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<document id="79EF7EFEB8297976AE8C530F2EF8EEBC" ID-ISSN="1536-9307" ID-ZooBank="C9EFE1F7-D1E6-4820-A5A7-5FE940A9033" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638257832623" checkinUser="felipe" docAuthor="Kovařík, František, Ranawana, Kithsiri B., Sanjeewa Jayarathne, V. A., Karunarathna, Sanjaya &amp; Ullrich, Alexander" docDate="2018" docId="9A623F52FFC1CA74FC44F97C05B0FBBB" docLanguage="en" docName="Euscorpius.258.1-5.pdf" docOrigin="Euscorpius 258" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Liocheles australasiae" docType="treatment" docVersion="3" lastPageNumber="5" masterDocId="665B472AFFC3CA72FFC4FFE3072CFF82" masterDocTitle="Scorpions of Sri Lanka (Arachnida, Scorpiones). Part II. Family Hormuridae" masterLastPageNumber="5" masterPageNumber="1" pageNumber="1" updateTime="1699436158509" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="68F25FEC7856BAF36C1828F32728EAD4">Scorpions of Sri Lanka (Arachnida, Scorpiones). Part II. Family Hormuridae</mods:title>
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<mods:namePart id="462080B09086DB1B43BFE09400177033">Kovařík, František</mods:namePart>
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<mods:namePart id="21CC55893CAA14F8FD5F416F9F9F1D8E">Sanjeewa Jayarathne, V. A.</mods:namePart>
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<mods:namePart id="10EE698788D151F4823182E3BD049079">Karunarathna, Sanjaya</mods:namePart>
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<mods:title id="D87088CB82654675FC00A14D7CDDABFC">Euscorpius</mods:title>
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<mods:date id="BAC578C636F58EF55FED5353716F90C7">2018</mods:date>
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<treatment id="9A623F52FFC1CA74FC44F97C05B0FBBB" ID-DOI="http://doi.org/10.5281/zenodo.5741206" ID-Zenodo-Dep="5741206" LSID="urn:lsid:plazi:treatment:9A623F52FFC1CA74FC44F97C05B0FBBB" httpUri="http://treatment.plazi.org/id/9A623F52FFC1CA74FC44F97C05B0FBBB" lastPageId="6" lastPageNumber="5" pageId="2" pageNumber="1">
<subSubSection id="5AD1DDCFFFC1CA70FC44F97C0272F938" box="[896,1374,1695,1722]" pageId="2" pageNumber="1" type="nomenclature">
<paragraph id="12748E44FFC1CA70FC44F97C0272F938" blockId="2.[896,1374,1695,1752]" box="[896,1374,1695,1722]" pageId="2" pageNumber="1">
<heading id="493C3928FFC1CA70FC44F97C0272F938" box="[896,1374,1695,1722]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="1" reason="4">
<taxonomicName id="D5CBF5C7FFC1CA70FC44F97C0272F938" ID-CoL="3V5CL" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[896,1374,1695,1722]" class="Arachnida" family="Hormuridae" genus="Liocheles" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="australasiae">
<emphasis id="20BF5256FFC1CA70FC44F97C03A2F938" bold="true" box="[896,1166,1695,1722]" italics="true" pageId="2" pageNumber="1">Liocheles australasiae</emphasis>
(Fabricius, 1775)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5AD1DDCFFFC1CA70FBEAF922039DF95A" box="[1070,1201,1729,1752]" pageId="2" pageNumber="1" type="description">
<paragraph id="12748E44FFC1CA70FBEAF922039DF95A" blockId="2.[896,1374,1695,1752]" box="[1070,1201,1729,1752]" pageId="2" pageNumber="1">
(
<figureCitation id="8AF092C1FFC1CA70FBF1F9220384F95A" box="[1077,1192,1729,1752]" captionStart-0="Figures 19" captionStart-1="Figures 1012" captionStart-2="Figures 1314" captionStartId-0="3.[192,278,1837,1861]" captionStartId-1="4.[192,278,1866,1890]" captionStartId-2="5.[192,278,1809,1833]" captionTargetBox-0="[223,1399,170,1804]" captionTargetBox-1="[191,1440,165,1849]" captionTargetBox-2="[191,1440,191,1780]" captionTargetId-0="figure-6@3.[192,1439,168,1819]" captionTargetId-1="figure-13@4.[192,1439,166,1848]" captionTargetId-2="figure-6@5.[192,1439,192,1779]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figures 19: Liocheles australasiae from Sri Lanka, pedipalp segments. Chela dorsal (1), external (2) and ventrointernal (3). Femur dorsal (4), external (5) and ventral (6). Patella dorsal (7) and ventral (8). Movable finger (9). Trichobothrial pattern is indicated." captionText-1="Figures 1012: Liocheles australasiae from Sri Lanka. Chelicerae, carapace and tergite I (10). Coxosternal area and sternite III (11). Female in vivo habitus (12)." captionText-2="Figures 1314: Liocheles australasiae, the Sri Lankan locality." pageId="2" pageNumber="1">Figs. 114</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5AD1DDCFFFC1CA70FCFAF8E6038FF8DB" pageId="2" pageNumber="1" type="reference_group">
