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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2011" docId="7D2EE251673097B211B91B09C7E1DEA8" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha fluminoris Hoffman 1977" docType="treatment" docVersion="6" lastPageNumber="68" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="65" updateTime="1668152297771" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>131</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>161</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152031129" LSID="urn:lsid:plazi:treatment:7D2EE251673097B211B91B09C7E1DEA8" httpUri="http://treatment.plazi.org/id/7D2EE251673097B211B91B09C7E1DEA8" lastPageId="67" lastPageNumber="68" pageId="64" pageNumber="65">
<subSubSection pageId="64" pageNumber="65" type="nomenclature">
<paragraph pageId="64" pageNumber="65">
<taxonomicName LSID="http://species-id.net/wiki/Orthomorpha_fluminoris" authority="Hoffman, 1977" authorityName="Hoffman" authorityYear="1977" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris Hoffman, 1977</taxonomicName>
Figs 5556
</paragraph>
</subSubSection>
<subSubSection pageId="64" pageNumber="65" type="reference_group">
<paragraph pageId="64" pageNumber="65">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris</taxonomicName>
<bibRefCitation author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">Hoffman 1977</bibRefCitation>
: 701 (D).
</paragraph>
<paragraph pageId="64" pageNumber="65">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris</taxonomicName>
-
<bibRefCitation pageId="64" pageNumber="65">Golovatch 1998</bibRefCitation>
: 42 (D, M).
</paragraph>
</subSubSection>
<subSubSection pageId="64" pageNumber="65" type="holotype">
<paragraph pageId="64" pageNumber="65">Holotype.</paragraph>
<paragraph pageId="64" pageNumber="65">♂ (MHNG), Malaysia, environs of Kuala Lumpur, Batu Caves, 24.02.1975, leg. P. Strinati.</paragraph>
</subSubSection>
<subSubSection lastPageId="67" lastPageNumber="68" pageId="64" pageNumber="65" type="redescription">
<paragraph pageId="64" pageNumber="65">Redescription.</paragraph>
<paragraph pageId="64" pageNumber="65">Length 39 mm, width of midbody pro- and metazona 2.9 and 4.4 mm, respectively. Coloration of alcohol material upon long-term preservation dark castaneous brown with contrasting yellowish paraterga; venter, legs yellownish and tip of epiproct light red-brown, antennae also light red-brown, but antenomere 6 slightly infuscate, brownish, antenomere 7 dark brown, and tip of antennae pallid (Fig. 55).</paragraph>
<paragraph lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65">
Clypeolabral region densely setose, vertex sparsely so, epicranial suture distinct. Antennae rather short, clavate (antennomere 6 broadest), extending behind body segment 2 dorsally. Head in width &lt;collum &lt;segment 2 &lt;3 &lt;4 &lt;5-16; thereafter body gently and gradually tapering. Collum with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3 posterior; caudal corner of paraterga nearly pointed, slightly declined ventrally and continuing
<normalizedToken originalValue="collums">collum's</normalizedToken>
convexity (Fig. 55A &amp; B); paraterga acutangular (ca 50°), slightly extending behind tergal margin, posterior
<pageBreakToken pageId="65" pageNumber="66" start="start">edge</pageBreakToken>
slightly concave (Fig. 55A &amp; B). Tegument smooth and poorly shining, prozona very finely shagreened, metazona leathery, rugulose, below paraterga microgranular. Metaterga 2-19 with two transverse rows of setae: 2+2 in anterior (pre-sulcus) row and 3+3 in posterior (postsulcus) one, all mostly abraded, but still traceable as insertion points. Tergal setae simple, moderately long, about 1/4 metatergal length. Axial line traceable. Paraterga very well developed (Fig. 55A-G), all lying below dorsum (at about 1/4 body height), slightly upturned to subhorizontal, in lateral view moderately strongly enlarged on pore-bearing segments, thinner on poreless ones; shoulders always present, broadly rounded and narrowly bordered, fused to callus; anterior edge of paraterga 2 straight, following paraterga rounded, caudal corner of all paraterga rounded, extending invariably and increasingly beyond rear tergal margin. Calluses delimited by a sulcus both dorsally and ventrally, rather broad. Posterior edge of paraterga evidently concave, especially strongly so on segments 16-19. Ozopores evident, lateral, lying in an ovoid groove at about 1/4 in front of caudal corner. Transverse sulcus complete on metaterga 5-18, incomplete on segments 4 and 19, very narrow, shallow, not reaching bases of paraterga, very finely ribbed at bottom (Fig. 55A, C
