345 lines
34 KiB
XML
345 lines
34 KiB
XML
<document id="8BAB710EADC4C95DE4F5C59161A1A490" ID-DOI="10.5281/zenodo.197793" ID-GBIF-Dataset="8f84f706-94cc-43a5-9027-efd2138aa784" ID-ISSN="1175-5326" ID-Zenodo-Dep="197793" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460278674581" checkinUser="plazi" docAuthor="Braasch, Dietrich & Boonsoong, Boonsatien" docDate="2010" docId="716BF555FF87FFDA8FF0F9907FEF642D" docLanguage="en" docName="zt02610p026.pdf" docOrigin="Zootaxa 2610" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Epeorus (Belovius) khayengensis Boonsoong and Braasch, sp. nov." docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="8D528D2DFF8BFFCA8F67FF8F7D58656A" masterDocTitle="A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia" masterLastPageNumber="26" masterPageNumber="1" pageNumber="13" updateTime="1698249041875" updateUser="plazi">
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<mods:mods id="E6811BB2056C09F25CEBB61DB6D0910F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D7AF13C78983B0E16D46A072FB23AF56">
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<mods:title id="9B1A377EAACC70CF2CBA12F9609B3741">A contribution to the Heptageniidae (Insecta, Ephemeroptera) of Thailand and Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name id="3536B162108167542520F21A38B30175" type="personal">
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<mods:role id="8FE1B398B2584F794225F36D21B463B4">
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<mods:roleTerm id="71522700BD7B1DFA6F45D9162A4E14C9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E6E4ABF5E2E4B69402929654C679575B">Braasch, Dietrich</mods:namePart>
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</mods:name>
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<mods:name id="5EE872D450B16B9786FEF26B07E29119" type="personal">
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<mods:role id="35BF628163750384561BF358CB80E215">
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<mods:roleTerm id="EBF7BB9C0F9B1281B69B560A75054641">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9426E7191E4D8FCEE1F79E0860494953">Boonsoong, Boonsatien</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="48AC67619C91B19B6906D2C267E65B93">text</mods:typeOfResource>
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<mods:relatedItem id="A2C1F349467043E3EC78D4EFBFA5645F" type="host">
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<mods:titleInfo id="3D489A704B6CE12C036952BCE86E454D">
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<mods:title id="476672D1849AAE4E7798C5AFB2D89F4F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0BDEB7A718157CA2B3E591781C98513B">
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<mods:date id="1B9971E990AE80415F204F215460D593">2010</mods:date>
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<mods:detail id="7AE7BAF9A40A1A2FA2628C4E56F6DB7C" type="volume">
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<mods:number id="CA686A4D3C909090875DCA7B6784352F">2610</mods:number>
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</mods:detail>
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<mods:extent id="E5A08738DFD8F158307156751C797C69" unit="page">
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<mods:start id="DA9D280A357DC605696DA473D9829B90">1</mods:start>
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<mods:end id="6F4A802E4588B1583541398B5659C0A0">26</mods:end>
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</mods:extent>
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<mods:classification id="A23FBF1200CF8A1E8A1DCF6E41841D0F">journal article</mods:classification>
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<mods:identifier id="B63409E8C900087662AA4BA325433C39" type="DOI">10.5281/zenodo.197793</mods:identifier>
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<mods:identifier id="60674A92F93AA532A86357AD743CA810" type="GBIF-Dataset">8f84f706-94cc-43a5-9027-efd2138aa784</mods:identifier>
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<mods:identifier id="A399C655E35BDD344047E3D60AAE0A12" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BA6E6F0F517BA4F0B05BA2848036F4C8" type="Zenodo-Dep">197793</mods:identifier>
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</mods:mods>
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<treatment id="716BF555FF87FFDA8FF0F9907FEF642D" ID-DOI="http://doi.org/10.5281/zenodo.6211085" ID-GBIF-Taxon="127804275" ID-Zenodo-Dep="6211085" LSID="urn:lsid:plazi:treatment:716BF555FF87FFDA8FF0F9907FEF642D" httpUri="http://treatment.plazi.org/id/716BF555FF87FFDA8FF0F9907FEF642D" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
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<subSubSection id="B1D817C8FF87FFC68FF0F9907E9A6353" box="[151,962,1567,1593]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="F97D4443FF87FFC68FF0F9907E9A6353" blockId="12.[151,962,1567,1593]" box="[151,962,1567,1593]" pageId="12" pageNumber="13">
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<heading id="A235F32FFF87FFC68FF0F9907E9A6353" bold="true" box="[151,962,1567,1593]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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<emphasis id="CBB69851FF87FFC68FF0F9907E9A6353" bold="true" box="[151,962,1567,1593]" pageId="12" pageNumber="13">
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<taxonomicName id="3EC23FC0FF87FFC68FF0F9907E056353" ID-CoL="6FNWJ" authority="Boonsoong and Braasch" authorityName="Boonsoong and Braasch" box="[151,861,1567,1593]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="khayengensis" status="sp. nov." subGenus="Belovius">
