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<document ID-DOI="http://doi.org/10.5281/zenodo.6280338" ID-GBIF-Dataset="647046fe-0ee5-401d-b4f8-aea0355fde7f" ID-GBIF-Taxon="124407469" ID-Zenodo-Dep="6280338" checkinTime="1471630429166" checkinUser="plazi" docAuthor="Piotr Naskrecki" docDate="2000" docId="634387D1A304FFE016EDFC48FBAA3A6F" docLanguage="en" docName="Naskrecki_KCR_2000.pdf.imf" docOrigin="Philadelphia, PA: The Orthopterists Society at the Academy of Natural Sciences of Philadelphia," docStyle="DocumentStyle{}" docTitle="Copiphora ottei Naskrecki, sp. n." docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="9F7AFFA9A32AFFCE167AFFC3FFF13C6E" masterDocTitle="Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato)." pageId="46" pageNumber="43" updateTime="1645797358202" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato).</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Piotr Naskrecki</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:originInfo>
<mods:dateIssued>2000</mods:dateIssued>
<mods:publisher>The Orthopterists Society at the Academy of Natural Sciences of Philadelphia,</mods:publisher>
<mods:place>
<mods:placeTerm>Philadelphia, PA</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:classification>book</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.270035</mods:identifier>
<mods:identifier type="GBIF-Dataset">647046fe-0ee5-401d-b4f8-aea0355fde7f</mods:identifier>
<mods:identifier type="Zenodo-Dep">270035</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6280338" ID-GBIF-Taxon="124407469" ID-Zenodo-Dep="6280338" LSID="urn:lsid:plazi:treatment:634387D1A304FFE016EDFC48FBAA3A6F" httpUri="http://treatment.plazi.org/id/634387D1A304FFE016EDFC48FBAA3A6F" lastPageNumber="43" pageId="46" pageNumber="43">
<subSubSection box="[151,503,907,930]" pageId="46" pageNumber="43" type="nomenclature">
<paragraph blockId="46.[151,526,907,991]" box="[151,503,907,930]" pageId="46" pageNumber="43">
<taxonomicName authority="Naskrecki" authorityName="Naskrecki" box="[151,438,907,930]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="43" phylum="Arthropoda" rank="species" species="ottei" status="sp. n.">
<emphasis bold="true" box="[151,322,907,929]" italics="true" pageId="46" pageNumber="43">Copiphora ottei</emphasis>
Naskrecki
</taxonomicName>
,
<taxonomicNameLabel box="[447,503,908,929]" pageId="46" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="43" type="description">
<paragraph blockId="46.[151,526,907,991]" box="[151,526,938,960]" pageId="46" pageNumber="43">Common name: Ottes spear bearer</paragraph>
<paragraph blockId="46.[151,526,907,991]" box="[151,391,969,991]" pageId="46" pageNumber="43">
<figureCitation box="[151,219,969,991]" captionStart="FIG. 9" captionStartId="40.[151,195,1684,1706]" captionTargetBox="[136,1376,204,1621]" captionTargetId="figure@40.[136,1377,204,1622]" captionTargetPageId="40" captionText="FIG. 9. Copiphora ottei. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. male cerci, dorsal view." httpUri="https://zenodo.org/record/270044/files/figure.png" pageId="46" pageNumber="43">Figs. 9</figureCitation>
A-D, 42F,
<figureCitation box="[324,391,969,991]" captionStart="MAPS 1 - 6" captionStartId="157.[174,246,1655,1677]" captionTargetBox="[264,1285,204,1639]" captionTargetId="figure@157.[264,1285,204,1639]" captionTargetPageId="157" captionText="MAPS 1 - 6. Distribution of species of Conocephalinae s. l. in Costa Rica. 1. Conocephalus angustifrons, C. cinereus, 2. Conocephalus saltator, C. ictus, 3. Conocephalus magdalenae, Orchelimum fraternum, Lipotactomimus rowelli, 4. Lirometopum coronatum, Acantheremus colwelli, 5. Copiphora rhinoceros, C. cochleata, C. brevicauda costaricensis, 6. Copiphora hastata, C. cultricornis, C. ottei." httpUri="https://zenodo.org/record/270095/files/figure.png" pageId="46" pageNumber="43">Map 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="43" type="materials_examined">
<paragraph blockId="46.[151,706,1029,1112]" pageId="46" pageNumber="43">
<typeStatus box="[151,203,1030,1051]" pageId="46" pageNumber="43">Type</typeStatus>
locality:
<collectingCountry box="[299,414,1029,1051]" name="Costa Rica" pageId="46" pageNumber="43">Costa Rica</collectingCountry>
:
<collectingRegion box="[422,542,1030,1051]" country="Costa Rica" name="Puntarenas" pageId="46" pageNumber="43">Puntarenas</collectingRegion>
Prov., Golfito;
<typeStatus box="[209,256,1060,1081]" pageId="46" pageNumber="43">type</typeStatus>
depository: Academy of Natural Sciences, Philadelphia
<typeStatus box="[368,462,1090,1112]" pageId="46" pageNumber="43">holotype</typeStatus>
