245 lines
26 KiB
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245 lines
26 KiB
XML
<document ID-CLB-Dataset="51924" ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-GBIF-Dataset="c2e00414-9633-4890-ab72-cb2aa3e9d174" ID-PMC="PMC10208988" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-PubMed="37250203" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak & Sutcharit, Chirasak" docDate="2023" docId="984CE46B31C6567DAD46BBBB30C83240" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma gnomus Tongkerd & Panha 2023, sp. nov." docType="treatment" docUuid="FEE72641-8C04-4BF7-A1F7-D7204FD45C29" docUuidSource="ZooBank" docVersion="5" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1732652259105" updateUser="ExternalLinkService">
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<mods:mods id="637F0DA1F1A67CDEF154405AA5D1FC2F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1BCC77988FC4458D7FE7DC9E461B21FE">
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<mods:title id="A17AAF30FCE0B9566E6DBE430E53D956">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:name id="64E23BF4EE8AEF191BBD37470FB71F2F" type="personal">
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<mods:roleTerm id="1107BE1EA7815FF9C00C173D8DE634F8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="133C570CC3A7567F45F989A3D2FAD1EE">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="B30DEE3EFAD4876A69C6F3AD3A17B38F">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="4BB1E2F672F7F7E9C9A9364C74C5D243" type="personal">
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<mods:roleTerm id="5B1EE4CD1A4C696D605D159CBE0DC59D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="882136CB80B1C429E088F9A5D45CD7F4">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="F0D9B3B39782CFD7285BE73680C14C89">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="3B40CBC264081D14DC8FC16BDCCBC87E" type="personal">
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<mods:role id="1D88BBDEA27571F781C4BF138AAC1B01">
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<mods:roleTerm id="C84511729FED5A3B567C67670DB4E09F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BF81DED73CE76FD8C98185C4CF2C2D56">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="A21B833E24DAE54F49D6F4646357D8D3">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:name id="144595341F970123F6944E7BAD615281" type="personal">
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<mods:roleTerm id="DC123ADAD9894BA256A97BD557C82CDB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="323567AC7A9F4A7396C8E9A490DC21E8">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="BC7D2EC4968522E8E70BEC67A24F0A30">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="B2A2B6B1F5E216D8DB933B835FE92F01" type="personal">
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</mods:role>
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<mods:namePart id="89AD9BF1F9F5D0F27FD7C71898569AB1">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="FC1EC755B9C1E12294E596169602AFD6">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="6B3185BC0B1844AC21E48BFA0FF4F724" type="personal">
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<mods:roleTerm id="1DA4B702E40CEE636FAEE912F942CD13">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="44A7908EC5E1873D5A7E9D896F73F271">Panha, Somsak</mods:namePart>
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<mods:affiliation id="F69C1BC4BBF2C75F180CEAFAC267E5C4">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="6EEE06B683FE330D98D6B6BC6CE8BB6F" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:name id="057C3391DCE94C9339EDC0AF4EF429BC" type="personal">
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</mods:role>
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<mods:namePart id="C51E96ED6A3803F8F26BBA2A3268A4F3">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="7B94D201AE2D38D7DE2CD8AF05896042">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="7A5250076EB30A48783DCBF18AD7D3B4" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="1C8333735EEFCBABC6A042FDF042D495">text</mods:typeOfResource>
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<mods:titleInfo id="AE8933E94386133D54DD41C98A884855">
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<mods:title id="774283182BA77F78E52650266F38130F">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="CD3863AAF9D1B3522A56D214BBC1C89B">
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<mods:date id="EDD80BFC1BCAF675ECE9ECA5E448E709">2023</mods:date>
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<mods:number id="10518AC66DF683FFE8942063EDA584DB">2023-01-24</mods:number>
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</mods:detail>
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<mods:detail id="D1BDBB78C8D346C102FC5D911E2123CF" type="volume">
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<mods:number id="9782772C81109C8ACB14709A7D357F7E">1142</mods:number>