<paragraph id="12748E44FFC1CA70FCFAF8E6038FF8DB" blockId="2.[830,1440,1797,1881]" pageId="2" pageNumber="1">
<treatmentCitationGroup id="32DBA96AFFC1CA70FCFAF8E6038FF8DB" pageId="2" pageNumber="1">
<treatmentCitation id="936AA855FFC1CA70FCFAF8E6021FF89E" author="FET" box="[830,1331,1797,1820]" page="397" pageId="2" pageNumber="1" year="2000">
<taxonomicName id="D5CBF5C7FFC1CA70FCFAF8E6021FF89E" ID-CoL="3V5CL" authority=": Fet, 2000: 395 - 397" authorityName=": Fet" authorityPageNumber="395 - 397" authorityYear="2000" box="[830,1331,1797,1820]" class="Arachnida" family="Hormuridae" genus="Liocheles" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="australasiae">
<emphasis id="20BF5256FFC1CA70FCFAF8E60315F89E" box="[830,1081,1797,1820]" italics="true" pageId="2" pageNumber="1">Liocheles australasiae</emphasis>
:
<bibRefCitation id="765AF3B5FFC1CA70FB94F8E6021FF89E" author="FET" box="[1104,1331,1797,1820]" pageId="2" pageNumber="1" refId="ref1080" refString="FET, V. 2000. Family Ischnuridae Simon, 1879. Pp. 383 - 408 in Fet, V., W. D. Sissom, G. Lowe &amp; M. E. Braunwalder. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." type="book" year="2000">Fet, 2000: 395397</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part, complete reference and synonymy list until 2000); Monod &amp; Prendini, 2014: 2526.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="46B4DECCFFC0CA71FF04F8CE0632F8F5" pageId="3" pageNumber="2" startId="3.[192,278,1837,1861]" targetBox="[223,1399,170,1804]" targetPageId="3">
<paragraph id="12748E44FFC0CA71FF04F8CE0632F8F5" blockId="3.[192,1440,1837,1911]" pageId="3" pageNumber="2">
<emphasis id="20BF5256FFC0CA71FF04F8CE067CF8C7" bold="true" box="[192,336,1837,1861]" pageId="3" pageNumber="2">Figures 19:</emphasis>
<taxonomicName id="D5CBF5C7FFC0CA71FE93F8D3051EF8C7" authorityName=": Fet" authorityYear="2000" box="[343,562,1840,1861]" class="Arachnida" family="Hormuridae" genus="Liocheles" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="australasiae">
<emphasis id="20BF5256FFC0CA71FE93F8D3051EF8C7" box="[343,562,1840,1861]" italics="true" pageId="3" pageNumber="2">Liocheles australasiae</emphasis>
</taxonomicName>
from Sri Lanka, pedipalp segments. Chela dorsal (1), external (2) and ventrointernal (3). Femur dorsal (4), external (5) and ventral (6). Patella dorsal (7) and ventral (8). Movable finger (9). Trichobothrial pattern is indicated.
</paragraph>
</caption>
<caption id="46B4DECCFFC7CA76FF04F8A9052FF8F9" pageId="4" pageNumber="3" startId="4.[192,278,1866,1890]" targetBox="[191,1440,165,1849]" targetPageId="4">
<paragraph id="12748E44FFC7CA76FF04F8A9052FF8F9" blockId="4.[192,1440,1866,1915]" pageId="4" pageNumber="3">
<emphasis id="20BF5256FFC7CA76FF04F8A90644F8E0" bold="true" box="[192,360,1866,1890]" pageId="4" pageNumber="3">Figures 1012:</emphasis>
<taxonomicName id="D5CBF5C7FFC7CA76FEABF8AE0565F8E0" authorityName=": Fet" authorityYear="2000" box="[367,585,1869,1890]" class="Arachnida" family="Hormuridae" genus="Liocheles" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="australasiae">
<emphasis id="20BF5256FFC7CA76FEABF8AE0565F8E0" box="[367,585,1869,1890]" italics="true" pageId="4" pageNumber="3">Liocheles australasiae</emphasis>
</taxonomicName>
from Sri Lanka. Chelicerae, carapace and tergite I (10). Coxosternal area and sternite III (11). Female in vivo habitus (12).
