22FE6BFB6F" blockId="25.[264,1323,1173,1279]" box="[264,486,1173,1198]" pageId="25" pageNumber="26">
<emphasis id="1AF2EAF1FF85F97A1730FB22FE6BFB6F" box="[264,486,1173,1198]" italics="true" pageId="25" pageNumber="26">Material examined</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="609C6568FF85F97A1700FB0AFAA4FB3E" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="283936E3FF85F97A1700FB0AFAA4FB3E" blockId="25.[264,1323,1173,1279]" pageId="25" pageNumber="26">
<materialsCitation id="98EE3CBEFF85F97A1700FB0AFBC6FB16" box="[312,1099,1213,1239]" collectionCode="IV" collectorName="de Kaw" country="French Guiana" location="Route de Kaw" pageId="25" pageNumber="26" specimenCount="1">
<collectingCountry id="50917673FF85F97A1700FB0AFDA3FB16" box="[312,558,1213,1239]" name="French Guiana" pageId="25" pageNumber="26">FRENCH GUIANA</collectingCountry>
. Cayenne (
<location id="2D596038FF85F97A148DFB0AFCE8FB16" LSID="urn:lsid:plazi:treatment:A02F87F5FF8BF97A1730FCC3FAA4FB3E:2D596038FF85F97A148DFB0AFCE8FB16" box="[693,869,1213,1239]" country="French Guiana" name="Route de Kaw" pageId="25" pageNumber="26">
Route
<collectorName id="85735335FF85F97A153FFB0AFCE8FB16" box="[775,869,1213,1239]" pageId="25" pageNumber="26">de Kaw</collectorName>
</location>
, 33 Km), 1 M,
<date id="5C381023FF85F97A1212FB0AFBC6FB16" box="[1066,1099,1213,1239]" pageId="25" pageNumber="26" value="1993-04">
<collectionCode id="4E97AE26FF85F97A1212FB0AFBC6FB16" box="[1066,1099,1213,1239]" pageId="25" pageNumber="26">IV</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="98EE3CBEFF85F97A1272FB0AFE2CFB3E" collectionCode="CPMT" collectorName="Marek-Seidl" country="French Guiana" pageId="25" pageNumber="26" specimenCount="1">
.1993,
<collectorName id="85735335FF85F97A12A1FB0AFAA6FB16" box="[1177,1323,1213,1239]" pageId="25" pageNumber="26">Marek­Seidl</collectorName>
leg. (
<collectionCode id="4E97AE26FF85F97A177BFB52FE14FB3E" box="[323,409,1253,1279]" pageId="25" pageNumber="26">CPMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="98EE3CBEFF85F97A1794FB52FAA8FB3E" box="[428,1317,1253,1279]" collectingDate="1974-10" collectionCode="MNRJ" collectorName="Silva" country="Brazil" location="Sinop" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Mato Grosso">
<collectingCountry id="50917673FF85F97A1794FB52FD97FB3E" box="[428,538,1253,1279]" name="Brazil" pageId="25" pageNumber="26">BRAZIL</collectingCountry>
.
<emphasis id="1AF2EAF1FF85F97A141DFB52FD30FB3F" box="[549,701,1253,1278]" italics="true" pageId="25" pageNumber="26">
<collectingRegion id="EA42F801FF85F97A141DFB52FD30FB3F" box="[549,701,1253,1278]" country="Brazil" name="Mato Grosso" pageId="25" pageNumber="26">Mato Grosso</collectingRegion>
</emphasis>
:
<location id="2D596038FF85F97A14F5FB52FC9CFB3E" LSID="urn:lsid:plazi:treatment:A02F87F5FF8BF97A1730FCC3FAA4FB3E:2D596038FF85F97A14F5FB52FC9CFB3E" box="[717,785,1253,1279]" country="Brazil" name="Sinop" pageId="25" pageNumber="26" stateProvince="Mato Grosso">Sinop</location>
(Faz. Teles), 1 F,
<date id="5C381023FF85F97A15DCFB52FBB3FB3E" box="[996,1086,1253,1279]" pageId="25" pageNumber="26" value="1974-10">
<collectingDate id="4C7CE9CBFF85F97A15DCFB52FBB3FB3E" box="[996,1086,1253,1279]" pageId="25" pageNumber="26" value="1974-10">X.1974</collectingDate>
</date>
,
<collectorName id="85735335FF85F97A1272FB52FB0BFB3E" box="[1098,1158,1253,1279]" pageId="25" pageNumber="26">Silva</collectorName>
leg. (
<collectionCode id="4E97AE26FF85F97A12F0FB52FA90FB3E" box="[1224,1309,1253,1279]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="26">MNRJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>