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<emphasis id="CBB69851FF87FFC68FF0F9907F716353" bold="true" box="[151,553,1567,1593]" italics="true" pageId="12" pageNumber="13">Epeorus (Belovius) khayengensis</emphasis>
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Boonsoong and Braasch
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</taxonomicName>
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<taxonomicNameLabel id="D085252AFF87FFC68C03F9907E9A6353" box="[868,962,1567,1593]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B1D817C8FF87FFC48FF0F9E578116322" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
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<paragraph id="F97D4443FF87FFC68FF0F9E57FD463C0" blockId="12.[151,1437,1642,2004]" pageId="12" pageNumber="13">
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<emphasis id="CBB69851FF87FFC68FF0F9E57C9063EE" bold="true" box="[151,456,1642,1668]" pageId="12" pageNumber="13">Description. Mature La:</emphasis>
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Body length
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<quantity id="3E3AE9A6FF87FFC68D00F9E57F8A63EE" box="[615,722,1642,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="12" pageNumber="13" unit="mm" value="10.5">10.5 mm</quantity>
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; length of caudal filaments
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<quantity id="3E3AE9A6FF87FFC68B70F9E579DB63EE" box="[1047,1155,1642,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="13" unit="mm" value="11.5">11.5 mm</quantity>
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. Body generally brown, with lighter and darker markings (Fig. 53).
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</paragraph>
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<paragraph id="F97D4443FF87FFC68FA2F93A78C462BE" blockId="12.[151,1437,1642,2004]" pageId="12" pageNumber="13">
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<emphasis id="CBB69851FF87FFC68FA2F93A7C4E63A5" bold="true" box="[197,278,1717,1743]" pageId="12" pageNumber="13">Head:</emphasis>
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Length
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<quantity id="3E3AE9A6FF87FFC68E19F93A7CB963A5" box="[382,481,1717,1743]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="mm" value="1.3">1.3 mm</quantity>
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, width
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<quantity id="3E3AE9A6FF87FFC68D59F93A7FF963A5" box="[574,673,1717,1743]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="13" unit="mm" value="2.1">2.1 mm</quantity>
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; brown, with conspicuous marks (Fig. 54); subquadrangular; anterior and lateral margins convex, with rows of dense hairlike setae; posterior margin slightly concave. Compound eyes black. Antennal scapes and pedicels brown, flagellae pale. Labrum (Fig. 55) with 3 large ridges on apical margin, with long hairlike setae on lateral margin, and with notch on anterior margin. Mandible (Figs. 56–57) with scattered setae along the molar area; incisor slightly developed. Maxillae (Fig. 58) each without armature on crown of galea-lacinia, apically with 3 acute teeth; medially with fine setae; terminal segment of maxillary palp with dense rows of hairlike setae. Hypopharynx (Fig. 59): Lingua with anterolateral lobes; superlinguae slightly expanded, laterally with rows of dense hairlike setae. Labium (Fig.
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</paragraph>
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<paragraph id="F97D4443FF86FFC78FF0FF1779ED65BD" blockId="13.[151,1437,152,626]" pageId="13" pageNumber="14">60) with broad V-shaped separation of glossae; glossae subtriangular; paraglossae slightly expanded laterally; apical segment of each palp acutely pointed, dorsal surface with dense row of comb setae.</paragraph>
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<paragraph id="F97D4443FF86FFC78FA2FF6D7EC464F8" blockId="13.[151,1437,152,626]" pageId="13" pageNumber="14">
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<emphasis id="CBB69851FF86FFC78FA2FF6D7C766596" bold="true" box="[197,302,226,252]" pageId="13" pageNumber="14">Thorax:</emphasis>
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Pronotum with some dark marks (Fig. 53). Legs (Fig. 61) with 2 dark brown, median spots, lateral spots greyish, scattered short, ovate bristles on dorsal surfaces (Fig. 62), distally dark brown; tibiae basally brown, having long brown band in the middle, tarsi darkened, fringe of stout long hairs at posterior margins of femora, tibiae and tarsi. Length of femur 2.57, width
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<quantity id="3E3AE9A6FF86FFC78CCFFEDD794F6406" box="[936,1047,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="13" pageNumber="14" unit="mm" value="1.17">1.17 mm</quantity>
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, length of tibia 2.34, width
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<quantity id="3E3AE9A6FF86FFC78A0FFEDD7D9064F8" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="13" pageNumber="14" unit="mm" value="0.18">0.18 mm</quantity>
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, length of tarsus
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<quantity id="3E3AE9A6FF86FFC78EE8FEF77CA264F8" box="[399,506,376,402]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="13" pageNumber="14" unit="mm" value="0.59">0.59 mm</quantity>
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; claws with 4–5 denticles (Fig. 63).
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</paragraph>
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<paragraph id="F97D4443FF86FFC78FABFE1278286718" blockId="13.[151,1437,152,626]" pageId="13" pageNumber="14">
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<emphasis id="CBB69851FF86FFC78FABFE127C0864DD" bold="true" box="[204,336,413,439]" pageId="13" pageNumber="14">Abdomen:</emphasis>
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Abdominal terga brown, terga without spines, but with striking pattern (Fig. 65); hind margin of tergum provided with tooth-like bristles (Fig. 64); terga II–VII with long postero-lateral extensions (Fig. 65); sterna light yellowish without markings but partially visible muscle impressions. Lamellae of gill I somewhat extended beneath abdomen (Fig. 66). Gills II–VII with anal ribs arched and last rib straight (Figs. 67–68, see arrow); middle pairs large with wide rounded anal-proximal projections; gill faces with exception of marginal parts reddish tinged, scattered with very fine dotting; cerci with row of hairlike setae dorsally.
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</paragraph>
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<paragraph id="F97D4443FF86FFC78FF0F85979976287" blockId="13.[151,1231,2006,2030]" box="[151,1231,2006,2030]" pageId="13" pageNumber="14">
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<emphasis id="CBB69851FF86FFC78FF0F8597CE46284" bold="true" box="[151,444,2006,2030]" pageId="13" pageNumber="14">PLATE VIII. FIGURE 53</emphasis>
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. Habitus of
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<taxonomicName id="3EC23FC0FF86FFC78D27F8587EFA6284" box="[576,930,2006,2030]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="khayengensis" status="sp. nov." subGenus="Belovius">
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<emphasis id="CBB69851FF86FFC78D27F8587FC16284" box="[576,665,2007,2030]" italics="true" pageId="13" pageNumber="14">Epeorus</emphasis>
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||
(
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<emphasis id="CBB69851FF86FFC78DCEF8587E5D6284" box="[681,773,2007,2030]" italics="true" pageId="13" pageNumber="14">Belovius</emphasis>
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||
)
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||
<emphasis id="CBB69851FF86FFC78C73F8587EFA6284" box="[788,930,2007,2030]" italics="true" pageId="13" pageNumber="14">khayengensis</emphasis>
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</taxonomicName>
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<emphasis id="CBB69851FF86FFC78CCEF85979476284" bold="true" box="[937,1055,2006,2030]" pageId="13" pageNumber="14">
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||
<taxonomicNameLabel id="D085252AFF86FFC78CCEF8597EA66284" box="[937,1022,2006,2030]" pageId="13" pageNumber="14" rank="species">
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sp.
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<date id="8D7C6283FF86FFC78CA8F8597EA66284" box="[975,1022,2006,2030]" pageId="13" pageNumber="14" value="1953-11">nov.</date>
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</taxonomicNameLabel>
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53
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</emphasis>
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, Dorsal habitus.
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</paragraph>
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<paragraph id="F97D4443FF85FFC48FF0FA7E78116322" blockId="14.[151,1437,1521,1609]" pageId="14" pageNumber="15">
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<emphasis id="CBB69851FF85FFC48FF0FA7E7C866363" bold="true" box="[151,478,1521,1545]" pageId="14" pageNumber="15">PLATE IX. FIGURES 54–60</emphasis>
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. Larva of
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<taxonomicName id="3EC23FC0FF85FFC48D2BFA7D7EF56363" box="[588,941,1521,1545]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="khayengensis" status="sp. nov." subGenus="Belovius">
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<emphasis id="CBB69851FF85FFC48D2BFA7D7FFD6363" box="[588,677,1522,1545]" italics="true" pageId="14" pageNumber="15">Epeorus</emphasis>
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(
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<emphasis id="CBB69851FF85FFC48DD3FA7D7E486363" box="[692,784,1522,1545]" italics="true" pageId="14" pageNumber="15">Belovius</emphasis>
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)
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<emphasis id="CBB69851FF85FFC48C78FA7D7EF56363" box="[799,941,1522,1545]" italics="true" pageId="14" pageNumber="15">khayengensis</emphasis>
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</taxonomicName>
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<emphasis id="CBB69851FF85FFC48CD4FA7E79716363" bold="true" box="[947,1065,1521,1545]" pageId="14" pageNumber="15">
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<taxonomicNameLabel id="D085252AFF85FFC48CD4FA7E79516363" box="[947,1033,1521,1545]" pageId="14" pageNumber="15" rank="species">
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sp.
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<date id="8D7C6283FF85FFC48CBDFA7E79516363" box="[986,1033,1521,1545]" pageId="14" pageNumber="15" value="1954-11">nov.</date>
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</taxonomicNameLabel>
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54
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</emphasis>
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, Head capsule.
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<emphasis id="CBB69851FF85FFC48BB5FA7E79B56363" bold="true" box="[1234,1261,1521,1545]" pageId="14" pageNumber="15">55</emphasis>
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, Labrum, dorsal view (left) and ventral view (right).
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<emphasis id="CBB69851FF85FFC48D58F99E7F046343" bold="true" box="[575,604,1553,1577]" pageId="14" pageNumber="15">56</emphasis>
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, Left mandible.
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<emphasis id="CBB69851FF85FFC48C78F99E7E636343" bold="true" box="[799,827,1553,1577]" pageId="14" pageNumber="15">57</emphasis>
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, Right mandible.
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<emphasis id="CBB69851FF85FFC48B69F99E79726343" bold="true" box="[1038,1066,1553,1577]" pageId="14" pageNumber="15">58</emphasis>
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, Left maxilla, ventral view.
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<emphasis id="CBB69851FF85FFC48A1EF99E78CD6343" bold="true" box="[1401,1429,1553,1577]" pageId="14" pageNumber="15">59</emphasis>
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, Hypopharynx, dorsal view (left) and ventral view (right).
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<emphasis id="CBB69851FF85FFC48D98F9BE7E416323" bold="true" box="[767,793,1585,1609]" pageId="14" pageNumber="15">60</emphasis>
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, Labium, dorsal view (left) and ventral view (right).
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</paragraph>
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</subSubSection>
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<subSubSection id="B1D817C8FF85FFC48FA2F9F97C5F63DC" pageId="14" pageNumber="15" type="etymology">
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<paragraph id="F97D4443FF85FFC48FA2F9F97C5F63DC" blockId="14.[151,1437,1654,2017]" pageId="14" pageNumber="15">
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<emphasis id="CBB69851FF85FFC48FA2F9F97C0E63FA" bold="true" box="[197,342,1654,1680]" pageId="14" pageNumber="15">Etymology:</emphasis>
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This species is named for the stream, where the
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<typeStatus id="2679FAE1FF85FFC48CEEF9F87EE463FB" box="[905,956,1655,1681]" pageId="14" pageNumber="15">type</typeStatus>
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material was collected: Khayeng stream,
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<collectingCountry id="81D504D3FF85FFC48FF0F9137C5B63DC" box="[151,259,1692,1718]" name="Thailand" pageId="14" pageNumber="15">Thailand</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B1D817C8FF85FFC48FA2F94E7FEA6221" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="F97D4443FF85FFC48FA2F94E7FEA6221" blockId="14.[151,1437,1654,2017]" pageId="14" pageNumber="15">
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<emphasis id="CBB69851FF85FFC48FA2F94E7C1F63B1" bold="true" box="[197,327,1729,1755]" pageId="14" pageNumber="15">Diagnosis:</emphasis>
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<taxonomicName id="3EC23FC0FF85FFC48E29F94E7F8963B0" box="[334,721,1729,1755]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="khayengensis" status="sp. nov." subGenus="Belovius">
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<emphasis id="CBB69851FF85FFC48E29F94E7CE863B0" box="[334,432,1729,1754]" italics="true" pageId="14" pageNumber="15">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="CBB69851FF85FFC48EA7F94E7F7D63B0" box="[448,549,1729,1754]" italics="true" pageId="14" pageNumber="15">Belovius</emphasis>
|
||
)
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<emphasis id="CBB69851FF85FFC48D52F94E7F8963B0" box="[565,721,1729,1754]" italics="true" pageId="14" pageNumber="15">khayengensis</emphasis>
|
||
</taxonomicName>
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||
<emphasis id="CBB69851FF85FFC48DBFF94E7E6D63B1" bold="true" box="[728,821,1729,1755]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="D085252AFF85FFC48DBFF94E7E6D63B1" box="[728,821,1729,1755]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
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||
is the only species of this subgenus from South-East Asia having no spines on its terga. It is easily separated from
|
||
<taxonomicName id="3EC23FC0FF85FFC48C05F9687D8F624C" authority="Braasch & Soldan, 1984" authorityName="Braasch & Soldan" authorityYear="1984" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinatus" subGenus="Belovius">
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<emphasis id="CBB69851FF85FFC48C05F9687E9C626A" box="[866,964,1767,1792]" italics="true" pageId="14" pageNumber="15">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="CBB69851FF85FFC48CB2F9687962626A" box="[981,1082,1767,1792]" italics="true" pageId="14" pageNumber="15">Belovius</emphasis>
|
||
)
|
||
<emphasis id="CBB69851FF85FFC48B2DF96879E1626A" box="[1098,1209,1767,1792]" italics="true" pageId="14" pageNumber="15">carinatus</emphasis>
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Braasch & Soldán, 1984
|
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</taxonomicName>