</paragraph>
</caption>
<caption id="46B4DECCFFC6CA77FF04F8F20418F8AA" box="[192,820,1809,1833]" pageId="5" pageNumber="4" startId="5.[192,278,1809,1833]" targetBox="[191,1440,191,1780]" targetPageId="5">
<paragraph id="12748E44FFC6CA77FF04F8F20418F8AA" blockId="5.[192,820,1809,1833]" box="[192,820,1809,1833]" pageId="5" pageNumber="4">
<emphasis id="20BF5256FFC6CA77FF04F8F20644F8AB" bold="true" box="[192,360,1809,1833]" pageId="5" pageNumber="4">Figures 1314:</emphasis>
<taxonomicName id="D5CBF5C7FFC6CA77FEAAF8F0056BF8AA" authorityName=": Fet" authorityYear="2000" box="[366,583,1811,1832]" class="Arachnida" family="Hormuridae" genus="Liocheles" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="australasiae">
<emphasis id="20BF5256FFC6CA77FEAAF8F0056BF8AA" box="[366,583,1811,1832]" italics="true" pageId="5" pageNumber="4">Liocheles australasiae</emphasis>
</taxonomicName>
, the Sri Lankan locality.
</paragraph>
</caption>
<subSubSection id="5AD1DDCFFFC5CA74FF04FF24057BFDAD" pageId="6" pageNumber="5" type="materials_examined">
<paragraph id="12748E44FFC5CA74FF04FF2406C9FE99" blockId="6.[192,783,199,283]" pageId="6" pageNumber="5">
<materialsCitation id="A2A38419FFC5CA74FF04FF240682FF5C" box="[192,430,199,222]" collectionCode="LOCALITY, AND" pageId="6" pageNumber="5" specimenCount="1" typeStatus="holotype">
<typeStatus id="CD7030E6FFC5CA74FF04FF2407D4FF5C" box="[192,248,199,222]" pageId="6" pageNumber="5">TYPE</typeStatus>
<collectionCode id="74DA1681FFC5CA74FEC3FF280682FF5C" box="[263,430,203,222]" pageId="6" pageNumber="5">LOCALITY AND</collectionCode>
</materialsCitation>
<materialsCitation id="A2A38419FFC5CA74FE78FF2806CCFE99" collectionCode="REPOSITORY, BMNH" country="United Kingdom" location="The Natural History Museum" municipality="London" pageId="6" pageNumber="5" specimenCount="1" typeStatus="holotype">
<typeStatus id="CD7030E6FFC5CA74FE78FF2806DDFF5C" box="[444,497,203,222]" pageId="6" pageNumber="5">TYPE</typeStatus>
<collectionCode id="74DA1681FFC5CA74FDC4FF2805AFFF5C" box="[512,643,203,222]" pageId="6" pageNumber="5">REPOSITORY</collectionCode>
. &quot;in insulis Oceani pacifi&quot;;
<collectionCode id="74DA1681FFC5CA74FEAEFF050696FF7F" box="[362,442,230,253]" pageId="6" pageNumber="5">BMNH</collectionCode>
(
<location id="1714D89FFFC5CA74FE0DFF050427FF7F" LSID="urn:lsid:plazi:treatment:9A623F52FFC1CA74FC44F97C05B0FBBB:1714D89FFFC5CA74FE0DFF050427FF7F" box="[457,779,230,253]" country="United Kingdom" municipality="London" name="The Natural History Museum" pageId="6" pageNumber="5">The Natural History Museum</location>
,
<collectingMunicipality id="F210143EFFC5CA74FF04FEE7063AFE99" box="[192,278,260,283]" pageId="6" pageNumber="5">London</collectingMunicipality>
,
<collectingCountry id="6ADCCED4FFC5CA74FEE4FEE706F6FE99" box="[288,474,260,283]" name="United Kingdom" pageId="6" pageNumber="5">United Kingdom</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="12748E44FFC5CA74FF04FEA1057BFDAD" blockId="6.[192,783,322,559]" pageId="6" pageNumber="5">
<materialsCitation id="A2A38419FFC5CA74FF04FEA105EFFE14" collectingDate="2016-12-18" collectionCode="SRI, LANKAN, MATERIAL, EXAMINED" country="Sri Lanka" elevation="3" latitude="6.407639" location="Bentota" longLatPrecision="1" longitude="80.02478" municipality="Galle District" pageId="6" pageNumber="5" specimenCount="3" specimenCount-female="3" stateProvince="Southern">
<collectionCode id="74DA1681FFC5CA74FF04FEA107C8FEDA" box="[192,228,322,345]" country="Tanzania" httpUri="http://grbio.org/cool/z5p4-dpar" name="Serengetti Research Institute" pageId="6" pageNumber="5">SRI</collectionCode>
<collectionCode id="74DA1681FFC5CA74FF34FEA10569FEDA" box="[240,581,322,345]" pageId="6" pageNumber="5">LANKAN MATERIAL EXAMINED</collectionCode>
.