<pageBreakToken pageId="66" pageNumber="67" start="start">&amp;</pageBreakToken>
amp; F). Stricture between pro- and metazona narrow and shallow, faintly beaded at bottom down to base of paraterga (Fig. 55A-F). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2-4, a low and interrupted swelling supplied with a small tooth caudally on segments 5-7, retained only as small denticles on segments 8-14, thereafter missing (Fig. 55B, D &amp; E). Epiproct (Fig. 55E-G) conical, flattened dorsoventrally, with two small apical papillae directed ventrocaudally, subtruncate at tip; pre-apical papillae small, lying close to tip. Hypoproct (Fig. 55G) nearly semi-circular, caudal tip very broadly rounded, setiferous knobs at caudal edge very small and well-separated.
</paragraph>
<paragraph pageId="66" pageNumber="67">Sterna sparsely setose, without modifications; cross-impressions shallow; a pair of strongly separated, anteroventrally directed, narrowly rounded, setose cones between ♂ coxae 4 (Fig. 55H &amp; I). A paramedian pair of small tubercles in front of gonopod aperture. Legs long and slender, midbody ones ca 1.3-1.5 times as long as body height, prefemora without modifications, ♂ tarsal brushes present until on legs of segment 17.</paragraph>
<paragraph lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
Gonopods (Fig. 56) a little more complex. Coxa long and slender, with several strong setae distodorsally. Prefemur densely setose, about half the length of femorite +
<normalizedToken originalValue="“postfemoral”">&quot;postfemoral&quot;</normalizedToken>
part. Femorite slender, slightly curved and nearly not enlarged distad, with a small, but conspicuous ventral knob near base of lamina lateralis, with a
<pageBreakToken pageId="67" pageNumber="68" start="start">&quot;</pageBreakToken>
postfemoral&quot; part demarcated by a distinct, oblique, lateral sulcus; solenophore with a tridentate tip, middle denticle shorter than terminal tooth, but longer than a small subterminal lobule; solenomere long, flagelliform, a short tip exposed.
</paragraph>
<caption pageId="67" pageNumber="68">
<paragraph pageId="67" pageNumber="68">
Figure 55.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris</taxonomicName>
Hoffman, 1977, ♂ holotype. A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively
<normalizedToken originalValue="EG">E-G</normalizedToken>
posterior part of body, lateral, dorsal and ventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<caption pageId="67" pageNumber="68">
<paragraph pageId="67" pageNumber="68">
Figure 56.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha fluminoris" order="Polydesmida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="fluminoris">Orthomorpha fluminoris</taxonomicName>
Hoffman, 1977, ♂ holotype.
<normalizedToken originalValue="AC">A-C</normalizedToken>
right gonopod, lateral, mesal and subcaudal views, respectively. Scale bar: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="67" pageNumber="68" type="remarks">
<paragraph pageId="67" pageNumber="68">Remarks.</paragraph>
<paragraph pageId="67" pageNumber="68">
The original description is fairly complete, also illustrated in due detail (
<bibRefCitation author="Hoffman, RL" journalOrPublisher="Journal of Natural History" pageId="114" pageNumber="115" pagination="361 - 389" title="Descriptions and allocation of new or poorly known genera and species of Paradoxosomatidae from south-eastern Asia (Diplopoda: Polydesmida)." url="doi: 10.1080/00222937300770281" volume="7" year="1973">Hoffman 1973</bibRefCitation>
). The above redescription, also based on the holotype, is only meant to bring it to our standard and to present additional illustrations.
</paragraph>
</subSubSection>
</treatment>
</document>