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</mods:detail>
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<mods:extent id="1795905031AF37E33631387402E18891" unit="page">
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<mods:start id="84819FC7B12AF915F1284C49A5651CDF">1</mods:start>
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<mods:end id="A1A2AC87DF6D967D5513B788C2146505">144</mods:end>
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<mods:url id="34EB7A1CA6F66ABF7559E97106CCBE77">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:url>
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</mods:location>
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<mods:classification id="5B4AE16329628DCE795659F8DDBAB581">journal article</mods:classification>
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<mods:identifier id="D081DD27B36C544F176CF3657528C31B" type="DOI">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:identifier>
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<mods:identifier id="11C7A17290BCA8D060BFB957205B12A9" type="Pensoft-Pub">1313-2970-1142-1</mods:identifier>
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<mods:identifier id="0E9015E2B554C979FC33DC87C3949227" type="ZooBank">A1129EE50F9941CFB73AE771B66E2486</mods:identifier>
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<mods:identifier id="64CD8A2B1F1C501B9143D3DA94486EE3" type="Pensoft-UUID">1D4BDF04F72B59B9984E00D0B4FB338D</mods:identifier>
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</mods:mods>
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<treatment id="984CE46B31C6567DAD46BBBB30C83240" ID-GBIF-Taxon="209102273" LSID="urn:lsid:zoobank.org:act:FEE72641-8C04-4BF7-A1F7-D7204FD45C29" httpUri="http://treatment.plazi.org/id/984CE46B31C6567DAD46BBBB30C83240" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Cyclophoridae" scope_subClass="Caenogastropoda" scope_subFamily="Cyclophorinae">
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<subSubSection id="640E76EE3B8BFB24E4013A0CB31A91A9" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="8836FD5A22B99744E6E4436DA2ABE993" pageId="0" pageNumber="1">
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26.
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<taxonomicName id="4BA6236428179B1566C797BD1B1159CC" ID-CoL="B3CKV" LSID="https://zoobank.org/FEE72641-8C04-4BF7-A1F7-D7204FD45C29" authority="Tongkerd & Panha" authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma gnomus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gnomus" status="sp. nov.">Rhiostoma gnomus Tongkerd & Panha</taxonomicName>
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<taxonomicNameLabel id="0DABA0893F743AF04E5CF9DBF9652244" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="1B60EFDD03541EF41ABF564A80CEFAC1" pageId="0" pageNumber="1" type="description">
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<paragraph id="8150B5C352CBCF0891609B2B3E8C78F9" pageId="0" pageNumber="1">
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<figureCitation id="F325D8A961A03E885B36F30F3ECA23B6" captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Distribution map of Rhiostoma morleti - species group." figureDoi="10.3897/zookeys.1142.90097.figure44" httpUri="https://binary.pensoft.net/fig/799902" pageId="0" pageNumber="1">Figs 44</figureCitation>
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<figureCitation id="27764D9E38CCD52EBAC1C4F162F6C662" captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Shell of Rhiostoma gnomus sp. nov. A holotype CUMZ 4716 from Pha Chu, Na Noi, Nan B-D paratypes CUMZ 4717 and E paratype CUMZ 4849 from type locality F paratype CUMZ 4718 from type locality shows albinistic shell form." figureDoi="10.3897/zookeys.1142.90097.figure51" httpUri="https://binary.pensoft.net/fig/799909" pageId="0" pageNumber="1">, 51</figureCitation>
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<figureCitation id="D10B0CC79BFB1C6FE7BB0626C2FDC120" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Radula morphology A Rhiostoma cambodjense, specimens CUMZ 4714 from Khao Chakan, Srakeo B Rhiostoma cheliopegma sp. nov., paratype CUMZ CUMZ 3985 / 2 Tham Takien, Khao Chamao, Rayong C Rhiostoma gnomus sp. nov. paratype CUMZ 4717 from Pha Chu, Na Noi, Nan D Rhiostoma laosense sp. nov. specimen CUMZ 10205 from Tad Muang, Khammouane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure53" httpUri="https://binary.pensoft.net/fig/799911" pageId="0" pageNumber="1">, 53C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E049D05CE1B6B6255B5AD970CC8F0E54" pageId="0" pageNumber="1" type="type material">
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<paragraph id="9286E0B788C6B782A777F1E66B1A0FA3" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="DE9FE1D6C79767B69440257568D1D4EF" pageId="0" pageNumber="1">
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<emphasis id="DA9EE7FC12A1785B29B8D53A8E941DF5" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 4716 (W 16.4 mm, H 10.7; Fig.