<collectingCountry id="6ADCCED4FFC5CA74FD9DFEA105E7FEDB" box="[601,715,322,345]" name="Sri Lanka" pageId="6" pageNumber="5">Sri Lanka</collectingCountry>
,
<collectingRegion id="D00F40A6FFC5CA74FD18FEA10648FEF5" country="Sri Lanka" name="Southern" pageId="6" pageNumber="5">Southern Province</collectingRegion>
,
<collectingMunicipality id="F210143EFFC5CA74FEB7FE830526FEF5" box="[371,522,352,375]" pageId="6" pageNumber="5">Galle District</collectingMunicipality>
,
<location id="1714D89FFFC5CA74FDDDFE830543FEF5" LSID="urn:lsid:plazi:treatment:9A623F52FFC1CA74FC44F97C05B0FBBB:1714D89FFFC5CA74FDDDFE830543FEF5" box="[537,623,352,375]" country="Sri Lanka" latitude="6.407639" longLatPrecision="1" longitude="80.02478" municipality="Galle District" name="Bentota" pageId="6" pageNumber="5" stateProvince="Southern">Bentota</location>
,
<geoCoordinate id="77FFE883FFC5CA74FDBAFE830423FEF5" box="[638,783,352,375]" degrees="06" direction="north" minutes="24" orientation="latitude" pageId="6" pageNumber="5" precision="1" seconds="27.5" value="6.407639">06°24'27.5&quot;N</geoCoordinate>
<geoCoordinate id="77FFE883FFC5CA74FF04FE9C067DFE14" box="[192,337,383,406]" degrees="80" direction="east" minutes="01" orientation="longitude" pageId="6" pageNumber="5" precision="1" seconds="29.2" value="80.02478">80°01'29.2&quot;E</geoCoordinate>
,
<elevation id="99E66977FFC5CA74FEA4FE9C06E5FE14" box="[352,457,383,406]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="5" unit="m" value="3.0">
<quantity id="D53323A1FFC5CA74FEA4FE9C06A7FE14" box="[352,395,383,406]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="5" unit="m" value="3.0">3 m</quantity>
a.s.l.
</elevation>
,
<date id="6675A884FFC5CA74FE1DFE9C0572FE14" box="[473,606,383,406]" pageId="6" pageNumber="5" value="2016-12-18">
<collectingDate id="7631516CFFC5CA74FE1DFE9C0572FE14" box="[473,606,383,406]" pageId="6" pageNumber="5" value="2016-12-18">18.XII.2016</collectingDate>
</date>
,
<specimenCount id="04CD45CDFFC5CA74FDA9FE9C05A2FE14" box="[621,654,383,406]" pageId="6" pageNumber="5" type="female">3♀</specimenCount>
, leg
</materialsCitation>
.
<materialsCitation id="A2A38419FFC5CA74FD15FE9C0681FE37" collectionCode="V" collectorName="A. Sanjeewa Jayarathne" country="Sri Lanka" pageId="6" pageNumber="5" specimenCount="1" stateProvince="Southern Province">
<collectionCode id="74DA1681FFC5CA74FD15FE9C05CBFE14" box="[721,743,383,406]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="5" type="Museum">V</collectionCode>
.
<collectorName id="BF3EEB92FFC5CA74FD31FE9C0681FE37" pageId="6" pageNumber="5">A. Sanjeewa Jayarathne</collectorName>
</materialsCitation>
,
<collectionCode id="74DA1681FFC5CA74FE07FE7D052AFE37" box="[451,518,414,437]" pageId="6" pageNumber="5">FKCP</collectionCode>
(the first author collection). The specimens were found inside a stone wall and under old bark of branches and in dry leaves on the land near a domestic tea cultivation field; also under the flower pots in home garden.
</paragraph>
</subSubSection>
<subSubSection id="5AD1DDCFFFC5CA74FF04FDB505B0FBBB" pageId="6" pageNumber="5" type="diagnosis">
<paragraph id="12748E44FFC5CA74FF04FDB505B0FBBB" blockId="6.[192,802,598,1081]" pageId="6" pageNumber="5">
DIAGNOSIS. Total length
<quantity id="D53323A1FFC5CA74FE0AFDB50568FDEF" box="[462,580,598,621]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.6" metricValueMin="2.2" pageId="6" pageNumber="5" unit="mm" value="29.0" valueMax="36.0" valueMin="22.0">2236 mm</quantity>
. Patella of pedipalps with 3 ventral trichobothria (
<figureCitation id="8AF092C1FFC5CA74FE33FD970515FD09" box="[503,569,628,651]" captionStart="Figures 19" captionStartId="3.[192,278,1837,1861]" captionTargetBox="[223,1399,170,1804]" captionTargetId="figure-6@3.[192,1439,168,1819]" captionTargetPageId="3" captionText="Figures 19: Liocheles australasiae from Sri Lanka, pedipalp segments. Chela dorsal (1), external (2) and ventrointernal (3). Femur dorsal (4), external (5) and ventral (6). Patella dorsal (7) and ventral (8). Movable finger (9). Trichobothrial pattern is indicated." pageId="6" pageNumber="5">Fig. 3</figureCitation>
); trichobothrium
<emphasis id="20BF5256FFC5CA74FD33FD97040EFD09" box="[759,802,628,651]" italics="true" pageId="6" pageNumber="5">Eb3</emphasis>
located in base of chela near trichobothria
<emphasis id="20BF5256FFC5CA74FD4DFD700586FD28" box="[649,682,659,682]" italics="true" pageId="6" pageNumber="5">Db</emphasis>
,
<emphasis id="20BF5256FFC5CA74FD73FD7005CEFD28" box="[695,738,659,682]" italics="true" pageId="6" pageNumber="5">Eb1</emphasis>
,
<emphasis id="20BF5256FFC5CA74FD34FD700437FD28" box="[752,795,659,682]" italics="true" pageId="6" pageNumber="5">Eb2</emphasis>
, and
<emphasis id="20BF5256FFC5CA74FF35FD510635FD4B" box="[241,281,690,713]" italics="true" pageId="6" pageNumber="5">Esb</emphasis>
(
<figureCitation id="8AF092C1FFC5CA74FEEFFD51065CFD4B" box="[299,368,690,713]" captionStart="Figures 19" captionStartId="3.[192,278,1837,1861]" captionTargetBox="[223,1399,170,1804]" captionTargetId="figure-6@3.[192,1439,168,1819]" captionTargetPageId="3" captionText="Figures 19: Liocheles australasiae from Sri Lanka, pedipalp segments. Chela dorsal (1), external (2) and ventrointernal (3). Femur dorsal (4), external (5) and ventral (6). Patella dorsal (7) and ventral (8). Movable finger (9). Trichobothrial pattern is indicated." pageId="6" pageNumber="5">Fig. 2</figureCitation>
). Color uniformly reddish to yellowish brown, telson always yellow. Not troglobitic, median and three lateral pigmented eyes present. Chelicerae yellowish brown, reticulate. Carapace lacks carinae, punctate and bears straight median longitudinal groove. Pectinal teeth number 48. Metasomal segments sparsely setose and finely punctate, with smooth and rounded dorsal and lateral margins. Pedipalps covered by granules, dorsal surface of chela and femur of pedipalp granulated densely by bigger granules. Mainly ventral surface of pedipalp punctate (
<figureCitation id="8AF092C1FFC5CA74FDCFFC07055AFC79" box="[523,630,996,1019]" captionStart="Figures 19" captionStartId="3.[192,278,1837,1861]" captionTargetBox="[223,1399,170,1804]" captionTargetId="figure-6@3.[192,1439,168,1819]" captionTargetPageId="3" captionText="Figures 19: Liocheles australasiae from Sri Lanka, pedipalp segments. Chela dorsal (1), external (2) and ventrointernal (3). Femur dorsal (4), external (5) and ventral (6). Patella dorsal (7) and ventral (8). Movable finger (9). Trichobothrial pattern is indicated." pageId="6" pageNumber="5">Figs. 18</figureCitation>
). In contrast to female, in males fingers of chela con-spicuously flexed, but most of populations are parthenogenetic.
</paragraph>
</subSubSection>
</treatment>
</document>