male
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="43" type="description">
<paragraph blockId="46.[151,740,1151,1720]" pageId="46" pageNumber="43">
<emphasis box="[174,420,1151,1173]" italics="true" pageId="46" pageNumber="43">Diagnostic description.—</emphasis>
Body small compared to other members of the genus, with wings in both sexes extending beyond apex of abdomen by only about 1/4 of their length (
<figureCitation box="[238,303,1242,1264]" captionStart="FIG. 9" captionStartId="40.[151,195,1684,1706]" captionTargetBox="[136,1376,204,1621]" captionTargetId="figure@40.[136,1377,204,1622]" captionTargetPageId="40" captionText="FIG. 9. Copiphora ottei. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. male cerci, dorsal view." httpUri="https://zenodo.org/record/270044/files/figure.png" pageId="46" pageNumber="43">Fig. 9</figureCitation>
A). Fastigium of vertex forming small, trituberculate horn, with lateral ocelli situated on conspicuous lateral protuberances; fastigium about as long as eye diameter; dorsal part of fastigium smooth or with two rows of poorly developed, small tubercles (
<figureCitation box="[651,710,1364,1386]" captionStart="FIG. 9" captionStartId="40.[151,195,1684,1706]" captionTargetBox="[136,1376,204,1621]" captionTargetId="figure@40.[136,1377,204,1622]" captionTargetPageId="40" captionText="FIG. 9. Copiphora ottei. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. male cerci, dorsal view." httpUri="https://zenodo.org/record/270044/files/figure.png" pageId="46" pageNumber="43">Fig. 9</figureCitation>
C). Face flat; genal carinae of head well developed, granulose. Pronotum smooth, flat dorsally; anterior dorsal margin straight to weakly concave, posterior one weakly convex; lateral lobe of pronotum about 1.8 times longer than high. Male stridulatory file nearly straight,
<quantity box="[151,234,1547,1568]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="46" pageNumber="43" unit="mm" value="2.3">2.3 mm</quantity>
long, 223 μm wide, with 214 thin, wide, and evenly distributed teeth (
<figureCitation box="[421,495,1577,1599]" captionStart="FIG. 42" captionStartId="140.[151,195,1684,1706]" captionTargetBox="[204,1305,204,1639]" captionTargetId="figure@140.[204,1306,204,1640]" captionTargetPageId="140" captionText="FIG. 42. Stridulatory files of species of Lirometopum and Copiphora. A. L. coronatum, B. C. cultricornis, C. C. hastata, D. C. brevicauda brevicauda, E. C. brevicauda costaricensis, F. C. ottei." httpUri="https://zenodo.org/record/270078/files/figure.png" pageId="46" pageNumber="43">Fig. 42</figureCitation>
F); mirror nearly circular; small, parallel vein next to AA1 reduced to hardly discernible fold.
</paragraph>
<paragraph blockId="46.[151,740,1151,1720]" lastBlockId="46.[771,1360,208,747]" pageId="46" pageNumber="43">
Ventral spines of
<taxonomicName box="[373,450,1668,1690]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="43" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
relatively short and thin, green; all genicular lobes, except for posterior lobe of fore
<taxonomicName box="[771,848,208,230]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="43" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
, armed with sharp spines. Male cercus thick (although not as much as in other species of the genus), blunt, and distinctly incurved; ventral, subapical spine of cercus long, blunt apically (
<figureCitation box="[1092,1152,299,321]" captionStart="FIG. 9" captionStartId="40.[151,195,1684,1706]" captionTargetBox="[136,1376,204,1621]" captionTargetId="figure@40.[136,1377,204,1622]" captionTargetPageId="40" captionText="FIG. 9. Copiphora ottei. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. male cerci, dorsal view." httpUri="https://zenodo.org/record/270044/files/figure.png" pageId="46" pageNumber="43">Fig. 9</figureCitation>
D). Male subgenital plate weakly narrowed toward apex, apex weakly emarginated; styli short, about 3 times as long as wide. Female subgenital plate approximately triangular, with minute, triangular apical incision. Ovipositor long, straight, ratio ovipositor/hind femur 1.4-1.6 (
<figureCitation box="[1250,1311,451,473]" captionStart="FIG. 9" captionStartId="40.[151,195,1684,1706]" captionTargetBox="[136,1376,204,1621]" captionTargetId="figure@40.[136,1377,204,1622]" captionTargetPageId="40" captionText="FIG. 9. Copiphora ottei. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. male cerci, dorsal view." httpUri="https://zenodo.org/record/270044/files/figure.png" pageId="46" pageNumber="43">Fig. 9</figureCitation>
).
</paragraph>
<paragraph blockId="46.[771,1360,208,747]" pageId="46" pageNumber="43">
<emphasis box="[794,897,482,503]" italics="true" pageId="46" pageNumber="43">Coloration</emphasis>
.— General coloration light olive green. Fastigium of vertex, face and mouthparts yellowish-orange to pinkish-brown, face often with 4 or 6 small, dark dots (
<figureCitation box="[779,837,573,595]" captionStart="FIG. 9" captionStartId="40.[151,195,1684,1706]" captionTargetBox="[136,1376,204,1621]" captionTargetId="figure@40.[136,1377,204,1622]" captionTargetPageId="40" captionText="FIG. 9. Copiphora ottei. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. male cerci, dorsal view." httpUri="https://zenodo.org/record/270044/files/figure.png" pageId="46" pageNumber="43">Fig. 9</figureCitation>
B). Pronotum brown to dark olive green. Tegmina green, with conspicuous yellow venation; edges of tegmina red, especially in their apical portions; male stridulatory file green; legs brown, with green femoral spines; hind
<taxonomicName box="[827,902,694,716]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="43" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
usually with well developed reticulate pattern on their outer surfaces. Abdomen green.
</paragraph>
<paragraph blockId="46.[794,1053,786,808]" box="[794,1053,786,808]" pageId="46" pageNumber="43">
<emphasis box="[794,936,786,807]" italics="true" pageId="46" pageNumber="43">Measurements</emphasis>
.—
<tableCitation box="[973,1050,786,808]" captionStart="TABLE 4" captionStartId="33.[174,254,207,228]" captionTargetBox="[174,1380,265,1093]" captionTargetPageId="33" captionText="TABLE 4. Body measurements of species of Copiphora (all measurements are lengths in mm: range, mean ± SD)" httpUri="http://table.plazi.org/id/BF95664FA30BFFEF16D4FF0CFAC63C8B" pageId="46" pageNumber="43" tableUuid="BF95664FA30BFFEF16D4FF0CFAC63C8B">Table 4</tableCitation>
.
</paragraph>
<paragraph blockId="46.[794,1108,846,868]" box="[794,1108,846,868]" pageId="46" pageNumber="43">
<emphasis box="[794,944,847,868]" italics="true" pageId="46" pageNumber="43">Bioacoustics.—</emphasis>
Call unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="43" type="distribution">
<paragraph blockId="46.[771,1360,907,1051]" pageId="46" pageNumber="43">
<emphasis box="[794,946,907,928]" italics="true" pageId="46" pageNumber="43">Distribution.—</emphasis>
This new species seems to be restricted in its distribution to the southernmost portion of
<collectingRegion box="[771,894,969,990]" country="Costa Rica" name="Puntarenas" pageId="46" pageNumber="43">Puntarenas</collectingRegion>
Prov. (Osa Peninsula and adjacent areas) (
<figureCitation box="[779,848,998,1020]" captionStart="MAPS 1 - 6" captionStartId="157.[174,246,1655,1677]" captionTargetBox="[264,1285,204,1639]" captionTargetId="figure@157.[264,1285,204,1639]" captionTargetPageId="157" captionText="MAPS 1 - 6. Distribution of species of Conocephalinae s. l. in Costa Rica. 1. Conocephalus angustifrons, C. cinereus, 2. Conocephalus saltator, C. ictus, 3. Conocephalus magdalenae, Orchelimum fraternum, Lipotactomimus rowelli, 4. Lirometopum coronatum, Acantheremus colwelli, 5. Copiphora rhinoceros, C. cochleata, C. brevicauda costaricensis, 6. Copiphora hastata, C. cultricornis, C. ottei." httpUri="https://zenodo.org/record/270095/files/figure.png" pageId="46" pageNumber="43">Map 6</figureCitation>
). It is possible, however, that this species also occurs in the north-western part of
<collectingCountry box="[1122,1211,1029,1050]" name="Panama" pageId="46" pageNumber="43">
<collectingRegion box="[1122,1211,1029,1050]" country="Panama" name="Panama" pageId="46" pageNumber="43">Panama</collectingRegion>
</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="43" type="materials_examined">
<paragraph blockId="46.[771,1362,1089,1416]" pageId="46" pageNumber="43">
<emphasis box="[794,1005,1090,1111]" italics="true" pageId="46" pageNumber="43">Material examined.—</emphasis>
<emphasis bold="true" box="[1010,1354,1089,1111]" pageId="46" pageNumber="43">
<collectingCountry box="[1010,1163,1089,1111]" name="Costa Rica" pageId="46" pageNumber="43">COSTA RICA</collectingCountry>
:
<collectingRegion box="[1172,1295,1090,1111]" country="Costa Rica" name="Puntarenas" pageId="46" pageNumber="43">Puntarenas</collectingRegion>
Prov.
</emphasis>
, Corcovado N. P., Península de Osa,
<date box="[1143,1303,1120,1141]" pageId="46" pageNumber="43" value="1978-08-14">14 August 1978</date>
(coll. D.H. Janzen) -
<specimenCount box="[921,1008,1150,1172]" pageId="46" pageNumber="43" type="female">1 female</specimenCount>
(
<typeStatus box="[1020,1115,1151,1173]" pageId="46" pageNumber="43">paratype</typeStatus>
) (
<collectionCode box="[1134,1200,1150,1172]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="46" pageNumber="43">INBio</collectionCode>
); Estac. Aguias, Río Agujas, Sendero Ajo, elev.
<quantity box="[1085,1148,1181,1202]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="46" pageNumber="43" unit="m" value="300.0">300 m</quantity>
,
<date box="[1157,1304,1181,1203]" pageId="46" pageNumber="43" value="1997-08-05">5 August 1997</date>
(coll. A. Azofeifa) -
<specimenCount box="[915,1003,1211,1233]" pageId="46" pageNumber="43" type="female">1 female</specimenCount>
(
<typeStatus box="[1015,1111,1211,1233]" pageId="46" pageNumber="43">paratype</typeStatus>
) (
<collectionCode box="[1131,1198,1211,1233]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="46" pageNumber="43">INBio</collectionCode>
); Estac. Aguias, Río Agujas, Sendero Purruja, elev.
<quantity box="[1129,1193,1242,1263]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="46" pageNumber="43" unit="m" value="300.0">300 m</quantity>
,
<date pageId="46" pageNumber="43" value="1997-09-01" valueMax="1997-09-06" valueMin="1997-09-01">1 - 6 September 1997</date>
(coll. A. Azofeifa) -
<specimenCount box="[1050,1143,1272,1294]" pageId="46" pageNumber="43" type="female">1 female</specimenCount>
(
<typeStatus box="[1160,1258,1272,1294]" pageId="46" pageNumber="43">paratype</typeStatus>
) (
<collectionCode box="[1281,1349,1272,1294]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="46" pageNumber="43">INBio</collectionCode>
); Golfito,
<date box="[860,1014,1302,1324]" pageId="46" pageNumber="43" value="1957-08-09">9 August 1957</date>
(coll. A. Menke) -
<specimenCount box="[1217,1289,1302,1324]" pageId="46" pageNumber="43" type="male">1 male</specimenCount>
(
<typeStatus pageId="46" pageNumber="43">holotype</typeStatus>
) (
<collectionCode box="[839,909,1333,1355]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="46" pageNumber="43">ANSP</collectionCode>
); Península de
<taxonomicName authority="Corcovado N. P., Estac. Sirena" authorityName="Corcovado N. P., Estac. Sirena" class="Magnoliopsida" family="Rubiaceae" genus="Osa" kingdom="Plantae" order="Gentianales" pageId="46" pageNumber="43" phylum="Tracheophyta" rank="genus">Osa, Corcovado N. P., Estac. Sirena</taxonomicName>
,
<date box="[862,1045,1363,1385]" pageId="46" pageNumber="43" value="1980-08-13">13 August 1980</date>
(coll. D.H. Janzen and W. Hallwachs) -
<specimenCount box="[912,1001,1394,1416]" pageId="46" pageNumber="43" type="female">1 female</specimenCount>
(
<typeStatus box="[1015,1103,1394,1416]" pageId="46" pageNumber="43">allotype</typeStatus>
) (
<collectionCode box="[1123,1193,1394,1416]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="46" pageNumber="43">ANSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="43" type="etymology">
<paragraph blockId="46.[771,1360,1455,1537]" pageId="46" pageNumber="43">
<emphasis box="[794,928,1455,1476]" italics="true" pageId="46" pageNumber="43">Etymology.—</emphasis>
This new species is named in honor of Daniel Otte, for his prodigious contributions to biology, and to entomology in particular.
</paragraph>
</subSubSection>
</treatment>
</document>