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<figureCitation id="06491B8A2C3B4B27ECEECA7AF3223487" captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Shell of Rhiostoma gnomus sp. nov. A holotype CUMZ 4716 from Pha Chu, Na Noi, Nan B-D paratypes CUMZ 4717 and E paratype CUMZ 4849 from type locality F paratype CUMZ 4718 from type locality shows albinistic shell form." figureDoi="10.3897/zookeys.1142.90097.figure51" httpUri="https://binary.pensoft.net/fig/799909" pageId="0" pageNumber="1">51A</figureCitation>
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).
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<emphasis id="CBFC121E58BE049117160E6F4A42B6D5" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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CUMZ 3922 (55 adults + 31 juveniles), CUMZ 4717 (3 shells; Figs
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<figureCitation id="861B728FC848449573D68FF7720BF39E" captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Shell of Rhiostoma gnomus sp. nov. A holotype CUMZ 4716 from Pha Chu, Na Noi, Nan B-D paratypes CUMZ 4717 and E paratype CUMZ 4849 from type locality F paratype CUMZ 4718 from type locality shows albinistic shell form." figureDoi="10.3897/zookeys.1142.90097.figure51" httpUri="https://binary.pensoft.net/fig/799909" pageId="0" pageNumber="1">51B-D</figureCitation>
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,
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<figureCitation id="76EAA1EF86945FF6686F1A3FED984FD8" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Radula morphology A Rhiostoma cambodjense, specimens CUMZ 4714 from Khao Chakan, Srakeo B Rhiostoma cheliopegma sp. nov., paratype CUMZ CUMZ 3985 / 2 Tham Takien, Khao Chamao, Rayong C Rhiostoma gnomus sp. nov. paratype CUMZ 4717 from Pha Chu, Na Noi, Nan D Rhiostoma laosense sp. nov. specimen CUMZ 10205 from Tad Muang, Khammouane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure53" httpUri="https://binary.pensoft.net/fig/799911" pageId="0" pageNumber="1">53C</figureCitation>
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), CUMZ 4718 (3 shells; Fig.
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<figureCitation id="84EB9D1CE5F8816B971F4E4047CE5A58" captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Shell of Rhiostoma gnomus sp. nov. A holotype CUMZ 4716 from Pha Chu, Na Noi, Nan B-D paratypes CUMZ 4717 and E paratype CUMZ 4849 from type locality F paratype CUMZ 4718 from type locality shows albinistic shell form." figureDoi="10.3897/zookeys.1142.90097.figure51" httpUri="https://binary.pensoft.net/fig/799909" pageId="0" pageNumber="1">51F</figureCitation>
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), CUMZ 4849 (9 shells; Fig.
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<figureCitation id="C744AB695BBED437EC746EEC86807FEF" captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Shell of Rhiostoma gnomus sp. nov. A holotype CUMZ 4716 from Pha Chu, Na Noi, Nan B-D paratypes CUMZ 4717 and E paratype CUMZ 4849 from type locality F paratype CUMZ 4718 from type locality shows albinistic shell form." figureDoi="10.3897/zookeys.1142.90097.figure51" httpUri="https://binary.pensoft.net/fig/799909" pageId="0" pageNumber="1">51E</figureCitation>
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), CUMZ 4872 (16 adults + 2 juveniles), NHMUK 20220447 (5 shells), and SMF 368682 (5 shells). All paratypes are from the type locality.