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, which has a row of submedian paired spines and from
|
||
<taxonomicName id="3EC23FC0FF85FFC48CE7F8837935624F" box="[896,1133,1804,1830]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="unicornutus" subGenus="Belovius">
|
||
<emphasis id="CBB69851FF85FFC48CE7F8837EC2624F" box="[896,922,1804,1829]" italics="true" pageId="14" pageNumber="15">E.</emphasis>
|
||
(
|
||
<emphasis id="CBB69851FF85FFC48CCBF8837E9E624F" box="[940,966,1804,1829]" italics="true" pageId="14" pageNumber="15">B.</emphasis>
|
||
)
|
||
<emphasis id="CBB69851FF85FFC48CBEF8837935624F" box="[985,1133,1804,1829]" italics="true" pageId="14" pageNumber="15">unicornutus</emphasis>
|
||
</taxonomicName>
|
||
, having a median row of single spines on the margins of the hind terga.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1D817C8FF85FFC58FA2F8D97F2962F2" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
|
||
<paragraph id="F97D4443FF85FFC58FA2F8D9797B64B6" blockId="14.[151,1437,1654,2017]" lastBlockId="15.[151,1436,152,476]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
<emphasis id="CBB69851FF85FFC48FA2F8D97C0A621A" bold="true" box="[197,338,1878,1904]" pageId="14" pageNumber="15">Discussion.</emphasis>
|
||
<bibRefCitation id="9D5339B2FF85FFC48E3CF8D87CA6621B" author="Kluge" box="[347,510,1879,1905]" pageId="14" pageNumber="15" refString="Kluge, N. (2004) The Phylogenetic System of Ephemeroptera. Kluwer Academic Publishers, Dordrecht, Boston, London, 442 pp." type="book" year="2004">Kluge (2004)</bibRefCitation>
|
||
, understanding
|
||
<taxonomicName id="3EC23FC0FF85FFC48DDBF8D87EAA621B" authority="Tshernova, 1981" authorityName="Tshernova" authorityYear="1981" box="[700,1010,1879,1905]" class="Insecta" family="Heptageniidae" genus="Belovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF85FFC48DDBF8D87E7A621A" box="[700,802,1879,1904]" italics="true" pageId="14" pageNumber="15">Belovius</emphasis>
|
||
<bibRefCitation id="9D5339B2FF85FFC48C4CF8D87EAA621B" author="Tshernova" box="[811,1010,1879,1905]" pageId="14" pageNumber="15" refString="Tshernova, O. A. (1981) K sistematike imago podenok roda Epeorus Eaton, 1881 (Ephemeroptea, Heptageniidae). Entomologicheskoe Obozrenie, 60, 323 - 336." type="journal article" year="1981">Tshernova, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
as a subgenus, transferred
|
||
<taxonomicName id="3EC23FC0FF85FFC48A5EF8D87C5462FF" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinatus">
|
||
<emphasis id="CBB69851FF85FFC48A5EF8D87C5462FF" italics="true" pageId="14" pageNumber="15">Epeorus carinatus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="81D504D3FF85FFC48E3EF8F37C9962FC" box="[345,449,1916,1942]" name="Vietnam" pageId="14" pageNumber="15">Vietnam</collectingCountry>
|
||
to
|
||
<taxonomicName id="3EC23FC0FF85FFC48E8DF8F37E5262FF" box="[490,778,1916,1942]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinatus" subGenus="Belovius">
|
||
<emphasis id="CBB69851FF85FFC48E8DF8F37F5D62FF" box="[490,517,1916,1941]" italics="true" pageId="14" pageNumber="15">E.</emphasis>
|
||
(
|
||
<emphasis id="CBB69851FF85FFC48D7FF8F37FDA62FF" box="[536,642,1916,1941]" italics="true" pageId="14" pageNumber="15">Belovius</emphasis>
|
||
)
|
||
<emphasis id="CBB69851FF85FFC48DF2F8F37E5262FF" box="[661,778,1916,1941]" italics="true" pageId="14" pageNumber="15">carinatus</emphasis>
|
||
</taxonomicName>
|
||
, characterized
|
||
<emphasis id="CBB69851FF85FFC48CAFF8F3796462FF" box="[968,1084,1916,1941]" italics="true" pageId="14" pageNumber="15">inter alia</emphasis>
|
||
by anal ribs on gills II–VII. Originally,
|
||
<bibRefCitation id="9D5339B2FF85FFC48E7BF82E7CB262D1" author="Tshernova" box="[284,490,1953,1979]" pageId="14" pageNumber="15" refString="Tshernova, O. A. (1981) K sistematike imago podenok roda Epeorus Eaton, 1881 (Ephemeroptea, Heptageniidae). Entomologicheskoe Obozrenie, 60, 323 - 336." type="journal article" year="1981">Tshernova (1981)</bibRefCitation>
|
||
published
|
||
<taxonomicName id="3EC23FC0FF85FFC48D0DF82E7F9762D0" box="[618,719,1953,1978]" class="Insecta" family="Heptageniidae" genus="Belovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF85FFC48D0DF82E7F9762D0" box="[618,719,1953,1978]" italics="true" pageId="14" pageNumber="15">Belovius</emphasis>
|
||
</taxonomicName>
|
||
as a distinct genus, later documented by
|
||
<bibRefCitation id="9D5339B2FF85FFC48BCAF82E78C362D1" author="Hubbard" box="[1197,1435,1953,1979]" pageId="14" pageNumber="15" refString="Hubbard, M. D. (1990) Mayflies of the world: A catalogue of the family and genus group taxa (Insecta: Ephemeroptera). Sandhill Crane Press, Gainesville, Florida, 119 pp." type="book" year="1990" yearSuffix="p">Hubbard (1990: p.7)</bibRefCitation>
|
||
in his ‘Mayflies of the World’ as “
|
||
<taxonomicName id="3EC23FC0FF85FFC48D4AF8487E0B628B" authority="Tshernova (As)" authorityName="Tshernova (As" box="[557,851,1991,2017]" class="Insecta" family="Heptageniidae" genus="Belovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF85FFC48D4AF8487FCA628A" box="[557,658,1991,2016]" italics="true" pageId="14" pageNumber="15">Belovius</emphasis>
|
||
Tshernova (As)
|
||
</taxonomicName>
|
||
”.
|
||
<bibRefCitation id="9D5339B2FF85FFC48C0BF848799F628B" author="Webb" box="[876,1223,1991,2017]" pageId="14" pageNumber="15" refString="Webb, J. M. & McCafferty, W. P. (2008) Heptageniidae of the World Part II: Key to the Genera. Canadian Journal of Arthropod Identification, 7, 1 - 55." type="journal article" year="2008">Webb and McCafferty (2008)</bibRefCitation>
|
||
did not recognize the subgenera of
|
||
<taxonomicName id="3EC23FC0FF84FFC58E04FF177C9D65DB" box="[355,453,152,177]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF84FFC58E04FF177C9D65DB" box="[355,453,152,177]" italics="true" pageId="15" pageNumber="16">Epeorus</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="3EC23FC0FF84FFC58D6EFF177F3665DB" box="[521,622,152,177]" class="Insecta" family="Heptageniidae" genus="Belovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF84FFC58D6EFF177F3665DB" box="[521,622,152,177]" italics="true" pageId="15" pageNumber="16">Belovius</emphasis>
|
||
</taxonomicName>
|
||
) and other for seeing them as paraphyletic. In our opinion, we agree with a statement by
|
||
<bibRefCitation id="9D5339B2FF84FFC58EF7FF327E1265BD" author="Wang" box="[400,842,189,215]" pageId="15" pageNumber="16" refString="Wang, T. - Q. & McCafferty, W. P. (2004) Heptageniidae (Ephemeroptera) of the World. Part I: Phylogenetic Higher Classification. Transactions of the American Entomological Society, 130, 1, 11 - 45." type="journal article" year="2004" yearSuffix="p">Wang and McCafferty (2004: p. 27)</bibRefCitation>
|
||
in their ‘Phylogenetic Higher Classification’ on
|
||
<taxonomicName id="3EC23FC0FF84FFC58FF0FF6D7C1E6596" box="[151,326,226,252]" class="Insecta" family="Heptageniidae" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Heptageniidae</taxonomicName>
|
||
: “As mentioned previously, the genus
|
||
<taxonomicName id="3EC23FC0FF84FFC58C76FF6D7E2B6591" box="[785,883,226,251]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF84FFC58C76FF6D7E2B6591" box="[785,883,226,251]" italics="true" pageId="15" pageNumber="16">Epeorus</emphasis>
|
||
</taxonomicName>
|
||
does not appear to be divisible into additional cladistically sound species groups, each with exclusive synapomorphies. However, the group is in severe need of a global revision at the species level, which could possibly yield data revealing such additional groupings. Within the strict rules of phylogenetic classification (
|
||
<bibRefCitation id="9D5339B2FF84FFC58C65FEDD7ED76406" author="Wiley" box="[770,911,338,364]" pageId="15" pageNumber="16" refString="Wiley, E. O. (1981) Phylogenetics. The Theory and Practice of Phylogenetic Systematics. Wiley-Interscience, New York, 439 pp." type="book" year="1981">Wiley 1981</bibRefCitation>
|
||
), additional genera would only be possible if they were sequentially derived with
|
||
<taxonomicName id="3EC23FC0FF84FFC58D58FEF77FF964FB" box="[575,673,376,401]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF84FFC58D58FEF77FF964FB" box="[575,673,376,401]" italics="true" pageId="15" pageNumber="16">Epeorus</emphasis>
|
||
</taxonomicName>
|
||
…” So far as we know, that study has not occurred and therefore, provisionally, we hold on to the validity of subgenus
|
||
<taxonomicName id="3EC23FC0FF84FFC58C76FE127E2E64DC" box="[785,886,413,438]" class="Insecta" family="Heptageniidae" genus="Belovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF84FFC58C76FE127E2E64DC" box="[785,886,413,438]" italics="true" pageId="15" pageNumber="16">Belovius</emphasis>
|
||
</taxonomicName>
|
||
within genus
|
||
<taxonomicName id="3EC23FC0FF84FFC58B78FE1279D964DC" box="[1055,1153,413,438]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF84FFC58B78FE1279D964DC" box="[1055,1153,413,438]" italics="true" pageId="15" pageNumber="16">Epeorus</emphasis>
|
||
</taxonomicName>
|
||
, but also to genus
|
||
<taxonomicName id="3EC23FC0FF84FFC58A04FE1278CD64DC" box="[1379,1429,413,438]" class="Insecta" family="Heptageniidae" genus="Iron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBB69851FF84FFC58A04FE1278CD64DC" box="[1379,1429,413,438]" italics="true" pageId="15" pageNumber="16">Iron</emphasis>
|
||
</taxonomicName>
|
||
. We are convinced that all conclusions regarding this are arbitrarily at present.
|
||
</paragraph>
|
||
<paragraph id="F97D4443FF84FFC58FF0F8CE7F2962F2" blockId="15.[151,1436,1857,1945]" pageId="15" pageNumber="16">
|
||
<emphasis id="CBB69851FF84FFC58FF0F8CE7C876233" bold="true" box="[151,479,1857,1881]" pageId="15" pageNumber="16">PLATE X. FIGURES 61–68</emphasis>
|
||
. Larva of
|
||
<taxonomicName id="3EC23FC0FF84FFC58D33F8CD7E9B6233" box="[596,963,1857,1881]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="khayengensis" status="sp. nov." subGenus="Belovius">
|
||
<emphasis id="CBB69851FF84FFC58D33F8CD7FE86233" box="[596,688,1858,1881]" italics="true" pageId="15" pageNumber="16">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="CBB69851FF84FFC58DA7F8CD7E476233" box="[704,799,1858,1881]" italics="true" pageId="15" pageNumber="16">Belovius</emphasis>
|
||
)
|
||
<emphasis id="CBB69851FF84FFC58C57F8CD7E9B6233" box="[816,963,1858,1881]" italics="true" pageId="15" pageNumber="16">khayengensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CBB69851FF84FFC58CACF8CE791F6233" bold="true" box="[971,1095,1857,1881]" pageId="15" pageNumber="16">
|
||
<taxonomicNameLabel id="D085252AFF84FFC58CACF8CE797C6233" box="[971,1060,1857,1881]" pageId="15" pageNumber="16" rank="species">
|
||
sp.
|
||
<date id="8D7C6283FF84FFC58C93F8CE797C6233" box="[1012,1060,1857,1881]" pageId="15" pageNumber="16" value="1961-11">nov.</date>
|
||
</taxonomicNameLabel>
|
||
61
|
||
</emphasis>
|
||
, Foreleg, left, dorsal view.
|
||
<emphasis id="CBB69851FF84FFC58A1DF8CE78CD6233" bold="true" box="[1402,1429,1857,1881]" pageId="15" pageNumber="16">62</emphasis>
|
||
, Bristles at face of foreleg femur.
|
||
<emphasis id="CBB69851FF84FFC58E99F8EE7F416213" bold="true" box="[510,537,1889,1913]" pageId="15" pageNumber="16">63</emphasis>
|
||
, Foreleg, tarsal claw.
|
||
<emphasis id="CBB69851FF84FFC58C6EF8EE7E7C6213" bold="true" box="[777,804,1889,1913]" pageId="15" pageNumber="16">64</emphasis>
|
||
, Abdominal tergum VII, posterior margin.
|
||
<emphasis id="CBB69851FF84FFC58B9DF8EE784D6213" bold="true" box="[1274,1301,1889,1913]" pageId="15" pageNumber="16">65</emphasis>
|
||
, Abdominal terga
|
||
<date id="8D7C6283FF84FFC58FB5F80E7C7C62F3" box="[210,292,1921,1945]" pageId="15" pageNumber="16" value="1966-01" valueMax="1966-10-31" valueMin="1966-01-00">
|
||
I–X.
|
||
<emphasis id="CBB69851FF84FFC58E6EF80E7C7C62F3" bold="true" box="[265,292,1921,1945]" pageId="15" pageNumber="16">66</emphasis>
|
||
</date>
|
||
, Gill
|
||
<date id="8D7C6283FF84FFC58E06F80E7CCA62F3" box="[353,402,1921,1945]" pageId="15" pageNumber="16" value="1967-01">
|
||
I.
|
||
<emphasis id="CBB69851FF84FFC58E10F80E7CCA62F3" bold="true" box="[375,402,1921,1945]" pageId="15" pageNumber="16">67</emphasis>
|
||
</date>
|
||
, Gill
|
||
<date id="8D7C6283FF84FFC58EA8F80E7F5F62F3" box="[463,519,1921,1945]" pageId="15" pageNumber="16" value="1968-05">
|
||
V.
|
||
<emphasis id="CBB69851FF84FFC58E8BF80E7F5F62F3" bold="true" box="[492,519,1921,1945]" pageId="15" pageNumber="16">68</emphasis>
|
||
</date>
|
||
, Gill VII.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1D817C8FF9BFFDA8FA2FF187FEF642D" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="F97D4443FF9BFFDA8FA2FF187FAF6596" blockId="16.[151,1438,151,327]" pageId="16" pageNumber="17">
|
||
<emphasis id="CBB69851FF9BFFDA8FA2FF187C9065DB" bold="true" box="[197,456,151,177]" pageId="16" pageNumber="17">Material examined:</emphasis>
|
||
<typeStatus id="2679FAE1FF9BFFDA8EB6FF177F2E65D8" box="[465,630,152,178]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
<emphasis id="CBB69851FF9BFFDA8DE5FF187E0465DB" bold="true" box="[642,860,151,177]" pageId="16" pageNumber="17">mature male La,</emphasis>
|
||
<collectingCountry id="81D504D3FF9BFFDA8C02FF177E8E65D8" box="[869,982,152,178]" name="Thailand" pageId="16" pageNumber="17">Thailand</collectingCountry>
|
||
, Kanchanaburi Province, Thong Pha Phum district, Huai Khayeng streams, alt.
|
||
<quantity id="3E3AE9A6FF9BFFDA8DB3FF327E7265BD" box="[724,810,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="16" pageNumber="17" unit="m" value="210.0">210 m</quantity>
|
||
,
|
||
<geoCoordinate id="9CF62284FF9BFFDA8C5DFF327E8E65BD" box="[826,982,189,215]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="14.585278">14º35'07"N</geoCoordinate>
|
||
, 98º34.9'37"E;
|
||
<date id="8D7C6283FF9BFFDA8BC0FF32781165BD" box="[1191,1353,189,215]" pageId="16" pageNumber="17" value="2008-12-13">13.XII.2008</date>
|
||
(BB);
|
||
<typeStatus id="2679FAE1FF9BFFDA8FF0FF6D7C666596" box="[151,318,226,252]" pageId="16" pageNumber="17" type="paratype">PARATYPES</typeStatus>
|
||
: 9 La (3 mature), same data as above.
|
||
</paragraph>
|
||
<paragraph id="F97D4443FF9BFFDA8FA2FE887FEF642D" blockId="16.[151,1438,151,327]" pageId="16" pageNumber="17">
|
||
<emphasis id="CBB69851FF9BFFDA8FA2FE887CCD644B" bold="true" box="[197,405,263,289]" pageId="16" pageNumber="17">
|
||
<typeStatus id="2679FAE1FF9BFFDA8FA2FE887C56644B" box="[197,270,263,289]" pageId="16" pageNumber="17">Types</typeStatus>
|
||
deposition
|
||
</emphasis>
|
||
:
|
||
<typeStatus id="2679FAE1FF9BFFDA8EC3FE877F1C6448" box="[420,580,264,290]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
<emphasis id="CBB69851FF9BFFDA8D29FE887E4D644B" bold="true" box="[590,789,263,289]" pageId="16" pageNumber="17">mature male La</emphasis>
|
||
(
|
||
<emphasis id="CBB69851FF9BFFDA8C41FE887ED9644B" bold="true" box="[806,897,263,289]" pageId="16" pageNumber="17">
|
||
<collectionCode id="9FD3DC86FF9BFFDA8C41FE887ED9644B" box="[806,897,263,289]" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="16" pageNumber="17">ZMKU</collectionCode>
|
||
</emphasis>
|
||
, 80 % alcohol);
|
||
<typeStatus id="2679FAE1FF9BFFDA8B25FE8779B16448" box="[1090,1257,264,290]" pageId="16" pageNumber="17" type="paratype">PARATYPES</typeStatus>
|
||
: 7 La (
|
||
<emphasis id="CBB69851FF9BFFDA8A5EFE8878CC644B" bold="true" box="[1337,1428,263,289]" pageId="16" pageNumber="17">
|
||
<collectionCode id="9FD3DC86FF9BFFDA8A5EFE8878CC644B" box="[1337,1428,263,289]" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="16" pageNumber="17">ZMKU</collectionCode>
|
||
</emphasis>
|
||
, 80 % alcohol);
|
||
<specimenCount id="EFC48FCAFF9BFFDA8E2DFEA27CC1642D" box="[330,409,301,327]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
|
||
La,
|
||
<specimenCount id="EFC48FCAFF9BFFDA8EA9FEA27F6C642D" box="[462,564,301,327]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
|
||
La (
|
||
<emphasis id="CBB69851FF9BFFDA8D0BFEA27FFF642D" bold="true" box="[620,679,301,327]" pageId="16" pageNumber="17">DBP</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |