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<document id="E18C11FFCA6C238D9108D95C026848C4" ID-DOI="10.3897/zse.100.130929" ID-ZooBank="4D6C62E4-224B-4AB5-A251-AA7ABEE66D74" ID-publisher-id="130929" URI-arpha="62A0369D-D7AA-53B4-8EA3-723D47A8742A" URI-zoobank="http://zoobank.org/4D6C62E4-224B-4AB5-A251-AA7ABEE66D74" article-type="research-article" checkinTime="1728062842149" checkinUser="pensoft" docAuthor="Jeratthitikul, Ekgachai, Sutcharit, Chirasak &amp; Prasankok, Pongpun" docDate="2024" docId="C0910C0350AE55BFB0CF0786B4F04852" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1333-1345" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/130929/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Isannaia Jeratthitikul, Sutcharit &amp; Prasankok, 2024, gen. nov." docType="treatment" docUuid="ADD0E6C8-E5BD-4E05-828B-FC89CA662F02" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="1345" masterDocId="62A0369DD7AA53B48EA3723D47A8742A" masterDocTitle="Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin" masterLastPageNumber="1345" masterPageNumber="1333" pageNumber="1333" updateTime="1728063243823" updateUser="ExternalLinkService">
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<mods:title id="18A8BCA7238EF575A7DA73DDEE0FC173">Two new genera and three new species of exceptionally rare and endemic freshwater mussels (Bivalvia, Unionidae) from the Mekong Basin</mods:title>
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<mods:namePart id="C00A033353749B6DD88D1E8D79E03B99">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="021008F9719A311FD02B7A236994C8D2">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="F4F115D45FCAD7D7395A4558F66AFD6B">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="A930B13528116AABE697F4F5C824E33D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="8E64EB97C8FC694C2235ECC7D308B637">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="EECF17F280C2D3C42720A53E201D2F1F">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:date id="65BBE8EC3AAD66CA7D3C07D20B1D1A48">2024</mods:date>
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<treatment id="C0910C0350AE55BFB0CF0786B4F04852" ID-DOI="http://doi.org/10.5281/zenodo.13891549" ID-Zenodo-Dep="13891549" ID-arpha="C0910C03-50AE-55BF-B0CF-0786B4F04852" ID-zoobank="https://zoobank.org/ADD0E6C8-E5BD-4E05-828B-FC89CA662F02" LSID="urn:lsid:zoobank.org:act:ADD0E6C8-E5BD-4E05-828B-FC89CA662F02" LSID-ZBK="urn:lsid:zoobank.org:act:ADD0E6C8-E5BD-4E05-828B-FC89CA662F02" httpUri="http://treatment.plazi.org/id/C0910C0350AE55BFB0CF0786B4F04852">
<subSubSection id="2B29A41C709E339A8D3C18C591B4F9A6" type="nomenclature">
<paragraph id="1C840AD57F1A9DC79EBD3F5C0649BBAE">
<heading id="FAAE197395FBA6415A00E5920C6AB77E" reason="label">Genus</heading>
<taxonomicName id="923C051A16CEBD79E327F350D37BE593" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="51CC5850DECB5196BEB080C787F2E550" italics="true">Isannaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33D2F26601E1EA72015EBE77A9E936CA" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFCBG" type="type species">
<paragraph id="81175635211C7C197B9B7DF66820E9B5">
<heading id="52FD04D19E8B6DD8E4A093EF3DFB8181" reason="title">
<typeStatus id="471EE57340C18091EFC9A6812ADB9281">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="12508E6298957498ED1941758F294892">
<taxonomicName id="63CFC620E481E36B3AA39C882D00CE15" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="fortunata" status="gen. et sp. nov.">
<emphasis id="779E143F22D6CA29B5ECE6F320E9303D" italics="true">Isannaia fortunata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="12C83BA1D141941D7DD626C41900192B" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWCBG" type="Species included">
<paragraph id="6E2EF7D163682D33EBB7BE4AB437E3AC">
<heading id="A8E14BB6D9E1F59D9002A352ED6ED2EF" reason="title">Species included.</heading>
</paragraph>
<paragraph id="A3FB9A7E0E4A5D09D1B02C8761664EE6">
The new genus is currently composed of two species, the
<typeStatus id="4C21197835D8BC02B7E543AA5E3EC8E9">type</typeStatus>
species and
<taxonomicName id="CE969F55B1948D23A2F7D9D38075B36D" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="occultata" status="sp. nov.">
<emphasis id="D1A437D026D0D905E40F4FF53F974D06" italics="true">I. occultata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="888E7F6C0C4B0D5F4C8BD6CAE4EC8A85" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHDBG" type="diagnosis">
<paragraph id="E19AC779C6DA330F0753DCC0FA76F623">
<heading id="3599EA1F412550829D490973ED8D1E65" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="1CAD0916A32CBFC319DF7A54657CEE40">
The new genus is distinguished from other genera in the
<taxonomicName id="0AF8F89D9FD97DD610189EEEC630F6EF" authorityName="Frierson" authorityYear="1927" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="tribe" tribe="Pseudodontini">Pseudodontini</taxonomicName>
by its thin and moderately inflated shell, and with rhomboidal to ovate outline (Fig.
<figureCitation id="EA8985393653CBBF1BCB24D71CF66832" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shells of Isannaia gen. nov., showing the inner side of the right valve, the outer side of the left valve, and the dorsal view of both valves. A. Holotype (MUMNH - UNI 1950) and B. Paratype (MUMNH - UNI 1952) of Isannaia fortunata sp. nov.; C. Holotype (MUMNH - UNI 2895) and D. Paratype (MUMNH - UNI 2897) of Isannaia occultata sp. nov. Scale bars: 10 mm." figureDoi="10.3897/zse.100.130929.figure4" httpUri="https://binary.pensoft.net/fig/1148329">4</figureCitation>
). The overall shape is somewhat elongated but relatively shorter when compared to those of
<taxonomicName id="E63C1042A819F842A6F65C92FEEA3108" authorityName="Simpson" authorityYear="1900" class="Bivalvia" family="Unionidae" genus="Bineurus" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
<emphasis id="0B7CCD7D67A6408B148C661CD554DAA2" italics="true">Bineurus</emphasis>
</taxonomicName>
,
<taxonomicName id="DD9A1C63017DACE65DE311976A403B34" authorityName="Simpson" authorityYear="1900" class="Bivalvia" family="Unionidae" genus="Pilsbryoconcha" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
<emphasis id="5B2DBFA586E10AD0ED83C5240A9D7015" italics="true">Pilsbryoconcha</emphasis>
</taxonomicName>
or
<taxonomicName id="905A6F28C96E3AC4E22C92545A149A52" class="Bivalvia" family="Unionidae" genus="Namkongnaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
<emphasis id="A975D5ABE5022D7669F95C10351E065C" italics="true">Namkongnaia</emphasis>
</taxonomicName>
. The anterior margin is rounded, shouldered, and the posterior margin is truncated to rounded. Its pseudocardinal teeth are rather small, one on each valve, hill-like or triangulate shape, and less developed on the left valve. Its umbo is slightly elevated and the umbo cavity moderately deep. It is also represented as a distinct clade in multi-locus phylogenetic analyses (Fig.
<figureCitation id="62E40868269A3D84C83296BD2678676A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood (ML) tree of freshwater mussels in subfamily Gonideinae based on 1,935 - bp concatenated alignment dataset of COI and 16 S rRNA mitochondrial genes, and 28 S rRNA nuclear gene. The numbers on nodes represent the bootstrap support (BS) from the ML analysis and the bipartition posterior probability (bpp) from the BI analysis, and are shown as ML / BI. Scale bar indicates the branch length." figureDoi="10.3897/zse.100.130929.figure1" httpUri="https://binary.pensoft.net/fig/1148326">1</figureCitation>
).
</paragraph>
<caption id="DCDBE545EB72FAD0F87084ACC7EC70DD" ID-DOI="10.3897/zse.100.130929.figure4" ID-arpha="B9A19AC2-518B-580E-A04F-10591B6E051D" httpUri="https://binary.pensoft.net/fig/1148329" startId="F4">
<paragraph id="091EA6A86033B03E00761A1549EF7451">
<label id="FB2C043E0251173423370B910BE4B363">Figure 4.</label>
</paragraph>
<paragraph id="8ED90E4160508865869DA9509C449603">
Shells of
<taxonomicName id="8BFE7D60565FDE0008243F493A4F6DFB" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="CC653F4A3D717D476D5AA098784DB103" italics="true">Isannaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="008AD851B97136247980EF19CC2C83FE" rank="genus">gen. nov.</taxonomicNameLabel>
, showing the inner side of the right valve, the outer side of the left valve, and the dorsal view of both valves.
<emphasis id="3D1A5B4007D66ECEEBBECA69B8059E2E" bold="true">A.</emphasis>
Holotype (
<named-content id="5B3DDA946FF41888F93DE84272FDBFBF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
- UNI 1950) and
<emphasis id="F8655D969412611EE0836A65EAEAA0D9" bold="true">B.</emphasis>
Paratype (
<named-content id="7144CA3FC7C2D8C7D23E7B784CB3D8F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
- UNI 1952) of
<taxonomicName id="3A28F3441FC76D6D8D77267D23197A50" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="fortunata" status="sp. nov.">
<emphasis id="3D0F08965D88ACF62B0B99528602A14C" italics="true">Isannaia fortunata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97981AB5AEB04B4AE37ACB6CF0CEECD4" rank="species">sp. nov.</taxonomicNameLabel>
;
<emphasis id="695ED0FB31E24CF3957AE3F8E0507317" bold="true">C.</emphasis>
Holotype (
<named-content id="7282B9823D0486DE2146B66DCEC08736" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
- UNI 2895) and
<emphasis id="DFF1FC8D1523A8E608CC25D7342EF9B3" bold="true">D.</emphasis>
Paratype (
<named-content id="3AE99F3DABAB87D11926C7CA507FA0FD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahidol-university" xlink_title="Mahidol University Museum of Natural History, Department of Biology, Faculty of Science, Mahidol University, Bangkok, Thailand">MUMNH</named-content>
- UNI 2897) of
<taxonomicName id="2FD43FF62CF8137E915910D6DFB1EE8A" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="occultata" status="sp. nov.">
<emphasis id="F40877F193CAE7D923251144AE6F30C4" italics="true">Isannaia occultata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E90368596752ACDCAC09E95D2A323A0" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EVGBG" type="description">
<paragraph id="F174D06EB8D6AC984DE8E4F5A1A05367">
<heading id="87E1A4F7AA59AF1802099F4ED8FF2F37" reason="title">Description.</heading>
</paragraph>
<paragraph id="7B39BC3CB771E28C42BDF57DE17653DB">Shell medium-sized, rather thin, rhomboidal to ovate outline, elongate, very inequilateral, moderately inflated; anterior margin rounded, shouldered; posterior margin truncated to rounded. Umbo slightly elevated and usually eroded. Ligament very narrow. One shallow hill-like or triangulate pseudocardinal tooth on right valve; left valve with one less developed hill-like to tubercule-like pseudocardinal tooth. End of hinge structure with V-shaped fossette. Anterior adductor muscle scar shallow to well-developed, ovate, contiguous with pedal retractor and anterior protractor muscle scars; posterior adductor muscle scar round, very shallow. Pallial line faint to well-marked, continuous. Umbo cavity moderately deep.</paragraph>
</subSubSection>
<subSubSection id="SECID0E1GBG" type="etymology">
<paragraph id="E898A9C13559316883F6C4E70CB5600D">
<heading id="02BAAD8ACF1EBF90D54D1B9C320F5C6E" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A27F6A00C41693907A398AF08EB663B5">
The generic name “
<emphasis id="6511D2CC1DAF2C2F19FF620F82EEF77D" italics="true">Isannaia</emphasis>
” is from the word “
<emphasis id="89783BD5D66DBA051D1659A8F0BFAB77" italics="true">Isan</emphasis>
”, the name of the region of northeastern
<collectingCountry id="634E731AC004F833DCF2F8E8EF18F019" name="Thailand">Thailand</collectingCountry>
, and Greek word “
<emphasis id="97F328ADA757490D5A625BFF5C0DC135" italics="true">naiad</emphasis>
” meaning freshwater mussel. The name of this genus thus means “ freshwater mussels from northeastern
<collectingCountry id="21EE5D1F2AFBCB5BE445A63187AD92BD" name="Thailand">Thailand</collectingCountry>
”. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQHBG" type="distribution">
<paragraph id="1EB9CB162E83212922D45D8ADBDD6197">
<heading id="A1BAAD912824AC704CAFDB6C007E9080" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A9591C2F6A6BA6AF96F91F740C968744">
Endemic to tributaries of the Mekong River in northeastern
<collectingCountry id="978EFF84D01E0F57533EBE4008F205C9" name="Thailand">Thailand</collectingCountry>
(Fig.
<figureCitation id="90ADA6410872511E1067DC0E55EBF358" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing type localities of the new species: circle Lannanaia kokensis sp. nov., triangle Isannaia fortunata sp. nov., and square Isannaia occultata sp. nov. Map was generated using QGIS v 3.24. 3 with the outline of Mekong basin from the Freshwater Ecoregions of the World (Abell et al. 2008), river and lake topology from the HydroSHEDS database (https://www.hydrosheds.org), and the map raster data from the NASA EARTHDATA (https://www.earthdata.nasa.gov/)." figureDoi="10.3897/zse.100.130929.figure3" httpUri="https://binary.pensoft.net/fig/1148328">3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1HBG" type="remarks">
<paragraph id="19080B9D0C7A1E48480B7B96ECBCFBA5">
<heading id="56AE1D1BE6F1E03C041C409485BBD35E" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="05A90026872ED7C8F1D5E20821ADFB1F">
Phylogenetic analyses have failed to recover a robustly supported position of
<taxonomicName id="876F6E2540F0A9D9B3B44BEAB2DF116D" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FEE4DBEB57931DAA7375D191424808D0" italics="true">Isannaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C80309BD92AF58172F996E8BC747E4DE" rank="genus">gen. nov.</taxonomicNameLabel>
among other
<taxonomicName id="40667A36925D89B2C7B100A2A0CC07DA" authorityName="Frierson" authorityYear="1927" class="Bivalvia" family="Unionidae" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="subTribe" subTribe="Pseudodontina">Pseudodontina</taxonomicName>
genera. However, in terms of genetic distance, it is nearest to
<taxonomicName id="9593A69C5CA9527F8CED76A81200AB62" class="Bivalvia" family="Unionidae" genus="Namkongnaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
<emphasis id="651DCA611190BB6291CFDA225AFC3F79" italics="true">Namkongnaia</emphasis>
</taxonomicName>
, with a separation of 11.77 % p-distance of the
<abbrev id="ABBRID0ETIBG" xlink_title="mitochondrial cytochrome c oxidase subunit-I gene">COI</abbrev>
gene (Table
<tableCitation id="E9D2751B13FB25DE93C049A5EB68F6A0" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Mean genetic distances (uncorrected p-distance: % ± SD) based on 660 bp COI gene fragment sequences among genera of the tribe Pseudodontini (below diagonal), and between species within each genus (in bold)." httpUri="http://table.plazi.org/id/B78670F787E884488518B3F8BC0B50EB" tableUuid="B78670F787E884488518B3F8BC0B50EB">1</tableCitation>
).
<taxonomicName id="77C4D9623BA851F99E2BF45DC5D40D2B" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="34C16887623F56E6554B7CF89567D490" italics="true">Isannaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BF665713C2AEA02A3DAD8B636E8EA37" rank="genus">gen. nov.</taxonomicNameLabel>
can be easily distinguished from
<taxonomicName id="5286F6661374496A1E715E909A1058A6" class="Bivalvia" family="Unionidae" genus="Namkongnaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
<emphasis id="E256CF61B8CC818C7F0D138684B8BADE" italics="true">Namkongnaia</emphasis>
</taxonomicName>
by having a much shorter shell (
<bibRefCitation id="820A40312AE77A864577632B40C45BBA" DOI="10.5852/ejt.2021.775.1553" author="Jeratthitikul E &amp; Sutcharit C &amp; Ngor PB &amp; Prasankok P" journalOrPublisher="European Journal of Taxonomy" pagination="119-142" refId="B21" refString="Jeratthitikul E, Sutcharit C, Ngor PB, Prasankok P (2021 b) Molecular phylogeny reveals a new genus of freshwater mussels from the Mekong River Basin (Bivalvia: Unionidae). European Journal of Taxonomy 775: 119142. https://doi.org/10.5852/ejt.2021.775.1553" title="Molecular phylogeny reveals a new genus of freshwater mussels from the Mekong River Basin (Bivalvia: Unionidae)." volume="775">Jeratthitikul et al. 2021 b</bibRefCitation>
).
</paragraph>
<paragraph id="AE19F05ED540BC0CA33533C3CA39446A">
In general,
<taxonomicName id="1685F78809B59D3C816DB278423F2006" authorityName="Jeratthitikul &amp; Sutcharit &amp; Prasankok" authorityYear="2024" class="Bivalvia" family="Unionidae" genus="Isannaia" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="34898A187662DCECA0D38E237501DE11" italics="true">Isannaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0A13F78EE5A1E22B3D04FCAC4106E4D" rank="genus">gen. nov.</taxonomicNameLabel>
bears conchological resemblance to certain species within the
<taxonomicName id="ADB47FEEF3C5FA0E538234AF31C349B6" class="Bivalvia" family="Unionidae" genus="Sundadontina" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="genus">
<emphasis id="B9112920395A223433BF1F1E6CF54224" italics="true">Sundadontina</emphasis>
</taxonomicName>
, such as
<taxonomicName id="63C1D2A86DA82D88DBCD106776F2AD16" authority="Bolotov et al., 2020" authorityName="Bolotov" authorityYear="2020" class="Bivalvia" family="Unionidae" genus="Sundadontina" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="brandti">
<emphasis id="FECE2D6DD4A580EE06FC1737B4962430" italics="true">S. brandti</emphasis>
<bibRefCitation id="4A0D6801582166A09CAD59A102C44B96" DOI="10.1038/s41598-020-63612-5" author="Bolotov" etAl="et al." firstAuthor="Bolotov" refId="B7" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Lopes-Lima M, Bogan AE, Lunn Z, Chan N, Win T, Aksenova OV, Tomilova AA, Tanmuangpak K, Tumpeesuwan S, Kondakov AV (2020) New freshwater mussel taxa discoveries clarify biogeographic division of Southeast Asia. Scientific Reports 10: 6616. https://doi.org/10.1038/s41598-020-63612-5" year="2020">Bolotov et al., 2020</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="501F68572977C4A34426C1E37459E608" authority="Konopleva et al., 2023" authorityName="Konopleva" authorityYear="2023" class="Bivalvia" family="Unionidae" genus="Sundadontina" kingdom="Animalia" order="Unionida" phylum="Mollusca" rank="species" species="plugpomenica">
<emphasis id="A32CC4BA05563BE2B862FA500316BE61" italics="true">S. plugpomenica</emphasis>
<bibRefCitation id="7C119504AEC8A491F8BEE23E9E32BA7F" DOI="10.3390/d15010010" author="Konopleva" etAl="et al." firstAuthor="Konopleva" refId="B28" refString="Konopleva ES, Lheknim V, Sriwoon R, Kondakov AV, Tomilova AA, Gofarov MY, Vikhrev IV, Bolotov IN (2023) Diversity and phylogenetics of freshwater mussels (Unionidae) from southern Thailand with the description of one new genus and five new species-group taxa. Diversity 15: 10. https://doi.org/10.3390/d15010010" year="2023">Konopleva et al., 2023</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="5BB588F82FB3DC9891CE576BC1345B00" DOI="10.1038/s41598-020-63612-5" author="Bolotov" etAl="et al." firstAuthor="Bolotov" refId="B7" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Lopes-Lima M, Bogan AE, Lunn Z, Chan N, Win T, Aksenova OV, Tomilova AA, Tanmuangpak K, Tumpeesuwan S, Kondakov AV (2020) New freshwater mussel taxa discoveries clarify biogeographic division of Southeast Asia. Scientific Reports 10: 6616. https://doi.org/10.1038/s41598-020-63612-5" year="2020">Bolotov et al. 2020</bibRefCitation>
;
<bibRefCitation id="AD9C4955B14320EECB68855591AEB095" DOI="10.3390/d15010010" author="Konopleva" etAl="et al." firstAuthor="Konopleva" refId="B28" refString="Konopleva ES, Lheknim V, Sriwoon R, Kondakov AV, Tomilova AA, Gofarov MY, Vikhrev IV, Bolotov IN (2023) Diversity and phylogenetics of freshwater mussels (Unionidae) from southern Thailand with the description of one new genus and five new species-group taxa. Diversity 15: 10. https://doi.org/10.3390/d15010010" year="2023">Konopleva et al. 2023</bibRefCitation>
). Nevertheless, its thin shell with less developed pseudocardinal teeth set it apart from the latter genus, which typically features a thicker and sturdier shell with stout and tubercular-like pseudocardinal teeth (
<bibRefCitation id="A0500D7C5C0C355796FFB5C63171A845" DOI="10.1038/s41598-020-63612-5" author="Bolotov" etAl="et al." firstAuthor="Bolotov" refId="B7" refString="Bolotov IN, Konopleva ES, Vikhrev IV, Gofarov MY, Lopes-Lima M, Bogan AE, Lunn Z, Chan N, Win T, Aksenova OV, Tomilova AA, Tanmuangpak K, Tumpeesuwan S, Kondakov AV (2020) New freshwater mussel taxa discoveries clarify biogeographic division of Southeast Asia. Scientific Reports 10: 6616. https://doi.org/10.1038/s41598-020-63612-5" year="2020">Bolotov et al. 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>