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</paragraph>
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<caption id="2A9F5AC697B32641CFF3E1E12EE089A0" doi="10.3897/zookeys.1142.90097.figure51" httpUri="https://binary.pensoft.net/fig/799909" pageId="0" pageNumber="1" start="Figure 51" startId="F51">
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<paragraph id="17B1CBF81889BC3817E72A68F73EAD0D" pageId="0" pageNumber="1">
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<emphasis id="90465EEBE084BC83592F6D47A92A70BB" bold="true" pageId="0" pageNumber="1">Figure 51.</emphasis>
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Shell of
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<taxonomicName id="4091FF88A8C5E52B2933C9FE8F99986D" authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma gnomus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gnomus">
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<emphasis id="3B5F8CB858CF5C41B75235200F36E9CD" italics="true" pageId="0" pageNumber="1">Rhiostoma gnomus</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="6CBF99E2CBD59AC1E04FF73F9F8D702A" bold="true" pageId="0" pageNumber="1">A</emphasis>
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holotype CUMZ 4716 from Pha Chu, Na Noi, Nan
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<emphasis id="B41277731D8E3DC5683AE65633F41BEF" bold="true" pageId="0" pageNumber="1">B-D</emphasis>
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paratypes CUMZ 4717 and
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<emphasis id="10A53ACF7D3E5A139584B6663366F9F1" bold="true" pageId="0" pageNumber="1">E</emphasis>
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paratype CUMZ 4849 from type locality
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<emphasis id="EEAFC7372EB8053EB5DBF6CE4B3541F9" bold="true" pageId="0" pageNumber="1">F</emphasis>
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paratype CUMZ 4718 from type locality shows albinistic shell form.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="B888FD532AAB9EE002BD42394B742916" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="45C8A34273CD323BBC8995FAE4F4B19F" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="5F797F4C0ED7BF82A000268642BD255E" pageId="0" pageNumber="1">
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Limestone hill with dry deciduous forest at Pha Chu (Srinan), Na Noi District, Nan Province, Thailand (
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<geoCoordinate id="B96053CD804C5892CDC48E09B3ABF6F6" degrees="18" direction="north" minutes="22" orientation="latitude" precision="1" seconds="4.6" value="18.367945">18°22'4.6"N</geoCoordinate>
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,
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<geoCoordinate id="7DB8FABB575B277A257940B1D2A0B933" degrees="100" direction="east" minutes="50" orientation="longitude" precision="1" seconds="23.4" value="100.839836">100°50'23.4"E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="F9B6AA25A67D19250EAF2F82DB7172FC" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="58D0A2EC36D64E45161D9120F03849AE" pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph id="6C6CE48C0D348312CDA08FF602FA8F1D" pageId="0" pageNumber="1">
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<materialsCitation id="CCB66CD1F81BA553EA09DAFA8C88C29F" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Nan" specimenCount="1" stateProvince="Nan">
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<emphasis id="69B3809524DD4D5E25A99F8EDD1519CD" bold="true" pageId="0" pageNumber="1">
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<collectingCountry id="CD52087C755057DE46693C7F74ACD5DF" name="Thailand">Thailand</collectingCountry>
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</emphasis>
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: Sri
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<collectingRegion id="A3F43273AAA05C5F8D598B8CA3D5A479" country="Thailand" name="Nan">Nan</collectingRegion>
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, Na Noi,
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<collectingRegion id="2883939232B3FA32BBF95B38F683D95E" country="Thailand" name="Nan">Nan</collectingRegion>
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: CUMZ 10035
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="43275AFB8A9166E96116378B06980CAA" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="180C86EC27778BCCA7C31964B69399BA" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="D3DABC15E88AA2D7AFA38621617FA641" pageId="0" pageNumber="1">Shell small and heliciform; detached whorl shorter than apertural width. Shell colour usually with brownish zigzag patterns. Breathing device notch-shaped. Operculum calcareous, low cup-shaped.</paragraph>
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</subSubSection>
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<subSubSection id="194466AC9EAD2ACE8E047201751AD006" pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph id="C29705D96C209AB794E7C93E8D78FA5D" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph id="F767A561D40D3E91FA7C467C7338D91A" pageId="0" pageNumber="1">
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This new species differs from all other members in this species group by having a small and heliciform shell, sometimes with a very short detached whorl and notch-shaped breathing device. Except for
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<taxonomicName id="7428C639E9A09F23912B928992EBD20B" lsidName="R. cambodjense" pageId="0" pageNumber="1" rank="species" species="cambodjense">
|
|
<emphasis id="15E040A1894A1D31EB3C308D7F06E6B3" italics="true" pageId="0" pageNumber="1">R. cambodjense</emphasis>
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</taxonomicName>
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|
, all other species have a flattened or sub-discoidal shell and a canal-shaped breathing device.
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<taxonomicName id="08E31A4CB7B8DD301372A5D973FD3041" authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma gnomus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gnomus">
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|
<emphasis id="26DFE970A971A4B6103339CA0C10A1B5" italics="true" pageId="0" pageNumber="1">Rhiostoma gnomus</emphasis>
|
|
</taxonomicName>
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|
sp. nov. differs from
|
|
<taxonomicName id="99877C37EB1315B03D37C8720E82B860" lsidName="R. cambodjense" pageId="0" pageNumber="1" rank="species" species="cambodjense">
|
|
<emphasis id="68FD72A93EDA911D6889C375C659D373" italics="true" pageId="0" pageNumber="1">R. cambodjense</emphasis>
|
|
</taxonomicName>
|
|
by having a relatively smaller shell size, lighter shell colour, and apex without dark colour. In contrast,
|
|
<taxonomicName id="0C75722AE979386575C16CE1CC1C5F3C" lsidName="R. cambodjense" pageId="0" pageNumber="1" rank="species" species="cambodjense">
|
|
<emphasis id="8EAA1AAE768D8D244118DF2997488588" italics="true" pageId="0" pageNumber="1">R. cambodjense</emphasis>
|
|
</taxonomicName>
|
|
tends to have a larger shell and darker shell colour.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E61B947AC119817E2B330C83D3653E8B" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="E62312C820F3924C7723DEF58591578F" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="60DDE1A3483FDCD6A21D697250B70054" pageId="0" pageNumber="1">
|
|
<emphasis id="880D976B6C796793C18838893C093BBD" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="8DD52EF86FDD3CBDB7503C131AC5FE99" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shell small, width 14.6-18.2 mm, height 8.9-11.1 mm, thin, and heliciform. Apex acute; spire elevated. Whorls 4 to 5, convex, increasing regularly; suture wide and deep; last whorl rounded. Shell surface smooth or with fine growth lines. Shell colour usually with brownish zigzag patterns or uniformly white to brown; narrow and brownish spiral band usually present (or absent in albinistic forms). Periostracum thin, corneous, and brownish colour. Detached whorl very short and slightly descending. Peristome circular and double; lip slightly thickened and rarely expanded. Aperture opened sub-laterally. Breathing device notch-shaped; outer lip protruding, with shallow groove; inner lip with shallow incision. Umbilicus widely opened and deep. Operculum calcareous, low cup-shaped, and multispiral (Fig.
|
|
<figureCitation id="CBD7FDC73D59C8BDFB20EBF93F7B55C0" captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Shell of Rhiostoma gnomus sp. nov. A holotype CUMZ 4716 from Pha Chu, Na Noi, Nan B-D paratypes CUMZ 4717 and E paratype CUMZ 4849 from type locality F paratype CUMZ 4718 from type locality shows albinistic shell form." figureDoi="10.3897/zookeys.1142.90097.figure51" httpUri="https://binary.pensoft.net/fig/799909" pageId="0" pageNumber="1">51</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="DCD5C5FCCA79CE9F6FCEB28C9C3CF60B" pageId="0" pageNumber="1">
|
|
<emphasis id="1EDAEF173A2BE9DBD92D3AF46592FA08" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="1226B16163605839289FD75B5A8148DA" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
|
.
|
|
</emphasis>
|
|
Teeth arrangement and shape are similar to those of
|
|
<taxonomicName id="DE29DAD398044714850BD0D7FA8546BA" lsidName="R. asiphon" pageId="0" pageNumber="1" rank="species" species="asiphon">
|
|
<emphasis id="8F70A9838EE5B1A081082FF027FC80B2" italics="true" pageId="0" pageNumber="1">R. asiphon</emphasis>
|
|
</taxonomicName>
|
|
. Central tooth with large central cusp and two lateral cusps on each side. Lateral teeth composed of four cusps; central cusp triangular; two small inner cusps with pointed tips and tapering in size, and one small outer cusp. Marginal teeth each composed of three cusps (Fig.
|
|
<figureCitation id="442385A121C7888CACA187D0556D8F8A" captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Radula morphology A Rhiostoma cambodjense, specimens CUMZ 4714 from Khao Chakan, Srakeo B Rhiostoma cheliopegma sp. nov., paratype CUMZ CUMZ 3985 / 2 Tham Takien, Khao Chamao, Rayong C Rhiostoma gnomus sp. nov. paratype CUMZ 4717 from Pha Chu, Na Noi, Nan D Rhiostoma laosense sp. nov. specimen CUMZ 10205 from Tad Muang, Khammouane, Laos." figureDoi="10.3897/zookeys.1142.90097.figure53" httpUri="https://binary.pensoft.net/fig/799911" pageId="0" pageNumber="1">53C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="8DE013E74728C95F8DA78C21BA126F77" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="2A78FD814677E0159862624B3343CDA6" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="96DE0167F36813AB4A2DA6AFE829AC2E" pageId="0" pageNumber="1">
|
|
The species name is derived from the Latin word
|
|
<emphasis id="DC3F72A206B500F9BE0328B9C614947B" italics="true" pageId="0" pageNumber="1">gnomus</emphasis>
|
|
, meaning dwarf. It refers to the small shell size of the new species.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="DFF5C409A381D29BD2A916FD3266C91B" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="0B99CE6732DFCCBF4259268772215B51" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="1C7B5C172B993E74ED40983C6C5366B9" pageId="0" pageNumber="1">
|
|
This new species is known only from the type locality at the base of a limestone cliff in Nan Province, Thailand, near the Thai-Laos border (Fig.
|
|
<figureCitation id="CFF00E7C4C7DB567A27ADC3120D1F48F" captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Distribution map of Rhiostoma morleti - species group." figureDoi="10.3897/zookeys.1142.90097.figure44" httpUri="https://binary.pensoft.net/fig/799902" pageId="0" pageNumber="1">44</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="905522EC87D2443FC56DD83A28F3EF56" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="3CAAF5DF5EB6C6F8479F842ADC3F1EF3" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="A31D648CB38753F09176A37B3F3B5C82" pageId="0" pageNumber="1">
|
|
Although this species is superficially similar to the genus
|
|
<taxonomicName id="C8E8C80708D14B85C3C9795A970E74B1" authorityName="Swainson" authorityYear="1840" class="Gastropoda" family="Cyclophoridae" genus="Cyclotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclotus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis id="4D00563D2A9873871E6FE15BB6D50C73" italics="true" pageId="0" pageNumber="1">Cyclotus</emphasis>
|
|
</taxonomicName>
|
|
, its calcareous, cup-shaped and multispiral operculum with elevated lamellae and its incomplete tube as breathing device on the apertural lip suggests it is a member of genus
|
|
<taxonomicName id="6F971C9BDCBBE5689041F3364651023D" authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis id="71B7E3BDC146A416893CB54767B4F613" italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
|
|
</taxonomicName>
|
|
. Shell colour variation is observed in the type population varied from a brownish zigzag pattern, pale brown blotches to monochrome whitish, or the albinistic shell form found sympatrically within the type population and here recognised as colour variation.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |