329 lines
37 KiB
XML
329 lines
37 KiB
XML
<document id="598F68EF5A8AFCDF71462E668ED3ECF1" ID-DOI="10.15298/rusentj.28.1.05" ID-ISSN="0132-8069" ID-Zenodo-Dep="10919898" checkinTime="1712198159115" checkinUser="felipe" docAuthor="Gildenkov, M. Yu." docDate="2019" docId="071D20116A68FF87FF123649EB89FCD4" docLanguage="en" docName="RussEntomolJ.28.1.30-35.pdf" docOrigin="Russian Entomological Journal 28 (1)" docSource="http://dx.doi.org/10.15298/rusentj.28.1.05" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docTitle="Carpelimus (Trogophloeus) vilisus Gildenkov 2019, sp.n." docType="treatment" docVersion="1" lastPageNumber="33" masterDocId="FB2458696A6AFF84FFDA3772EA12FFC2" masterDocTitle="Five new species of the Genus Carpelimus Leach, 1819 from Thailand and the Philippines (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="35" masterPageNumber="30" pageNumber="32" updateTime="1712198473155" updateUser="ExternalLinkService">
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<mods:titleInfo id="A9A7103D92FCEE708435D63FE0019A83">
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<mods:title id="8C00FEDD9EC88869A53372EA939D467E">Five new species of the Genus Carpelimus Leach, 1819 from Thailand and the Philippines (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:name id="7FE051B5EAA1CA723520E76DFD5D6378" type="personal">
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<mods:namePart id="9C206268C4FE8797E359ED9CBDF437F1">Gildenkov, M. Yu.</mods:namePart>
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<mods:title id="AB1CA2186D60332AE0B5D1B5C7A62241">Russian Entomological Journal</mods:title>
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<mods:part id="0BD3AE341A8F7CC0EFAEDE3C95C6A81C">
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<mods:date id="F58EEFCB2283CDBF40A4DE9E61708EAF">2019</mods:date>
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<mods:number id="73B8DEFFD704C4E823FF98A4969AC61C">2019-09-30</mods:number>
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<mods:number id="EF7E43246C0D9429C4622924392D1AD0">28</mods:number>
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<mods:url id="1DD694D016F4A70DEAB8E4B235C5D5CA">http://dx.doi.org/10.15298/rusentj.28.1.05</mods:url>
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<mods:identifier id="5C32B57CC15A14E6A26145B486674F64" type="DOI">10.15298/rusentj.28.1.05</mods:identifier>
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<mods:identifier id="92DABA445E22C3F884A417C7BD1EECE9" type="ISSN">0132-8069</mods:identifier>
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<mods:identifier id="0B6699D50676C5F5B04DF07D4B33AF01" type="Zenodo-Dep">10919898</mods:identifier>
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<treatment id="071D20116A68FF87FF123649EB89FCD4" LSID="urn:lsid:plazi:treatment:071D20116A68FF87FF123649EB89FCD4" httpUri="http://treatment.plazi.org/id/071D20116A68FF87FF123649EB89FCD4" lastPageId="3" lastPageNumber="33" pageId="2" pageNumber="32">
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<subSubSection id="C7AEC28C6A68FF86FF123649E8E9FE91" box="[200,763,315,339]" pageId="2" pageNumber="32" type="nomenclature">
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<paragraph id="8F0B91076A68FF86FF123649E8E9FE91" blockId="2.[189,776,315,819]" box="[200,763,315,339]" pageId="2" pageNumber="32">
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<heading id="D443266B6A68FF86FF123649E8E9FE91" box="[200,763,315,339]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="32" reason="8">
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<taxonomicName id="48B4EA846A68FF86FF123649E8A8FE91" ID-CoL="8XWNV" authority="Gildenkov" authorityName="Gildenkov" authorityYear="2019" box="[200,698,315,339]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="vilisus" status="sp. nov." subGenus="Trogophloeus">
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<emphasis id="BDC04D156A68FF86FF123649EB56FE91" box="[200,324,315,339]" italics="true" pageId="2" pageNumber="32">Carpelimus</emphasis>
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(
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<emphasis id="BDC04D156A68FF86FE8F3649EBFBFE91" box="[341,489,315,339]" italics="true" pageId="2" pageNumber="32">Trogophloeus</emphasis>
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)
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<emphasis id="BDC04D156A68FF86FE203649E82CFE91" box="[506,574,315,339]" italics="true" pageId="2" pageNumber="32">vilisus</emphasis>
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Gildenkov
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</taxonomicName>
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,
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||
<emphasis id="BDC04D156A68FF86FD1C3649E8E9FE91" bold="true" box="[710,763,315,339]" pageId="2" pageNumber="32">
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<taxonomicNameLabel id="A6F3F06E6A68FF86FD1C3649E8E9FE91" box="[710,763,315,339]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C7AEC28C6A68FF86FE78362BE833FEB3" box="[418,545,345,369]" pageId="2" pageNumber="32" type="description">
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<paragraph id="8F0B91076A68FF86FE78362BE833FEB3" blockId="2.[189,776,315,819]" box="[418,545,345,369]" pageId="2" pageNumber="32">
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<figureCitation id="178F8D826A68FF86FE78362BEBF4FEB3" box="[418,486,345,369]" captionStart="Figs 1–4" captionStartId="1.[241,278,1974,1993]" captionTargetBox="[192,1393,1029,1946]" captionTargetId="figure-534@1.[189,1399,1023,1948]" captionTargetPageId="1" captionText="Figs 1–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n. Рис. 1–4. Carpelimus (Trogophloeus) spp., голотипы, самцы, сверху: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n." figureDoi="http://doi.org/10.5281/zenodo.10919902" httpUri="https://zenodo.org/record/10919902/files/figure.png" pageId="2" pageNumber="32">Figs 2</figureCitation>
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,
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<figureCitation id="178F8D826A68FF86FE28362BE80FFEB3" box="[498,541,345,369]" captionStart="Figs 5–14" captionStartId="2.[226,263,1942,1961]" captionTargetBox="[193,1399,842,1891]" captionTargetId="figure-475@2.[189,1399,839,1891]" captionTargetPageId="2" captionText="Figs 5–14. Genitalia of Carpelimus (Trogophloeus) spp.: 5–7 — C. notumus, sp.n.; 8–9 — C. vilisus, sp.n.; 10–11 — C. plenus, sp.n.; 12–14 — C. notatus, sp.n.; 5, 8, 10, 12 — aedeagus, ventral view (holotypes); 6, 9, 11, 13 — aedeagus, lateral view (holotypes); 7, 14 – spermatheca (paratypes). Scale bar: 0.25 mm. Рис. 5–14. Гениталии Carpelimus (Trogophloeus) spp.: 5–7 — C. notumus, sp.n.; 8–9 — C. vilisus, sp.n.; 10–11 — C. plenus, sp.n.; 12– 14 — C. notatus, sp.n.; 5, 8, 10, 12 — Эдеагус, сниЗу (голотипы); 6, 9, 11, 13 — Эдеагус, сбоку (голотипы); 7, 14 — сперматека (паратипы). МасШтаб: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10919904" httpUri="https://zenodo.org/record/10919904/files/figure.png" pageId="2" pageNumber="32">8, 9</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C7AEC28C6A68FF86FF38360CE893FE1B" pageId="2" pageNumber="32" type="materials_examined">
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<paragraph id="8F0B91076A68FF86FF38360CE893FE1B" blockId="2.[189,776,315,819]" pageId="2" pageNumber="32">
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<materialsCitation id="3FDC9B5A6A68FF86FF38360CE86FFE1B" collectingDate="1995-11-16" collectingDateMax="1995-11-18" collectingDateMin="1995-11-16" collectionCode="NHMW" collectorName="Zettel" country="Thailand" elevation="350" location="Mae Khaem" municipality="Huai Kaet" pageId="2" pageNumber="32" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Phrae" typeStatus="holotype">
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MATERIAL.
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<typeStatus id="500F2FA56A68FF86FE81360CEBBFFE53" box="[347,429,382,401]" pageId="2" pageNumber="32" type="holotype">Holotype</typeStatus>
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||
,
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<specimenCount id="99B25A8E6A68FF86FE6F3609EBDBFE53" box="[437,457,379,401]" count="1" pageId="2" pageNumber="32" type="male">♂</specimenCount>
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“
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<collectingCountry id="F7A3D1976A68FF86FE0C360CE85BFE53" box="[470,585,382,401]" name="Thailand" pageId="2" pageNumber="32">THAILAND</collectingCountry>
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:17./
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<date id="FB0AB7C76A68FF86FDB6360FE8C3FE52" box="[620,721,381,400]" pageId="2" pageNumber="32" value="1995-11-18">
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<collectingDate id="EB4E4E2F6A68FF86FDB6360FE8C3FE52" box="[620,721,381,400]" pageId="2" pageNumber="32" value="1995-11-18">18.11.1995</collectingDate>
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</date>
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<collectingRegion id="4D705FE56A68FF86FD0F360CEAFCFE6B" country="Thailand" name="Phrae" pageId="2" pageNumber="32">Phrae Prov.</collectingRegion>
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,
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<collectingMunicipality id="6F6F0B7D6A68FF86FF2036E4EB45FE6B" box="[250,343,406,425]" pageId="2" pageNumber="32">Huai Kaet</collectingMunicipality>
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<quantity id="484C3CE26A68FF86FEBB36E7EB80FE6A" box="[353,402,405,424]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="32" unit="km" value="50.0">50km</quantity>
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NE
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<collectingRegion id="4D705FE56A68FF86FE1936E4EBE4FE6B" box="[451,502,406,425]" country="Thailand" name="Phrae" pageId="2" pageNumber="32">Phrae</collectingRegion>
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, leg.
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<collectorName id="2241F4D16A68FF86FDF736E4E84DFE6B" box="[557,607,406,425]" pageId="2" pageNumber="32">Zettel</collectorName>
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||
(17b)” (
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||
<collectionCode id="E9A509C26A68FF86FD6836E4E8ECFE6B" box="[690,766,406,425]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="32" type="Museum">NHMW</collectionCode>
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).
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<typeStatus id="500F2FA56A68FF86FF6736DCEB1EFE03" box="[189,268,430,449]" pageId="2" pageNumber="32" type="paratype">Paratype</typeStatus>
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||
:
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<specimenCount id="99B25A8E6A68FF86FECC36DCEB3FFE03" box="[278,301,427,449]" count="1" pageId="2" pageNumber="32" type="female">1♀</specimenCount>
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||
“
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||
<collectingCountry id="F7A3D1976A68FF86FEE436DCEBA3FE03" box="[318,433,430,449]" name="Thailand" pageId="2" pageNumber="32">THAILAND</collectingCountry>
|
||
:
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||
<date id="FB0AB7C76A68FF86FE6636DFE80FFE02" box="[444,541,429,448]" pageId="2" pageNumber="32" value="1995-11-16">
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||
<collectingDate id="EB4E4E2F6A68FF86FE6636DFE80FFE02" box="[444,541,429,448]" pageId="2" pageNumber="32" value="1995-11-16">16.11.1995</collectingDate>
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||
</date>
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||
<collectingRegion id="4D705FE56A68FF86FDFF36DCE882FE03" box="[549,656,430,449]" country="Thailand" name="Phrae" pageId="2" pageNumber="32">Phrae Prov.</collectingRegion>
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||
,
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||
<location id="8A6BC7DC6A68FF86FD4036DCE914FE03" LSID="urn:lsid:plazi:treatment:071D20116A68FF87FF123649EB89FCD4:8A6BC7DC6A68FF86FD4036DCE914FE03" box="[666,774,430,449]" country="Thailand" municipality="Huai Kaet" name="Mae Khaem" pageId="2" pageNumber="32" stateProvince="Phrae">Mae Khaem</location>
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||
<quantity id="484C3CE26A68FF86FF6736B7EAFCFE1A" box="[189,238,453,472]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="32" unit="km" value="15.0">15km</quantity>
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||
E
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<collectingRegion id="4D705FE56A68FF86FED236B4EB29FE1B" box="[264,315,454,473]" country="Thailand" name="Phrae" pageId="2" pageNumber="32">Phrae</collectingRegion>
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, ca.
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||
<quantity id="484C3CE26A68FF86FEBE36B7EB8AFE1A" box="[356,408,453,472]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="32" unit="m" value="350.0">
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<elevation id="049976346A68FF86FEBE36B7EB8AFE1A" box="[356,408,453,472]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="32" unit="m" value="350.0">350m</elevation>
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</quantity>
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, leg.
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<collectorName id="2241F4D16A68FF86FE1236B4EBE8FE1B" box="[456,506,454,473]" pageId="2" pageNumber="32">Zettel</collectorName>
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(16b)” (cMG)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C7AEC28C6A68FF87FF3836ACE894FD6E" lastPageId="3" lastPageNumber="33" pageId="2" pageNumber="32" type="description">
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<paragraph id="8F0B91076A68FF86FF3836ACE845FD81" blockId="2.[189,776,315,819]" pageId="2" pageNumber="32">
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DESCRIPTION (
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<typeStatus id="500F2FA56A68FF86FE5136AFEBF7FE31" box="[395,485,477,499]" pageId="2" pageNumber="32" type="holotype">holotype</typeStatus>
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||
). Length
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<quantity id="484C3CE26A68FF86FDE536ACE899FE31" box="[575,651,478,499]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="32" unit="mm" value="1.8">1.8 mm</quantity>
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. Colouration dark brown, legs and antennal bases yellow brown, darker to light brown towards antennal apex. Integument slightly shining, body with short, light-coloured hairs.
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</paragraph>
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<paragraph id="8F0B91076A68FF86FF38353AE91AFCF1" blockId="2.[189,776,315,819]" pageId="2" pageNumber="32">
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Head transverse, with a wide base, ratio of its length (from posterior margin of head to anterior margin of clypeus) to maximum width about 16:24. Neck constriction prominent. Eyes large, convex. Temples well-developed, round, eye diameter in dorsal view about 2.5 times as long as temple length. Head widest across eyes (
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<figureCitation id="178F8D826A68FF86FE6E35BFEBFDFD26" box="[436,495,717,740]" captionStart="Figs 1–4" captionStartId="1.[241,278,1974,1993]" captionTargetBox="[192,1393,1029,1946]" captionTargetId="figure-534@1.[189,1399,1023,1948]" captionTargetPageId="1" captionText="Figs 1–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n. Рис. 1–4. Carpelimus (Trogophloeus) spp., голотипы, самцы, сверху: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n." figureDoi="http://doi.org/10.5281/zenodo.10919902" httpUri="https://zenodo.org/record/10919902/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
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). Head surface with delicate, fine and dense punctation. Diameter of punctures about 3.0 times as small as eye facet. Distances between punctures about equal to their diameter, interspaces smooth, slightly shining.
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</paragraph>
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<paragraph id="8F0B91076A68FF86FEE73017E861F8BD" blockId="2.[317,627,1893,1919]" box="[317,627,1893,1919]" pageId="2" pageNumber="32">
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||
<emphasis id="BDC04D156A68FF86FEE73017E861F8BD" bold="true" box="[317,627,1893,1919]" pageId="2" pageNumber="32">8 9</emphasis>
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</paragraph>
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<paragraph id="8F0B91076A68FF86FCF63649EF2BFE63" blockId="2.[812,1399,315,817]" pageId="2" pageNumber="32">
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Antennae rather short, antennal segments 1–3 elongate; segments 4–7 about as long as wide; segments 8–10 slightly transverse; segment 11 elongate, conical. Last 3 segments more massive than others and form loose club (
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<figureCitation id="178F8D826A68FF86FB2836F9EF3FFE63" box="[1266,1325,395,417]" captionStart="Figs 1–4" captionStartId="1.[241,278,1974,1993]" captionTargetBox="[192,1393,1029,1946]" captionTargetId="figure-534@1.[189,1399,1023,1948]" captionTargetPageId="1" captionText="Figs 1–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n. Рис. 1–4. Carpelimus (Trogophloeus) spp., голотипы, самцы, сверху: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n." figureDoi="http://doi.org/10.5281/zenodo.10919902" httpUri="https://zenodo.org/record/10919902/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
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).
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</paragraph>
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<paragraph id="8F0B91076A68FF86FC8836D4EE98FD05" blockId="2.[812,1399,315,817]" pageId="2" pageNumber="32">
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Pronotum widest about 2/3 its length from base, then narrowed. Lateral margin with barely noticeable notch at base, then smoothly rounded (
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<figureCitation id="178F8D826A68FF86FBD536A9EE5AFE33" box="[1039,1096,475,497]" captionStart="Figs 1–4" captionStartId="1.[241,278,1974,1993]" captionTargetBox="[192,1393,1029,1946]" captionTargetId="figure-534@1.[189,1399,1023,1948]" captionTargetPageId="1" captionText="Figs 1–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n. Рис. 1–4. Carpelimus (Trogophloeus) spp., голотипы, самцы, сверху: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n." figureDoi="http://doi.org/10.5281/zenodo.10919902" httpUri="https://zenodo.org/record/10919902/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
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). Ratio of pronotum length to its maximum width about 19:26. Surface of pronotum with delicate, fine and dense punctation; punctation similar to that on head. Pronotal disc with 2 pairs of prominent, symmetrical depressions and 1 unpaired oval depression along midline near apex. Base of pronotal disc with narrow, crescent-shaped depressions separated by medial ridge. Central part of disc with rather deep, oval depressions fused across midline to single butterfly-shaped depression (
|
||
<figureCitation id="178F8D826A68FF86FB9835C3EE6FFD05" box="[1090,1149,689,711]" captionStart="Figs 1–4" captionStartId="1.[241,278,1974,1993]" captionTargetBox="[192,1393,1029,1946]" captionTargetId="figure-534@1.[189,1399,1023,1948]" captionTargetPageId="1" captionText="Figs 1–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n. Рис. 1–4. Carpelimus (Trogophloeus) spp., голотипы, самцы, сверху: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n." figureDoi="http://doi.org/10.5281/zenodo.10919902" httpUri="https://zenodo.org/record/10919902/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8F0B91076A68FF86FC8835B9EF65FCF3" blockId="2.[812,1399,315,817]" pageId="2" pageNumber="32">
|
||
Ratio of length of elytra to their combined width about 30:34. Scutellum with shallow, round depressions (
|
||
<figureCitation id="178F8D826A68FF86FAF63594EF78FD3E" box="[1324,1386,742,764]" captionStart="Figs 1–4" captionStartId="1.[241,278,1974,1993]" captionTargetBox="[192,1393,1029,1946]" captionTargetId="figure-534@1.[189,1399,1023,1948]" captionTargetPageId="1" captionText="Figs 1–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n. Рис. 1–4. Carpelimus (Trogophloeus) spp., голотипы, самцы, сверху: 1 — C. notumus, sp.n.; 2 — C. vilisus, sp.n.; 3 — C. plenus, sp.n.; 4 — C. notatus, sp.n." figureDoi="http://doi.org/10.5281/zenodo.10919902" httpUri="https://zenodo.org/record/10919902/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
|
||
). Elytra with rather delicate, fine and dense punctation. Diameter of punctures about 1.5 times as small as eye facet.
|
||
</paragraph>
|
||
<paragraph id="8F0B91076A68FF86FC493017EF01F8BD" blockId="2.[915,1299,1893,1919]" box="[915,1299,1893,1919]" pageId="2" pageNumber="32">
|
||
<emphasis id="BDC04D156A68FF86FC493017EF01F8BD" bold="true" box="[915,1299,1893,1919]" pageId="2" pageNumber="32">12 13</emphasis>
|
||
</paragraph>
|
||
<caption id="DBCBC18F6A68FF86FF3830E4EB63F7E7" ID-DOI="http://doi.org/10.5281/zenodo.10919904" ID-Zenodo-Dep="10919904" httpUri="https://zenodo.org/record/10919904/files/figure.png" pageId="2" pageNumber="32" startId="2.[226,263,1942,1961]" targetBox="[193,1399,842,1891]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8F0B91076A68FF86FF3830E4E842F81B" blockId="2.[189,1399,1941,2085]" pageId="2" pageNumber="32">
|
||
Figs 5–14. Genitalia of
|
||
<taxonomicName id="48B4EA846A68FF86FE6A30E4E8B2F86B" authorityName="Mannerheim" authorityYear="1830" box="[432,672,1942,1961]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="subGenus" subGenus="Trogophloeus">
|
||
<emphasis id="BDC04D156A68FF86FE6A30E4E806F86B" box="[432,532,1942,1961]" italics="true" pageId="2" pageNumber="32">Carpelimus</emphasis>
|
||
(
|
||
<emphasis id="BDC04D156A68FF86FDF830E4E88BF86B" box="[546,665,1942,1961]" italics="true" pageId="2" pageNumber="32">Trogophloeus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.: 5–7 —
|
||
<taxonomicName id="48B4EA846A68FF86FCCD30E4E968F86A" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[791,890,1942,1961]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="notumus" status="sp.n.">
|
||
<emphasis id="BDC04D156A68FF86FCCD30E4E937F86B" box="[791,805,1942,1961]" italics="true" pageId="2" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A68FF86FCEB30E4E968F86A" box="[817,890,1942,1960]" italics="true" pageId="2" pageNumber="32">notumus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BDC04D156A68FF86FC5F30E4E9A2F86B" bold="true" box="[901,944,1942,1961]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A6F3F06E6A68FF86FC5F30E4E9A2F86B" box="[901,944,1942,1961]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 8–9 —
|
||
<taxonomicName id="48B4EA846A68FF86FC2730E4EE5CF86B" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[1021,1102,1942,1961]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="vilisus" status="sp.n.">
|
||
<emphasis id="BDC04D156A68FF86FC2730E4EE19F86B" box="[1021,1035,1942,1961]" italics="true" pageId="2" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A68FF86FBCD30E4EE5CF86B" box="[1047,1102,1942,1961]" italics="true" pageId="2" pageNumber="32">vilisus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BDC04D156A68FF86FB8030E4EE96F86B" bold="true" box="[1114,1156,1942,1961]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A6F3F06E6A68FF86FB8030E4EE96F86B" box="[1114,1156,1942,1961]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 10–11 —
|
||
<taxonomicName id="48B4EA846A68FF86FB3230E4EF2BF86B" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[1256,1337,1942,1961]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="plenus" status="sp.n.">
|
||
<emphasis id="BDC04D156A68FF86FB3230E4EEE4F86B" box="[1256,1270,1942,1961]" italics="true" pageId="2" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A68FF86FADB30E4EF2BF86B" box="[1281,1337,1942,1961]" italics="true" pageId="2" pageNumber="32">plenus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BDC04D156A68FF86FA9F30E4EF62F86B" bold="true" box="[1349,1392,1942,1961]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A6F3F06E6A68FF86FA9F30E4EF62F86B" box="[1349,1392,1942,1961]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 12–14 —
|
||
<taxonomicName id="48B4EA846A68FF86FECF30DCEB7CF802" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[277,366,1966,1985]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="notatus" status="sp.n.">
|
||
<emphasis id="BDC04D156A68FF86FECF30DCEB31F803" box="[277,291,1966,1985]" italics="true" pageId="2" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A68FF86FEF530DCEB7CF802" box="[303,366,1966,1984]" italics="true" pageId="2" pageNumber="32">notatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BDC04D156A68FF86FEA130DCEBBCF803" bold="true" box="[379,430,1966,1985]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A6F3F06E6A68FF86FEA130DCEBBAF803" box="[379,424,1966,1985]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
|
||
;
|
||
</emphasis>
|
||
5, 8, 10, 12 — aedeagus, ventral view (holotypes); 6, 9, 11, 13 — aedeagus, lateral view (holotypes); 7, 14 – spermatheca (paratypes). Scale bar: 0.25 mm.
|
||
</paragraph>
|
||
<paragraph id="8F0B91076A68FF86FF383090EB63F7E7" blockId="2.[189,1399,1941,2085]" pageId="2" pageNumber="32">
|
||
Рис. 5–14. Гениталии
|
||
<taxonomicName id="48B4EA846A68FF86FE7D3093E886F836" authorityName="Mannerheim" authorityYear="1830" box="[423,660,2017,2036]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="subGenus" subGenus="Trogophloeus">
|
||
<emphasis id="BDC04D156A68FF86FE7D3093E81BF836" box="[423,521,2017,2036]" italics="true" pageId="2" pageNumber="32">Carpelimus</emphasis>
|
||
(
|
||
<emphasis id="BDC04D156A68FF86FDCC3093E89FF836" box="[534,653,2017,2036]" italics="true" pageId="2" pageNumber="32">Trogophloeus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.: 5–7 —
|
||
<taxonomicName id="48B4EA846A68FF86FCDD3090E974F836" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[775,870,2018,2037]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="notumus" status="sp.n.">
|
||
<emphasis id="BDC04D156A68FF86FCDD3090E907F837" box="[775,789,2018,2037]" italics="true" pageId="2" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A68FF86FCC53090E974F836" box="[799,870,2018,2036]" italics="true" pageId="2" pageNumber="32">notumus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BDC04D156A68FF86FCAB3093E989F836" bold="true" box="[881,923,2017,2036]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A6F3F06E6A68FF86FCAB3093E989F836" box="[881,923,2017,2036]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 8–9 —
|
||
<taxonomicName id="48B4EA846A68FF86FC3F3090EE26F836" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[997,1076,2017,2037]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="vilisus" status="sp.n.">
|
||
<emphasis id="BDC04D156A68FF86FC3F3090E9E1F837" box="[997,1011,2018,2037]" italics="true" pageId="2" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A68FF86FC243093EE26F836" box="[1022,1076,2017,2036]" italics="true" pageId="2" pageNumber="32">vilisus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BDC04D156A68FF86FBE43093EE7AF836" bold="true" box="[1086,1128,2017,2036]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A6F3F06E6A68FF86FBE43093EE7AF836" box="[1086,1128,2017,2036]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 10–11 —
|
||
<taxonomicName id="48B4EA846A68FF86FB1D3090EF05F836" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[1223,1303,2017,2037]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="plenus" status="sp.n.">
|
||
<emphasis id="BDC04D156A68FF86FB1D3090EEC7F837" box="[1223,1237,2018,2037]" italics="true" pageId="2" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A68FF86FB3A3093EF05F836" box="[1248,1303,2017,2036]" italics="true" pageId="2" pageNumber="32">plenus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BDC04D156A68FF86FAFB3093EF59F836" bold="true" box="[1313,1355,2017,2036]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A6F3F06E6A68FF86FAFB3093EF59F836" box="[1313,1355,2017,2036]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 12– 14 —
|
||
<taxonomicName id="48B4EA846A68FF86FF353088EB57F7CE" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[239,325,2042,2061]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="notatus" status="sp.n.">
|
||
<emphasis id="BDC04D156A68FF86FF353088EAEFF7CF" box="[239,253,2042,2061]" italics="true" pageId="2" pageNumber="32">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A68FF86FEDD3088EB57F7CE" box="[263,325,2042,2060]" italics="true" pageId="2" pageNumber="32">notatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="BDC04D156A68FF86FE94308BEB92F7CE" bold="true" box="[334,384,2041,2060]" pageId="2" pageNumber="32">
|
||
<taxonomicNameLabel id="A6F3F06E6A68FF86FE94308BEB69F7CE" box="[334,379,2041,2060]" pageId="2" pageNumber="32" rank="species">sp.n.</taxonomicNameLabel>
|
||
;
|
||
</emphasis>
|
||
5, 8, 10, 12 — Эдеагус, сниЗу (голотипы); 6, 9, 11, 13 — Эдеагус, сбоку (голотипы); 7, 14 — сперматека (паратипы). МасШтаб: 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8F0B91076A69FF87FF673649E863FEAE" blockId="3.[189,775,315,791]" pageId="3" pageNumber="33">Distances between punctures slightly smaller than their diameter, interspaces smooth, slightly shining.</paragraph>
|
||
<paragraph id="8F0B91076A69FF87FF383603E830FE45" blockId="3.[189,775,315,791]" box="[226,546,369,391]" pageId="3" pageNumber="33">Abdomen delicately shagreened.</paragraph>
|
||
<paragraph id="8F0B91076A69FF87FF3836F9E8A6FE63" blockId="3.[189,775,315,791]" box="[226,692,395,417]" pageId="3" pageNumber="33">
|
||
Aedeagus of characteristic structure (
|
||
<figureCitation id="178F8D826A69FF87FD9536F9E8BAFE63" box="[591,680,395,417]" captionStart="Figs 5–14" captionStartId="2.[226,263,1942,1961]" captionTargetBox="[193,1399,842,1891]" captionTargetId="figure-475@2.[189,1399,839,1891]" captionTargetPageId="2" captionText="Figs 5–14. Genitalia of Carpelimus (Trogophloeus) spp.: 5–7 — C. notumus, sp.n.; 8–9 — C. vilisus, sp.n.; 10–11 — C. plenus, sp.n.; 12–14 — C. notatus, sp.n.; 5, 8, 10, 12 — aedeagus, ventral view (holotypes); 6, 9, 11, 13 — aedeagus, lateral view (holotypes); 7, 14 – spermatheca (paratypes). Scale bar: 0.25 mm. Рис. 5–14. Гениталии Carpelimus (Trogophloeus) spp.: 5–7 — C. notumus, sp.n.; 8–9 — C. vilisus, sp.n.; 10–11 — C. plenus, sp.n.; 12– 14 — C. notatus, sp.n.; 5, 8, 10, 12 — Эдеагус, сниЗу (голотипы); 6, 9, 11, 13 — Эдеагус, сбоку (голотипы); 7, 14 — сперматека (паратипы). МасШтаб: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10919904" httpUri="https://zenodo.org/record/10919904/files/figure.png" pageId="3" pageNumber="33">Figs 8–9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8F0B91076A69FF87FF3836D4EB83FE33" blockId="3.[189,775,315,791]" pageId="3" pageNumber="33">
|
||
<emphasis id="BDC04D156A69FF87FF3836D4EB3FFE79" bold="true" box="[226,301,422,443]" pageId="3" pageNumber="33">Female</emphasis>
|
||
. Sexual dimorphism absent, female morphologically similar to male. Spermatheca failed to isolate and its structure is unknown.
|
||
</paragraph>
|
||
<paragraph id="8F0B91076A69FF87FF383684E894FD6E" blockId="3.[189,775,315,791]" pageId="3" pageNumber="33">
|
||
COMPARATIVE REMARKS. The new species is similar and closely related to
|
||
<taxonomicName id="48B4EA846A69FF87FE733563E804FDE4" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[425,534,529,550]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="notumus" status="sp.n.">
|
||
<emphasis id="BDC04D156A69FF87FE733563EBABFDE4" box="[425,441,529,550]" italics="true" pageId="3" pageNumber="33">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A69FF87FE1E3563E804FDE4" box="[452,534,529,550]" italics="true" pageId="3" pageNumber="33">notumus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BDC04D156A69FF87FDC13563E859FDE4" bold="true" box="[539,587,529,550]" pageId="3" pageNumber="33">
|
||
<taxonomicNameLabel id="A6F3F06E6A69FF87FDC13563E859FDE4" box="[539,587,529,550]" pageId="3" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="48B4EA846A69FF87FDAD3563E8C3FDE4" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" box="[631,721,529,550]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="plenus" status="sp.n.">
|
||
<emphasis id="BDC04D156A69FF87FDAD3563E895FDE4" box="[631,647,529,550]" italics="true" pageId="3" pageNumber="33">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A69FF87FD483563E8C3FDE4" box="[658,721,529,550]" italics="true" pageId="3" pageNumber="33">plenus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BDC04D156A69FF87FD0C3563E915FDE4" bold="true" box="[726,775,529,550]" pageId="3" pageNumber="33">
|
||
<taxonomicNameLabel id="A6F3F06E6A69FF87FD0C3563E915FDE4" box="[726,775,529,550]" pageId="3" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see descriptions above and below). It differs from
|
||
<taxonomicName id="48B4EA846A69FF87FD6C355EEAF6FD9E" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="notumus" status="sp.n.">
|
||
<emphasis id="BDC04D156A69FF87FD6C355EE8D4FD83" box="[694,710,556,577]" italics="true" pageId="3" pageNumber="33">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A69FF87FD0E355EEAF6FD9E" italics="true" pageId="3" pageNumber="33">notumus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BDC04D156A69FF87FF323534EB0BFD99" bold="true" box="[232,281,582,603]" pageId="3" pageNumber="33">
|
||
<taxonomicNameLabel id="A6F3F06E6A69FF87FF323534EB0BFD99" box="[232,281,582,603]" pageId="3" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having slightly more prominent eyes and from
|
||
<taxonomicName id="48B4EA846A69FF87FD2A3534EAEEFDB4" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="plenus" status="sp.n.">
|
||
<emphasis id="BDC04D156A69FF87FD2A3534E912FD99" box="[752,768,582,603]" italics="true" pageId="3" pageNumber="33">C</emphasis>
|
||
.
|
||
<emphasis id="BDC04D156A69FF87FF673513EAEEFDB4" box="[189,252,609,630]" italics="true" pageId="3" pageNumber="33">plenus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BDC04D156A69FF87FED93513EB26FDB4" bold="true" box="[259,308,609,630]" pageId="3" pageNumber="33">
|
||
<taxonomicNameLabel id="A6F3F06E6A69FF87FED93513EB26FDB4" box="[259,308,609,630]" pageId="3" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a significantly smaller body size and smaller punctation on the body surface. It differs markedly in the structure of the aedeagus (
|
||
<figureCitation id="178F8D826A69FF87FE3935E4E868FD6E" box="[483,634,662,684]" captionStart="Figs 5–14" captionStartId="2.[226,263,1942,1961]" captionTargetBox="[193,1399,842,1891]" captionTargetId="figure-475@2.[189,1399,839,1891]" captionTargetPageId="2" captionText="Figs 5–14. Genitalia of Carpelimus (Trogophloeus) spp.: 5–7 — C. notumus, sp.n.; 8–9 — C. vilisus, sp.n.; 10–11 — C. plenus, sp.n.; 12–14 — C. notatus, sp.n.; 5, 8, 10, 12 — aedeagus, ventral view (holotypes); 6, 9, 11, 13 — aedeagus, lateral view (holotypes); 7, 14 – spermatheca (paratypes). Scale bar: 0.25 mm. Рис. 5–14. Гениталии Carpelimus (Trogophloeus) spp.: 5–7 — C. notumus, sp.n.; 8–9 — C. vilisus, sp.n.; 10–11 — C. plenus, sp.n.; 12– 14 — C. notatus, sp.n.; 5, 8, 10, 12 — Эдеагус, сниЗу (голотипы); 6, 9, 11, 13 — Эдеагус, сбоку (голотипы); 7, 14 — сперматека (паратипы). МасШтаб: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10919904" httpUri="https://zenodo.org/record/10919904/files/figure.png" pageId="3" pageNumber="33">Figs 5–6, 8–11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C7AEC28C6A69FF87FF3835C3EBE5FD05" box="[226,503,689,711]" pageId="3" pageNumber="33" type="distribution">
|
||
<paragraph id="8F0B91076A69FF87FF3835C3EBE5FD05" blockId="3.[189,775,315,791]" box="[226,503,689,711]" pageId="3" pageNumber="33">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F7A3D1976A69FF87FE4035C3EBE1FD05" box="[410,499,689,711]" name="Thailand" pageId="3" pageNumber="33">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C7AEC28C6A69FF87FF3835B9EB89FCD4" pageId="3" pageNumber="33" type="etymology">
|
||
<paragraph id="8F0B91076A69FF87FF3835B9EB89FCD4" blockId="3.[189,775,315,791]" pageId="3" pageNumber="33">
|
||
ETYMOLOGY. From Latin “vilis” (worthless, insignificant) referring to a small difference in its external appearance from
|
||
<taxonomicName id="48B4EA846A69FF87FEFF3473EB87FCD4" authorityName="Gildenkov" authorityYear="2019" box="[293,405,769,790]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="notumus">
|
||
<emphasis id="BDC04D156A69FF87FEFF3473EB87FCD4" box="[293,405,769,790]" italics="true" pageId="3" pageNumber="33">C. notumus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |