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<document id="302E4A0A6E983C67A7F6F03A82D2B2A4" ID-CLB-Dataset="3434" ID-DOI="10.1111/zoj.12482" ID-GBIF-Dataset="ede2ef87-5522-4e98-97e5-9b689b99ac73" ID-ISSN="0024-4082" ID-Zenodo-Dep="5710946" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619627631287" checkinUser="carolina" docAuthor="Jaafar, Zeehan &amp; Parenti, Lynne R." docDate="2017" docId="E91587EB4D316B318557FE0544B69B04" docLanguage="en" docName="ZoolJLinnSoc.180.195-215.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 180 (5)" docSource="http://dx.doi.org/10.1111/zoj.12482" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Oxuderces NEXIPINNIS (CANTOR 1849" docType="treatment" docVersion="8" lastPageNumber="210" masterDocId="152CFF934D3D6B3E8638FFCE40629E43" masterDocTitle="Systematics of the mudskipper genus Oxuderces Eydoux &amp; Souleyet 1848 (Teleostei: Gobiidae: Oxudercinae) with resurrection from synonymy of O. nexipinnis (Cantor 1849)" masterLastPageNumber="215" masterPageNumber="195" pageNumber="207" updateTime="1698937812356" updateUser="ExternalLinkService">
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<mods:title id="C647695D7C774CB3D8DD41CB1A155E14">Systematics of the mudskipper genus Oxuderces Eydoux &amp; Souleyet 1848 (Teleostei: Gobiidae: Oxudercinae) with resurrection from synonymy of O. nexipinnis (Cantor 1849)</mods:title>
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<mods:namePart id="7ADEE55BD5A60E2B4C614B61BF5C30EC">Parenti, Lynne R.</mods:namePart>
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<mods:title id="A1F6E1CF1044BA1CCE340C627D404806">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="C4C55243D78328AF475ACE90BA1BFDE6">2017</mods:date>
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<treatment id="E91587EB4D316B318557FE0544B69B04" ID-DOI="http://doi.org/10.5281/zenodo.5710965" ID-GBIF-Taxon="190222511" ID-Zenodo-Dep="5710965" LSID="urn:lsid:plazi:treatment:E91587EB4D316B318557FE0544B69B04" httpUri="http://treatment.plazi.org/id/E91587EB4D316B318557FE0544B69B04" lastPageId="15" lastPageNumber="210" pageId="12" pageNumber="207">
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<taxonomicName id="A6BC4D7E4D316B328557FE05452A9FA3" ID-CoL="6B7Q" ID-ENA="1109016" authority="NEXIPINNIS (CANTOR, 1849)" authorityName="NEXIPINNIS (CANTOR" authorityYear="1849" box="[879,1352,454,481]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="207" phylum="Chordata" rank="genus">
<emphasis id="53C8EAEF4D316B328557FE0544E39F9D" box="[879,1153,459,480]" italics="true" pageId="12" pageNumber="207">OXUDERCES NEXIPINNIS</emphasis>
(
<bibRefCitation id="052D4B0C4D316B3282ACFE06455D9FA3" author="Cantor T" box="[1172,1343,454,481]" pageId="12" pageNumber="207" pagination="983 - 1443" refId="ref12023" refString="Cantor T. 1849. Catalog of Malayan fishes. Journal of the Asiatic Society of Bengal 18: 983 - 1443." type="journal article" year="1849">CANTOR, 1849</bibRefCitation>
)
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<paragraph id="610336FD4D316B32850FFE2945E29FBC" blockId="12.[823,1408,454,541]" box="[823,1408,485,512]" pageId="12" pageNumber="207">
<figureCitation id="F9872A784D316B32850FFE2943D79FBC" box="[823,949,485,511]" captionStart="Figure 2" captionStartId="1.[827,906,1821,1840]" captionTargetBox="[834,1435,1682,1787]" captionTargetId="figure-753@1.[834,1435,1682,1787]" captionTargetPageId="1" captionText="Figure 2. Oxuderces nexipinnis (Cantor, 1849), ZRC 53919, 53.2 mm SL, West Bengal, India. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5710952" httpUri="https://zenodo.org/record/5710952/files/figure.png" pageId="12" pageNumber="207">FIGURES 2</figureCitation>
,
<figureCitation id="F9872A784D316B3285FBFE2B43879FBC" box="[963,997,485,511]" captionStart="Figure 3" captionStartId="4.[146,225,1790,1809]" captionTargetId="figure-29@3.[174,1437,198,1838]" captionTargetPageId="3" captionText="Figure 3. Dorsal view of the head including interorbital region of (A) Oxuderces dentatus (ZRC 53939) and (B) Oxuderces nexipinnis (ZRC 53919). Arrow points to interorbital trough or groove. A sensory pore C sits in the anterior portion of the trough. C, Apocryptodon wirzi (AMS I23993.007), there is a single sensory pore D, and no trough; D, Apocryptodon glyphisodon (USNM 99874), there is a paired sensory pore E and no trough. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5729842" httpUri="https://zenodo.org/record/5729842/files/figure.png" pageId="12" pageNumber="207">3B</figureCitation>
,
<figureCitation id="F9872A784D316B3285CFFE2B447A9FBC" box="[1015,1048,485,511]" captionStart="Figure 4" captionStartId="6.[146,225,1791,1810]" captionTargetId="figure-29@5.[174,1440,198,1876]" captionTargetPageId="5" captionText="Figure 4. Histological sections through the head of (A) Oxuderces nexipinnis (USNM 427099) and (B) Apocryptodon wirzi (USNM 432509). The paired adductor arcus palatini lie on either side of the parasphenoid and the intermandibularis spans the lower jaw and connects the dentaries. Bone is bright red, cartilage light blue, muscle and connective tissue dark blue to purple. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5729846" httpUri="https://zenodo.org/record/5729846/files/figure.png" pageId="12" pageNumber="207">4A</figureCitation>
,
<figureCitation id="F9872A784D316B328212FE2B44299FBC" box="[1066,1099,485,511]" captionStart="Figure 5" captionStartId="7.[829,908,1716,1735]" captionTargetBox="[173,774,202,1847]" captionTargetId="figure-358@7.[173,774,199,1848]" captionTargetPageId="7" captionText="Figure 5. Enlargement of the histological sections of Figure 4 through the mid-dorsal region of (A) Oxuderces nexipinnis (USNM 427099) and (B) Apocryptodon wirzi (USNM 432509). Bone is bright red, cartilage light blue, muscle and connective tissue dark blue to purple." figureDoi="http://doi.org/10.5281/zenodo.5710954" httpUri="https://zenodo.org/record/5710954/files/figure.png" pageId="12" pageNumber="207">5A</figureCitation>
,
<figureCitation id="F9872A784D316B328265FE2B440D9FBC" box="[1117,1135,485,511]" captionStart="Figure 6" captionStartId="9.[166,245,730,749]" captionTargetBox="[173,774,198,697]" captionTargetId="figure-567@9.[173,774,198,697]" captionTargetPageId="9" captionText="Figure 6. Dermal invagination or pouch (arrow) at point of attachment of the pelvic-fin base in Oxuderces nexipinnis (ZRC 53919, 53.2 mm SL). Anterior is to the right." figureDoi="http://doi.org/10.5281/zenodo.5710958" httpUri="https://zenodo.org/record/5710958/files/figure.png" pageId="12" pageNumber="207">6</figureCitation>
,
<figureCitation id="F9872A784D316B328246FE2844FD9FBC" box="[1150,1183,486,512]" captionStart="Figure 7" captionStartId="9.[166,245,1762,1781]" captionTargetId="figure-35@10.[153,1388,312,1755]" captionTargetPageId="10" captionText="Figure 7. Line drawings of the jaw and jaw suspensorium of (A) Oxuderces nexipinnis (USNM 279359) and (B) Gnatholepis sp. (USNM 287166). Angle between metapterygoid–symplectic–quadrate strut and anguloarticular is acute in O. nexipinnis, in which the preopercle is relatively thin and subcrescent-shaped. In contrast, the angle between the strut and the anguloarticular is obtuse, and the preopercle is wider and broadly crescent-shaped in Gnatholepis sp. Bone is stippled, cartilage is represented by open circles." figureDoi="http://doi.org/10.5281/zenodo.5729848" httpUri="https://zenodo.org/record/5729848/files/figure.png" pageId="12" pageNumber="207">7A</figureCitation>
,
<figureCitation id="F9872A784D316B328289FE2B44B09FBC" box="[1201,1234,485,511]" captionStart="Figure 8" captionStartId="13.[829,908,1583,1602]" captionTargetBox="[173,774,345,1855]" captionTargetId="figure-420@13.[173,773,431,1848]" captionTargetPageId="13" captionText="Figure 8. Line drawings of the hyoid bar in (A) Oxuderces nexipinnis (USNM 279359), (B) Apocryptodon wirzi (USNM 287292) and (C) Gnatholepis sp. (USNM 287166). The anterior ceratohyal posterior to insertion of the fourth branchiostegal ray is elongate and notched (arrow) in O. nexipinnis, elongate and not notched (arrow) in A. wirzi, and not elongate and not notched in Gnatholepis sp. Branchiostegal rays 3–5 are distinctly thinner in A and B. Bone is stippled, cartilage is represented by open circles. Only the proximal portion of the five branchiostegal rays is drawn." figureDoi="http://doi.org/10.5281/zenodo.5710963" httpUri="https://zenodo.org/record/5710963/files/figure.png" pageId="12" pageNumber="207">8A</figureCitation>
,
<figureCitation id="F9872A784D316B3282DCFE2B45679FBC" box="[1252,1285,485,511]" captionStart="Figure 9" captionStartId="14.[146,225,1581,1600]" captionTargetBox="[156,752,291,1549]" captionTargetId="figure-949@14.[156,752,291,1461]" captionTargetPageId="14" captionText="Figure 9. Line drawings of several anterior vertebrae and associated elements of the first dorsal fin in (A) Oxuderces nexipinnis (USNM 279359) and (B) Gnatholepis sp. (USNM 287166). The neural spine of the fourth vertebra (arrow) is broad and spatulate and directed posteriorly in O. nexipinnis and relatively thin and pointed and directed dorsally in Gnatholepis. Bone is stippled, cartilage is represented by open circles." figureDoi="http://doi.org/10.5281/zenodo.5710967" httpUri="https://zenodo.org/record/5710967/files/figure.png" pageId="12" pageNumber="207">9A</figureCitation>
,
<figureCitation id="F9872A784D316B32832FFE2B452A9FBC" box="[1303,1352,485,511]" captionStart="Figure 10" captionStartId="15.[829,908,1600,1619]" captionTargetBox="[173,774,204,1740]" captionTargetId="figure-441@15.[176,766,275,1733]" captionTargetPageId="15" captionText="Figure 10. Line drawings of the last precaudal and the first two caudal vertebrae and associated elements of the anal fin in (A) Oxuderces dentatus (USNM 86954) and (B) Oxuderces nexipinnis (USNM 279359). In O. dentatus, the hemal spine extends to a point at about mid-length of the first anal-fin pterygiophore; in O. nexipinnis the hemal spine terminates closer to the proximal tip of the first anal-fin pterygiophore. Bone is stippled; cartilage is represented by open circles." figureDoi="http://doi.org/10.5281/zenodo.5710969" httpUri="https://zenodo.org/record/5710969/files/figure.png" pageId="12" pageNumber="207">10B</figureCitation>
,
<figureCitation id="F9872A784D316B328362FE2B451E9FBC" box="[1370,1404,485,511]" captionStart="Figure 14" captionStartId="18.[146,225,1675,1694]" captionTargetBox="[153,1415,198,1641]" captionTargetId="figure-93@18.[153,1415,198,1642]" captionTargetPageId="18" captionText="Figure 14. Apocryptes nexipinnis Cantor, 1849, syntypes, BMNH1860.3.19.568-69 (2), dried, filleted and stuffed with cotton wool. Anterior is to the left. Top: the larger of the two syntypes, approximately 67.8 mm SL. Middle: the same specimen as the top, rotated on the horizontal axis. Bottom: the smaller of the two syntypes, approximately 65 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5710977" httpUri="https://zenodo.org/record/5710977/files/figure.png" pageId="12" pageNumber="207">14</figureCitation>
;
</paragraph>
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<tableCitation id="2C3E03464D316B328212FDCB44EC9C5E" box="[1066,1166,515,541]" captionStart="Table 1" captionStartId="8.[88,154,502,523]" captionText="Table 1. Counts and measurements for Oxuderces dentatus and Oxuderces nexipinnis" pageId="12" pageNumber="207">TABLE 1</tableCitation>
</paragraph>
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<treatmentCitation id="E01D10EC4D316B328510FDF845659C09" author="Cantor T" box="[808,1287,565,587]" page="1170" pageId="12" pageNumber="207" year="1849">
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<emphasis id="53C8EAEF4D316B328510FDF844409C08" box="[808,1058,566,587]" italics="true" pageId="12" pageNumber="207">Apocryptes nexipinnis</emphasis>
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<bibRefCitation id="052D4B0C4D316B328214FDFB45659C09" author="Cantor T" box="[1068,1287,565,586]" pageId="12" pageNumber="207" pagination="983 - 1443" refId="ref12023" refString="Cantor T. 1849. Catalog of Malayan fishes. Journal of the Asiatic Society of Bengal 18: 983 - 1443." type="journal article" year="1849">Cantor, 1849: 1170</bibRefCitation>
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locality,
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,
<collectingCountry id="19AB766D4D316B328231FD9A44139C29" box="[1033,1137,596,618]" name="Malaysia" pageId="12" pageNumber="207">Malaysia</collectingCountry>
)
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<taxonomicName id="A6BC4D7E4D316B328579FDBD44479CCB" ID-CoL="FR3C" authorityName="Day" authorityYear="1871" box="[833,1061,627,648]" class="Actinopterygii" family="Gobiidae" genus="Apocryptes" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="207" phylum="Chordata" rank="species" species="cantoris">
<emphasis id="53C8EAEF4D316B328579FDBD44479CCB" box="[833,1061,627,648]" italics="true" pageId="12" pageNumber="207">Apocryptes cantoris</emphasis>
</taxonomicName>
not
<bibRefCitation id="052D4B0C4D316B32825FFDBD44829CC4" author="Day F" box="[1127,1248,626,648]" pageId="12" pageNumber="207" pagination="677 - 705" refId="ref12105" refString="Day F. 1871. On the fishes of the Andaman Islands. Proceedings of the General Meetings for Scientific Business of the Zoological Society London 1870: 677 - 705." type="journal article" year="1871">Day, 1871</bibRefCitation>
(in part
<bibRefCitation id="052D4B0C4D316B328362FDBD43039CE4" author="Day F" pageId="12" pageNumber="207" pagination="281 - 320" refId="ref12137" refString="Day F. 1876. The fishes of India, being a natural history of the fishes known to inhabit the seas and fresh waters of India, Burma, and Ceylon. 2: 281 - 320." type="journal article" year="1876">Day, 1876</bibRefCitation>
, specimens from Madras,
<collectingCountry id="19AB766D4D316B3282B6FD5F44A99CE4" box="[1166,1227,657,679]" name="India" pageId="12" pageNumber="207">India</collectingCountry>
)
</paragraph>
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<taxonomicName id="A6BC4D7E4D316B328579FD61452C9C86" ID-CoL="5VPM4" authority="Fowler, 1935: 162" authorityName="Fowler" authorityPageNumber="162" authorityYear="1935" box="[833,1358,687,709]" family="Gobiidae" genus="Apocryptichthys" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="207" phylum="Chordata" rank="species" species="livingstoni">
<emphasis id="53C8EAEF4D316B328579FD61441B9C87" box="[833,1145,687,708]" italics="true" pageId="12" pageNumber="207">Apocryptichthys livingstoni</emphasis>
<materialsCitation id="D1D43CA04D316B3282B9FD61452C9C86" ID-GBIF-Occurrence="3408764304" box="[1153,1358,687,709]" country="Thailand" location="Paknam" pageId="12" pageNumber="207" specimenCount="1">
<bibRefCitation id="052D4B0C4D316B3282B9FD61452C9C86" author="Fowler HW" box="[1153,1358,687,709]" pageId="12" pageNumber="207" pagination="89 - 163" refId="ref12293" refString="Fowler HW. 1935. Zoological results of the third DeSchauensee Siamese Expedition Part VI: fishes obtained in 1934. Proceedings of the Academy of Natural Sciences Philadelphia 87: 89 - 163." type="journal article" year="1935">Fowler, 1935: 162</bibRefCitation>
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(
<typeStatus id="BE07885F4D316B328365FD7E45ED9C86" box="[1373,1423,688,709]" pageId="12" pageNumber="207">type</typeStatus>
locality,
<location id="646360264D316B3285B2FD00438E9CA7" LSID="urn:lsid:plazi:treatment:E91587EB4D316B318557FE0544B69B04:646360264D316B3285B2FD00438E9CA7" box="[906,1004,718,740]" country="Thailand" name="Paknam" pageId="12" pageNumber="207">Paknam</location>
,
<collectingCountry id="19AB766D4D316B3285C0FD0044029CA7" box="[1016,1120,718,740]" name="Thailand" pageId="12" pageNumber="207">Thailand</collectingCountry>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="610336FD4D316B328510FCC245189D03" blockId="12.[808,1423,779,832]" pageId="12" pageNumber="207">
<emphasis id="53C8EAEF4D316B328510FCC2447D9D63" box="[808,1055,779,801]" italics="true" pageId="12" pageNumber="207">
<typeStatus id="BE07885F4D316B328510FCC243EF9D62" box="[808,909,780,801]" pageId="12" pageNumber="207" type="syntype">Syntypes</typeStatus>
examined:
</emphasis>
<collectionCode id="07ADAE384D316B328215FCC2441C9D62" box="[1069,1150,780,801]" pageId="12" pageNumber="207">BMNH</collectionCode>
1860.3.19.568-69, 2 undet., 65.0
<emphasis id="53C8EAEF4D316B328588FCE743DE9D03" box="[944,956,809,832]" italics="true" pageId="12" pageNumber="207">–</emphasis>
<quantity id="A6449B184D316B328584FCE444429D7C" box="[956,1056,810,832]" metricMagnitude="-2" metricUnit="m" metricValue="6.779999999999999" pageId="12" pageNumber="207" unit="mm" value="67.8">67.8 mm</quantity>
SL, Sea of
<collectingRegion id="A378F81F4D316B32829EFCE4449C9D7C" box="[1190,1278,810,831]" country="Malaysia" name="Pulau Pinang" pageId="12" pageNumber="207">Penang</collectingRegion>
,
<collectingCountry id="19AB766D4D316B328332FCE445179D03" box="[1290,1397,810,832]" name="Malaysia" pageId="12" pageNumber="207">Malaysia</collectingCountry>
.
</paragraph>
<paragraph id="610336FD4D316B328510FCA945099DF9" blockId="12.[808,1423,871,954]" pageId="12" pageNumber="207">
<emphasis id="53C8EAEF4D316B328510FCA943C39DD9" italics="true" pageId="12" pageNumber="207">
Other type material examined:
<taxonomicName id="A6BC4D7E4D316B3282E1FCA943C39DD9" authorityName="Fowler" authorityYear="1935" family="Gobiidae" genus="Apocryptichthys" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="207" phylum="Chordata" rank="species" species="livingstoni">Apocryptichthys livingstoni</taxonomicName>
</emphasis>
<typeStatus id="BE07885F4D316B328592FC4B446C9DD8" box="[938,1038,901,923]" pageId="12" pageNumber="207" type="holotype">holotype</typeStatus>
: ANSP 63091, male,
<quantity id="A6449B184D316B3282C6FC4B45029DD8" box="[1278,1376,901,923]" metricMagnitude="-2" metricUnit="m" metricValue="7.32" pageId="12" pageNumber="207" unit="mm" value="73.2">73.2 mm</quantity>
SL, mouth of Me Nam Chao Praya, Paknam,
<collectingCountry id="19AB766D4D316B3282C7FC6A450A9DF9" box="[1279,1384,932,954]" name="Thailand" pageId="12" pageNumber="207">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="610336FD4D316B328510FC2F449498D4" blockId="12.[808,1423,993,1687]" pageId="12" pageNumber="207">
<emphasis id="53C8EAEF4D316B328510FC2F44149DB5" box="[808,1142,993,1014]" italics="true" pageId="12" pageNumber="207">Other material examined:</emphasis>
Sixty-four specimens,
<quantity id="A6449B184D316B328510FBCE43AA9A56" box="[808,968,1022,1045]" metricMagnitude="-2" metricUnit="m" metricValue="5.025" metricValueMax="8.059999999999999" metricValueMin="1.9899999999999998" pageId="12" pageNumber="207" unit="mm" value="50.25" valueMax="80.6" valueMin="19.9">
19.9
<emphasis id="53C8EAEF4D316B328560FC3043069A56" box="[856,868,1022,1045]" italics="true" pageId="12" pageNumber="207">–</emphasis>
80.6 mm
</quantity>
SL. RMNH.PISC 12091, male (
<quantity id="A6449B184D316B328508FBD043FA9A77" box="[816,920,1054,1076]" metricMagnitude="-2" metricUnit="m" metricValue="6.68" pageId="12" pageNumber="207" unit="mm" value="66.8">66.8 mm</quantity>
),
<specimenCount id="77BAFD744D316B328594FBD0447B9A77" box="[940,1049,1054,1076]" pageId="12" pageNumber="207" type="female">2 females</specimenCount>
(65.9
<emphasis id="53C8EAEF4D316B328261FBD344079A77" box="[1113,1125,1053,1076]" italics="true" pageId="12" pageNumber="207">–</emphasis>
73.0 mm), east coast
<collectingRegion id="A378F81F4D316B328361FBD145ED9A77" box="[1369,1423,1055,1076]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="207">Java</collectingRegion>
Island,
<collectingCountry id="19AB766D4D316B3285B6FBF344629A10" box="[910,1024,1085,1107]" name="Indonesia" pageId="12" pageNumber="207">Indonesia</collectingCountry>
; RMNH.PISC 12092,
<specimenCount id="77BAFD744D316B328310FBF245ED9A10" box="[1320,1423,1084,1107]" pageId="12" pageNumber="207" type="male">5 males</specimenCount>
(63.0
<emphasis id="53C8EAEF4D316B328558FB94430E9A32" box="[864,876,1114,1137]" italics="true" pageId="12" pageNumber="207">–</emphasis>
<quantity id="A6449B184D316B328554FB9543B69A32" box="[876,980,1115,1137]" metricMagnitude="-2" metricUnit="m" metricValue="7.64" pageId="12" pageNumber="207" unit="mm" value="76.4">76.4 mm</quantity>
),
<specimenCount id="77BAFD744D316B3285CEFB95441F9A32" box="[1014,1149,1115,1137]" pageId="12" pageNumber="207" type="female">14 females</specimenCount>
(60.0
<emphasis id="53C8EAEF4D316B3282F3FB9444B59A32" box="[1227,1239,1114,1137]" italics="true" pageId="12" pageNumber="207">–</emphasis>
<quantity id="A6449B184D316B3282EFFB95455D9A32" box="[1239,1343,1115,1137]" metricMagnitude="-2" metricUnit="m" metricValue="7.99" pageId="12" pageNumber="207" unit="mm" value="79.9">79.9 mm</quantity>
), east coast
<collectingRegion id="A378F81F4D316B32854BFBB443CB9ACC" box="[883,937,1146,1167]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="207">Java</collectingRegion>
Island,
<collectingCountry id="19AB766D4D316B328221FBB444E99AD3" box="[1049,1163,1146,1168]" name="Indonesia" pageId="12" pageNumber="207">Indonesia</collectingCountry>
; RMNH.PISC 12433, female (
<quantity id="A6449B184D316B32859DFB56446F9AED" box="[933,1037,1176,1198]" metricMagnitude="-2" metricUnit="m" metricValue="5.56" pageId="12" pageNumber="207" unit="mm" value="55.6">55.6 mm</quantity>
),
<collectingRegion id="A378F81F4D316B32827AFB57441A9AED" box="[1090,1144,1177,1198]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="207">Java</collectingRegion>
Island,
<collectingCountry id="19AB766D4D316B328321FB5645E99AED" box="[1305,1419,1176,1198]" name="Indonesia" pageId="12" pageNumber="207">Indonesia</collectingCountry>
; RMNH.PISC 12750, female (
<quantity id="A6449B184D316B3282ACFB79449E9A8F" box="[1172,1276,1207,1228]" metricMagnitude="-2" metricUnit="m" metricValue="7.94" pageId="12" pageNumber="207" unit="mm" value="79.4">79.4 mm</quantity>
), Surabaya,
<collectingCountry id="19AB766D4D316B328510FB1B43F89AA8" box="[808,922,1237,1259]" name="Indonesia" pageId="12" pageNumber="207">Indonesia</collectingCountry>
; RMNH.PISC 33844, female (
<quantity id="A6449B184D316B328323FB1B45E19AA8" box="[1307,1411,1237,1259]" metricMagnitude="-2" metricUnit="m" metricValue="5.76" pageId="12" pageNumber="207" unit="mm" value="57.6">57.6 mm</quantity>
),
<collectingCountry id="19AB766D4D316B328510FB3A43F89B49" box="[808,922,1268,1290]" name="Indonesia" pageId="12" pageNumber="207">Indonesia</collectingCountry>
; RMNH.PISC 17382, female (72.0 mm), Pulau Weh,
<collectingRegion id="A378F81F4D316B3285F5FADD44519B6B" box="[973,1075,1299,1320]" country="Indonesia" name="Sumatera Utara" pageId="12" pageNumber="207">Sumatra</collectingRegion>
,
<collectingCountry id="19AB766D4D316B328273FADD44DF9B6A" box="[1099,1213,1299,1321]" name="Indonesia" pageId="12" pageNumber="207">Indonesia</collectingCountry>
;
<collectionCode id="07ADAE384D316B3282EDFADD45469B6B" box="[1237,1316,1299,1320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="207" type="Museum">USNM</collectionCode>
119547, female (64.0 mm), off Tachalon, Gulf of
<collectingCountry id="19AB766D4D316B328319FAFF45E99B04" box="[1313,1419,1329,1351]" name="Thailand" pageId="12" pageNumber="207">Thailand</collectingCountry>
;
<collectionCode id="07ADAE384D316B328510FA9E43159B26" box="[808,887,1360,1381]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="207" type="Museum">USNM</collectionCode>
278447, female (
<quantity id="A6449B184D316B328273FA9E44D19B26" box="[1099,1203,1360,1382]" metricMagnitude="-2" metricUnit="m" metricValue="4.36" pageId="12" pageNumber="207" unit="mm" value="43.6">43.6 mm</quantity>
), southern side of river mouth, Muar river,
<collectingRegion id="A378F81F4D316B32825BFAA044D09BC7" box="[1123,1202,1390,1412]" country="Malaysia" name="Johor" pageId="12" pageNumber="207">Johore</collectingRegion>
,
<collectingCountry id="19AB766D4D316B3282FCFAA0454D9BC7" box="[1220,1327,1390,1412]" name="Malaysia" pageId="12" pageNumber="207">Malaysia</collectingCountry>
;
<collectionCode id="07ADAE384D316B328378FAA045ED9BC0" box="[1344,1423,1390,1411]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="207" type="Museum">USNM</collectionCode>
279359,
<specimenCount id="77BAFD744D316B3285B4FA4343949BE0" box="[908,1014,1421,1443]" pageId="12" pageNumber="207" type="male">12 males</specimenCount>
(19.9
<emphasis id="53C8EAEF4D316B328201FA4244279BE0" box="[1081,1093,1420,1443]" italics="true" pageId="12" pageNumber="207">–</emphasis>
50.0 mm; 1 C&amp;S,
<quantity id="A6449B184D316B328324FA4345E19BE1" box="[1308,1411,1421,1443]" metricMagnitude="-2" metricUnit="m" metricValue="3.57" pageId="12" pageNumber="207" unit="mm" value="35.7">35.7 mm</quantity>
),
<specimenCount id="77BAFD744D316B328510FA6243C69B82" box="[808,932,1451,1473]" pageId="12" pageNumber="207" type="female">14 females</specimenCount>
(
<quantity id="A6449B184D316B32858EFA62443B9B82" box="[950,1113,1450,1473]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" metricValueMax="3.15" metricValueMin="2.21" pageId="12" pageNumber="207" unit="mm" value="26.8" valueMax="31.5" valueMin="22.1">
22.1
<emphasis id="53C8EAEF4D316B3285DEFA6443909B82" box="[998,1010,1450,1473]" italics="true" pageId="12" pageNumber="207">–</emphasis>
31.5 mm
</quantity>
; 1 C&amp;S,
<quantity id="A6449B184D316B3282FBFA6245499B82" box="[1219,1323,1452,1473]" metricMagnitude="-2" metricUnit="m" metricValue="4.79" pageId="12" pageNumber="207" unit="mm" value="47.9">47.9 mm</quantity>
);
<collectionCode id="07ADAE384D316B328378FA6245ED9B82" box="[1344,1423,1452,1473]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="207" type="Museum">USNM</collectionCode>
288662,
<specimenCount id="77BAFD744D316B3285A8FA0443819BA3" box="[912,995,1482,1504]" pageId="12" pageNumber="207" type="male">1 male</specimenCount>
(C&amp;S,
<quantity id="A6449B184D316B32827EFA0444CC9B9C" box="[1094,1198,1482,1503]" metricMagnitude="-2" metricUnit="m" metricValue="3.97" pageId="12" pageNumber="207" unit="mm" value="39.7">39.7 mm</quantity>
),
<specimenCount id="77BAFD744D316B3282F0FA04455E9BA3" box="[1224,1340,1482,1504]" pageId="12" pageNumber="207" type="female">2 females</specimenCount>
(C&amp;S,
<quantity id="A6449B184D316B328510FA2643AE9BBD" box="[808,972,1511,1534]" metricMagnitude="-2" metricUnit="m" metricValue="2.42" metricValueMax="2.68" metricValueMin="2.16" pageId="12" pageNumber="207" unit="mm" value="24.2" valueMax="26.8" valueMin="21.6">
21.6
<emphasis id="53C8EAEF4D316B328560FA2943069BBD" box="[856,868,1511,1534]" italics="true" pageId="12" pageNumber="207">–</emphasis>
26.8 mm
</quantity>
), southern side of river mouth, Muar river,
<collectingRegion id="A378F81F4D316B328548F9C943DD985E" box="[880,959,1543,1565]" country="Malaysia" name="Johor" pageId="12" pageNumber="207">Johore</collectingRegion>
,
<collectingCountry id="19AB766D4D316B3285F4F9C94455985E" box="[972,1079,1543,1565]" name="Malaysia" pageId="12" pageNumber="207">Malaysia</collectingCountry>
; ZRC 39777, male (
<quantity id="A6449B184D316B328323F9C945E1985E" box="[1307,1411,1543,1565]" metricMagnitude="-2" metricUnit="m" metricValue="8.059999999999999" pageId="12" pageNumber="207" unit="mm" value="80.6">80.6 mm</quantity>
), Sungai Selising, Matang mangroves,
<collectingRegion id="A378F81F4D316B32837CF9E845E8987F" box="[1348,1418,1574,1596]" country="Malaysia" name="Perak" pageId="12" pageNumber="207">Perak</collectingRegion>
,
<collectingCountry id="19AB766D4D316B328510F98A43F19819" box="[808,915,1604,1626]" name="Malaysia" pageId="12" pageNumber="207">Malaysia</collectingCountry>
; ZRC 53919,
<specimenCount id="77BAFD744D316B32827FF98A44CB9819" box="[1095,1193,1604,1626]" pageId="12" pageNumber="207" type="male">3 males</specimenCount>
(
<quantity id="A6449B184D316B3282F9F98A45079819" box="[1217,1381,1603,1626]" metricMagnitude="-2" metricUnit="m" metricValue="5.175" metricValueMax="5.32" metricValueMin="5.029999999999999" pageId="12" pageNumber="207" unit="mm" value="51.75" valueMax="53.2" valueMin="50.3">
50.3
<emphasis id="53C8EAEF4D316B3282C9F98D449F9819" box="[1265,1277,1603,1626]" italics="true" pageId="12" pageNumber="207">–</emphasis>
53.2 mm
</quantity>
),
<specimenCount id="77BAFD744D316B3283B9F98A431D983A" pageId="12" pageNumber="207" type="female">3 females</specimenCount>
(
<quantity id="A6449B184D316B3285AFF9AD445E983B" box="[919,1084,1634,1657]" metricMagnitude="-2" metricUnit="m" metricValue="4.334999999999999" metricValueMax="5.33" metricValueMin="3.34" pageId="12" pageNumber="207" unit="mm" value="43.349999999999994" valueMax="53.3" valueMin="33.4">
33.4
<emphasis id="53C8EAEF4D316B3285F0F9AC43B6983A" box="[968,980,1634,1657]" italics="true" pageId="12" pageNumber="207">–</emphasis>
53.3 mm
</quantity>
), Matla mudflats, Jharkali, South 24 Parganas,
<collectingRegion id="A378F81F4D316B32822AF94C44C798D4" box="[1042,1189,1665,1687]" country="India" name="West Bengal" pageId="12" pageNumber="207">West Bengal</collectingRegion>
,
<collectingCountry id="19AB766D4D316B32828AF94F449098D4" box="[1202,1266,1665,1687]" name="India" pageId="12" pageNumber="207">India</collectingCountry>
.
</paragraph>
<paragraph id="610336FD4D316B338510F97042049F7B" blockId="12.[808,1423,1726,1901]" lastBlockId="13.[166,781,199,312]" lastPageId="13" lastPageNumber="208" pageId="12" pageNumber="207">
<emphasis id="53C8EAEF4D316B328510F97045389897" box="[808,1370,1726,1748]" italics="true" pageId="12" pageNumber="207">
Differential diagnosis:
<taxonomicName id="A6BC4D7E4D316B328276F97045389897" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[1102,1370,1726,1748]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="207" phylum="Chordata" rank="species" species="nexipinnis">Oxuderces nexipinnis</taxonomicName>
</emphasis>
is differentiated from
<taxonomicName id="A6BC4D7E4D316B32821BF91344CB98B1" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[1059,1193,1757,1778]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="207" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D316B32821BF91344CB98B1" box="[1059,1193,1757,1778]" italics="true" pageId="12" pageNumber="207">O. dentatus</emphasis>
</taxonomicName>
by the following characters: first hemal spine terminating above midlength of first anal-fin pterygiophore (vs. extending ventrally below mid-length of first anal-fin pterygiophore;
<figureCitation id="F9872A784D316B3285CAF8994426992E" box="[1010,1092,1879,1901]" captionStart="Figure 10" captionStartId="15.[829,908,1600,1619]" captionTargetBox="[173,774,204,1740]" captionTargetId="figure-441@15.[176,766,275,1733]" captionTargetPageId="15" captionText="Figure 10. Line drawings of the last precaudal and the first two caudal vertebrae and associated elements of the anal fin in (A) Oxuderces dentatus (USNM 86954) and (B) Oxuderces nexipinnis (USNM 279359). In O. dentatus, the hemal spine extends to a point at about mid-length of the first anal-fin pterygiophore; in O. nexipinnis the hemal spine terminates closer to the proximal tip of the first anal-fin pterygiophore. Bone is stippled; cartilage is represented by open circles." figureDoi="http://doi.org/10.5281/zenodo.5710969" httpUri="https://zenodo.org/record/5710969/files/figure.png" pageId="12" pageNumber="207">Fig. 10</figureCitation>
); mouth superior (vs. terminal); conspicuous dermal invagination posterior to the point of attachment of pelvic-fin base (vs. dermal invagination absent;
<figureCitation id="F9872A784D306B3387C6FECA42219F59" box="[510,579,260,282]" captionStart="Figure 6" captionStartId="9.[166,245,730,749]" captionTargetBox="[173,774,198,697]" captionTargetId="figure-567@9.[173,774,198,697]" captionTargetPageId="9" captionText="Figure 6. Dermal invagination or pouch (arrow) at point of attachment of the pelvic-fin base in Oxuderces nexipinnis (ZRC 53919, 53.2 mm SL). Anterior is to the right." figureDoi="http://doi.org/10.5281/zenodo.5710958" httpUri="https://zenodo.org/record/5710958/files/figure.png" pageId="13" pageNumber="208">Fig. 6</figureCitation>
); head relatively long (27.3
<emphasis id="53C8EAEF4D306B33872FFEEF41419F7B" box="[279,291,289,312]" italics="true" pageId="13" pageNumber="208">–</emphasis>
30.3%SL vs. 25.1
<emphasis id="53C8EAEF4D306B3387DEFEEF41909F7B" box="[486,498,289,312]" italics="true" pageId="13" pageNumber="208">–</emphasis>
27.7%SL).
</paragraph>
</subSubSection>
<caption id="35C366754D306B338505F9E1439E9925" ID-DOI="http://doi.org/10.5281/zenodo.5710963" ID-Zenodo-Dep="5710963" httpUri="https://zenodo.org/record/5710963/files/figure.png" pageId="13" pageNumber="208" startId="13.[829,908,1583,1602]" targetBox="[173,774,345,1855]" targetPageId="13">
<paragraph id="610336FD4D306B338505F9E1439E9925" blockId="13.[829,1445,1583,1894]" pageId="13" pageNumber="208">
<emphasis id="53C8EAEF4D306B338505F9E143C59801" bold="true" box="[829,935,1583,1602]" pageId="13" pageNumber="208">Figure 8.</emphasis>
Line drawings of the hyoid bar in (A)
<taxonomicName id="A6BC4D7E4D306B33835CF9E14385981C" baseAuthorityName="Cantor" baseAuthorityYear="1849" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="208" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D306B33835CF9E14385981C" italics="true" pageId="13" pageNumber="208">Oxuderces nexipinnis</emphasis>
</taxonomicName>
(USNM 279359), (B)
<taxonomicName id="A6BC4D7E4D306B3382EEF98245C6981C" box="[1238,1444,1612,1631]" class="Actinopterygii" family="Gobiidae" genus="Apocryptodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="208" phylum="Chordata" rank="species" species="wirzi">
<emphasis id="53C8EAEF4D306B3382EEF98245C6981C" box="[1238,1444,1612,1631]" italics="true" pageId="13" pageNumber="208">Apocryptodon wirzi</emphasis>
</taxonomicName>
(USNM 287292) and (C)
<taxonomicName id="A6BC4D7E4D306B33827FF9A74489983E" box="[1095,1259,1641,1661]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="208" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="53C8EAEF4D306B33827FF9A744A7983F" box="[1095,1221,1641,1660]" italics="true" pageId="13" pageNumber="208">Gnatholepis</emphasis>
sp.
</taxonomicName>
(USNM 287166). The anterior ceratohyal posterior to insertion of the fourth branchiostegal ray is elongate and notched (arrow) in
<taxonomicName id="A6BC4D7E4D306B338559F90F438C9890" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[865,1006,1729,1748]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="208" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D306B338559F90F438C9890" box="[865,1006,1729,1748]" italics="true" pageId="13" pageNumber="208">O. nexipinnis</emphasis>
</taxonomicName>
, elongate and not notched (arrow) in
<taxonomicName id="A6BC4D7E4D306B338505F91043F398B2" box="[829,913,1758,1777]" class="Actinopterygii" family="Gobiidae" genus="Apocryptodon" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="208" phylum="Chordata" rank="species" species="wirzi">
<emphasis id="53C8EAEF4D306B338505F91043F398B2" box="[829,913,1758,1777]" italics="true" pageId="13" pageNumber="208">A. wirzi</emphasis>
</taxonomicName>
, and not elongate and not notched in
<taxonomicName id="A6BC4D7E4D306B33831EF9104339994C" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="208" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="53C8EAEF4D306B33831EF91045C698B2" box="[1318,1444,1758,1777]" italics="true" pageId="13" pageNumber="208">Gnatholepis</emphasis>
sp.
</taxonomicName>
Branchiostegal rays 3
<emphasis id="53C8EAEF4D306B33826FF9304400994D" box="[1111,1122,1790,1806]" italics="true" pageId="13" pageNumber="208">–</emphasis>
5 are distinctly thinner in A and B. Bone is stippled, cartilage is represented by open circles. Only the proximal portion of the five branchiostegal rays is drawn.
</paragraph>
</caption>
<subSubSection id="29A665764D306B3086CBF88844FB9EB8" lastPageId="14" lastPageNumber="209" pageId="13" pageNumber="208" type="description">
<paragraph id="610336FD4D306B3386CBF88841219927" blockId="13.[243,323,1862,1892]" box="[243,323,1862,1892]" pageId="13" pageNumber="208">
<quantity id="A6449B184D306B3386CBF88841219927" box="[243,323,1862,1892]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="208" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
<paragraph id="610336FD4D306B338505FF0945C69BA3" blockId="13.[829,1444,199,1504]" pageId="13" pageNumber="208">
<emphasis id="53C8EAEF4D306B338505FF0943A49E9F" box="[829,966,199,220]" italics="true" pageId="13" pageNumber="208">Description:</emphasis>
Head markedly depressed, wider than deep, head length 27.3
<emphasis id="53C8EAEF4D306B338268FF2A443E9EB8" box="[1104,1116,228,251]" italics="true" pageId="13" pageNumber="208">–</emphasis>
30.3%SL, head width 32.6
<emphasis id="53C8EAEF4D306B3383A0FF2A45C69EB8" box="[1432,1444,228,251]" italics="true" pageId="13" pageNumber="208">–</emphasis>
44.4%HL, head depth 30.4
<emphasis id="53C8EAEF4D306B338244FECD44EA9F59" box="[1148,1160,259,282]" italics="true" pageId="13" pageNumber="208">–</emphasis>
42.7%HL. Eye diameter 8.7
<emphasis id="53C8EAEF4D306B338567FEEF43099F7B" box="[863,875,289,312]" italics="true" pageId="13" pageNumber="208">–</emphasis>
16.3%HL; interorbital distance 3.0
<emphasis id="53C8EAEF4D306B33833BFEEF456D9F7B" box="[1283,1295,289,312]" italics="true" pageId="13" pageNumber="208">–</emphasis>
4.8%HL, eye without dermal cup. Snout profile pointed, snout length 12.0
<emphasis id="53C8EAEF4D306B3385F0FE9043B69F36" box="[968,980,350,373]" italics="true" pageId="13" pageNumber="208">–</emphasis>
15.5%HL. Mouth superior, lower jaw terminating just anterior to upper jaw, gape wide; jaw length 41.3
<emphasis id="53C8EAEF4D306B3385D7FE5543999FF1" box="[1007,1019,411,434]" italics="true" pageId="13" pageNumber="208">–</emphasis>
59.7%HL; distinct notch in middle of upper lip between two medial teeth. Teeth in both jaws caninoid, unevenly spaced, in single row; one or two prominent and elongate canines on each side of premaxillary symphysis, canine teeth extending slightly anteroventrally and reaching beyond lower jaw when mouth closed, teeth decreasing in length posteriorly; teeth in lower jaw more uniform in size, teeth all shorter than medial canine teeth of upper jaw, teeth absent posteriorly, no canine tooth on each side of symphysis internal to anterior margin of lower jaw. Body compressed and gracile; body depth at anus 8.6
<emphasis id="53C8EAEF4D306B3385FEFCC443B09D62" box="[966,978,778,801]" italics="true" pageId="13" pageNumber="208">–</emphasis>
12.9%SL, body width at anus 4.5
<emphasis id="53C8EAEF4D306B338359FCC4450F9D62" box="[1377,1389,778,801]" italics="true" pageId="13" pageNumber="208">–</emphasis>
7.3% SL. Predorsal long, predorsal length 28.2
<emphasis id="53C8EAEF4D306B338315FCE7455B9D03" box="[1325,1337,809,832]" italics="true" pageId="13" pageNumber="208">–</emphasis>
31.6%SL. D1 and D2 connected by membrane for entire height, base of D1 and D2 54.9
<emphasis id="53C8EAEF4D306B338259FCA8440F9D3E" box="[1121,1133,870,893]" italics="true" pageId="13" pageNumber="208">–</emphasis>
59.4%SL; base of anal fin 35.9
<emphasis id="53C8EAEF4D306B338555FC4A431B9DD8" box="[877,889,900,923]" italics="true" pageId="13" pageNumber="208">–</emphasis>
40.4%SL. No membrane connecting D2 or A to caudal fin. Pelvic fin short and rounded, not reaching genital papilla, pelvic-fin length 14.0
<emphasis id="53C8EAEF4D306B33836BFC0F453D9D9B" box="[1363,1375,961,984]" italics="true" pageId="13" pageNumber="208">–</emphasis>
18.2% SL. Pectoral-fin length moderate, 14.0
<emphasis id="53C8EAEF4D306B338315FC2E455B9DB4" box="[1325,1337,992,1015]" italics="true" pageId="13" pageNumber="208">–</emphasis>
19.2%SL. Caudal fin long, lanceolate, 20.8
<emphasis id="53C8EAEF4D306B3382F8FC3144AE9A55" box="[1216,1228,1023,1046]" italics="true" pageId="13" pageNumber="208">–</emphasis>
24.8%SL. D1 with six spinous dorsal rays (VI); all elements of D2 and A fins segmented; D2 with 25
<emphasis id="53C8EAEF4D306B33828BFBF244DD9A10" box="[1203,1215,1084,1107]" italics="true" pageId="13" pageNumber="208">–</emphasis>
26 elements, A fin with 24
<emphasis id="53C8EAEF4D306B3385ADFB9443C39A32" box="[917,929,1114,1137]" italics="true" pageId="13" pageNumber="208">–</emphasis>
26 elements; pectoral rays 21
<emphasis id="53C8EAEF4D306B3382C1FB9445679A32" box="[1273,1285,1114,1137]" italics="true" pageId="13" pageNumber="208">–</emphasis>
24; caudal fin segmented rays 17; dorsal procurrent rays 5, ventral procurrent rays 4. Two epural bones. Lateral longitudinal scale count 50
<emphasis id="53C8EAEF4D306B338241FB7844E79A8E" box="[1145,1157,1206,1229]" italics="true" pageId="13" pageNumber="208">–</emphasis>
54, scales on body small and embedded in skin anteriorly, and increasing in size posteriorly. All specimens without scales on dorsal region anterior to D1, ventral region of the head, cheek, operculum and pectoral-fin base except for largest specimen examined (ZRC 39777,
<quantity id="A6449B184D306B338505FAA043C09BC7" box="[829,930,1390,1412]" metricMagnitude="-2" metricUnit="m" metricValue="8.059999999999999" pageId="13" pageNumber="208" unit="mm" value="80.6">80.6 mm</quantity>
) which has cheek scales and 14 predorsal midline scales; all scales small and embedded. Raised and distinct free-neuromast rows on head. Conspicuous dermal invagination posterior to point
</paragraph>
<paragraph id="610336FD4D336B30869FFF0242929EBC" blockId="14.[167,752,204,255]" box="[167,752,204,255]" pageId="14" pageNumber="209">A First dorsal-fin anterior elements</paragraph>
<paragraph id="610336FD4D336B308510FF0944FB9EB8" blockId="14.[808,1423,199,251]" pageId="14" pageNumber="209">
of attachment of pelvic-fin base, invagination deep and appearing as a ‘pit’ (
<figureCitation id="F9872A784D336B30827FFF2B44EE9EB8" box="[1095,1164,229,251]" captionStart="Figure 6" captionStartId="9.[166,245,730,749]" captionTargetBox="[173,774,198,697]" captionTargetId="figure-567@9.[173,774,198,697]" captionTargetPageId="9" captionText="Figure 6. Dermal invagination or pouch (arrow) at point of attachment of the pelvic-fin base in Oxuderces nexipinnis (ZRC 53919, 53.2 mm SL). Anterior is to the right." figureDoi="http://doi.org/10.5281/zenodo.5710958" httpUri="https://zenodo.org/record/5710958/files/figure.png" pageId="14" pageNumber="209">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="29A665764D336B308510FEEC4412987F" pageId="14" pageNumber="209" type="discussion">
<paragraph id="610336FD4D336B308510FEEC4412987F" blockId="14.[808,1424,290,1596]" pageId="14" pageNumber="209">
<emphasis id="53C8EAEF4D336B308510FEEC43C59F74" box="[808,935,290,311]" italics="true" pageId="14" pageNumber="209">Coloration:</emphasis>
Preserved specimens uniformly light to dark brown with black spots on dorsal region of head and nape. Darkened brown to black upper lip and pectoral-fin base. Dark brown spot between 20
<superScript id="96C99BB54D336B308370FEB645389FC5" attach="left" box="[1352,1370,376,390]" fontSize="6" pageId="14" pageNumber="209">th</superScript>
and penultimate rays, positioned at mid-height of D2.
<bibRefCitation id="052D4B0C4D336B308510FE75446A9F92" author="Murdy EO" box="[808,1032,443,465]" pageId="14" pageNumber="209" pagination="1 - 93" refId="ref12918" refString="Murdy EO. 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Records of the Australian Museum 11 (Supplement): 1 - 93." type="journal article" year="1989">Murdy (1989: 20)</bibRefCitation>
based his description of live coloration on material from Peninsular
<collectingCountry id="19AB766D4D336B30831FFE1445ED9FB3" box="[1319,1423,474,496]" name="Malaysia" pageId="14" pageNumber="209">Malaysia</collectingCountry>
that he had identified as
<taxonomicName id="A6BC4D7E4D336B308269FE3744B59C4E" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[1105,1239,504,526]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D336B308269FE3744B59C4E" box="[1105,1239,504,526]" italics="true" pageId="14" pageNumber="209">O. dentatus</emphasis>
</taxonomicName>
(=
<taxonomicName id="A6BC4D7E4D336B3082CEFE3745ED9C4D" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[1270,1423,505,526]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D336B3082CEFE3745ED9C4D" box="[1270,1423,505,526]" italics="true" pageId="14" pageNumber="209">O. nexipinnis</emphasis>
</taxonomicName>
here): ‘Head and trunk greyish blue; 7 greyish-blue spots along lateral midline starting at a point equal to origin of first dorsal fin and ending at a point equal to about 75% of the length of D2; a suffusion of shiny, bluish scales on posterior half of trunk; 6 dusky, saddle-like blotches on dorsum starting at terminus of D1 with 4 across the base of D2 and 1 on caudal peduncle; D1 translucent; D2 translucent except for a thin, faint dusky stripe through the median part of fin and a large black ocellus near distal tips of last 4 rays; caudal, anal and pelvic fins translucent; pectoral fin translucent but with a large, dusky blotch on upper base; nape with dusky reticulations; anterior nostril with a black anterior margin; venter shiny white.’ In the
<collectingCountry id="19AB766D4D336B3082ECFC0D45529D9A" box="[1236,1328,963,985]" name="Malaysia" pageId="14" pageNumber="209">Malacca</collectingCountry>
Straits, along the western coast of Peninsular
<collectingCountry id="19AB766D4D336B30831AFC2F45E99DB4" box="[1314,1419,993,1015]" name="Malaysia" pageId="14" pageNumber="209">Malaysia</collectingCountry>
,
<bibRefCitation id="052D4B0C4D336B308510FBCE456D9A55" author="Takita T &amp; Agusnimar A &amp; Ali AB" box="[808,1295,1024,1046]" pageId="14" pageNumber="209" pagination="131 - 138" refId="ref13467" refString="Takita T, Agusnimar A, Ali AB. 1999. Distribution and habitat requirements of oxudercine gobies (Gobiidae: Oxudercinae) along the Straits of Malacca. Ichthyological Research 46: 131 - 138." type="journal article" year="1999">Takita, Agusnimar &amp; Ali (1999: 132)</bibRefCitation>
identified
<taxonomicName id="A6BC4D7E4D336B308510FBD143DD9A77" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[808,959,1055,1076]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D336B308510FBD143DD9A77" box="[808,959,1055,1076]" italics="true" pageId="14" pageNumber="209">O. nexipinnis</emphasis>
</taxonomicName>
(reported as
<taxonomicName id="A6BC4D7E4D336B3082B7FBD145709A70" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[1167,1298,1054,1076]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D336B3082B7FBD145709A70" box="[1167,1298,1054,1076]" italics="true" pageId="14" pageNumber="209">O. dentatus</emphasis>
</taxonomicName>
) by the characteristic ‘...bright green eyes and depressed head, and a dark spot posteriorly on the dorsal fin’. They reported that
<taxonomicName id="A6BC4D7E4D336B3085F2FBB444039AD3" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[970,1121,1146,1168]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D336B3085F2FBB444039AD3" box="[970,1121,1146,1168]" italics="true" pageId="14" pageNumber="209">O. nexipinnis</emphasis>
</taxonomicName>
occurs on the mudflats of Bengkalis and
<collectingRegion id="A378F81F4D336B3085E8FB5744479AED" box="[976,1061,1177,1198]" country="Malaysia" name="Pulau Pinang" pageId="14" pageNumber="209">Penang</collectingRegion>
islands in Peninsular
<collectingCountry id="19AB766D4D336B30831AFB5645E99AED" box="[1314,1419,1176,1198]" name="Malaysia" pageId="14" pageNumber="209">Malaysia</collectingCountry>
, at sites facing the sea and not influenced by freshwater inflow (
<bibRefCitation id="052D4B0C4D336B3085BFFB18440D9AAF" author="Takita T &amp; Agusnimar A &amp; Ali AB" box="[903,1135,1238,1260]" pageId="14" pageNumber="209" pagination="131 - 138" refId="ref13467" refString="Takita T, Agusnimar A, Ali AB. 1999. Distribution and habitat requirements of oxudercine gobies (Gobiidae: Oxudercinae) along the Straits of Malacca. Ichthyological Research 46: 131 - 138." type="journal article" year="1999">
Takita
<emphasis id="53C8EAEF4D336B3085DCFB18447F9AA8" box="[996,1053,1238,1259]" italics="true" pageId="14" pageNumber="209">et al.</emphasis>
, 1999
</bibRefCitation>
). At Teluk Bahang in
<collectingRegion id="A378F81F4D336B308510FB3B431F9B49" box="[808,893,1269,1290]" country="Malaysia" name="Pulau Pinang" pageId="14" pageNumber="209">Penang</collectingRegion>
Island, this species was found low on the intertidal zone, an area which was exposed for only several days per lunar cycle, prompting the hypothesis that
<taxonomicName id="A6BC4D7E4D336B308554FA9E44619B26" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[876,1027,1360,1381]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D336B308554FA9E44619B26" box="[876,1027,1360,1381]" italics="true" pageId="14" pageNumber="209">O. nexipinnis</emphasis>
</taxonomicName>
is probably active underwater (
<bibRefCitation id="052D4B0C4D336B308508FAA044629BC6" author="Takita T &amp; Agusnimar A &amp; Ali AB" box="[816,1024,1390,1413]" pageId="14" pageNumber="209" pagination="131 - 138" refId="ref13467" refString="Takita T, Agusnimar A, Ali AB. 1999. Distribution and habitat requirements of oxudercine gobies (Gobiidae: Oxudercinae) along the Straits of Malacca. Ichthyological Research 46: 131 - 138." type="journal article" year="1999">
Takita
<emphasis id="53C8EAEF4D336B3085B9FAA143D89BC0" box="[897,954,1390,1412]" italics="true" pageId="14" pageNumber="209">et al.</emphasis>
, 1999
</bibRefCitation>
). This hypothesis is corroborated by
<bibRefCitation id="052D4B0C4D336B308510FA4344259BE0" author="Polgar G &amp; Bartolino V" box="[808,1095,1421,1443]" pageId="14" pageNumber="209" pagination="199 - 212" refId="ref13245" refString="Polgar G, Bartolino V. 2010. Size variation of six species of oxudercine gobies along the intertidal zone in a Malayan coastal swamp. Marine Ecology Progress Series 409: 199 - 212." type="book chapter" year="2010">Polgar &amp; Bartolino (2010)</bibRefCitation>
who found larger individuals of
<taxonomicName id="A6BC4D7E4D336B30857DFA6243B99B82" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[837,987,1452,1473]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D336B30857DFA6243B99B82" box="[837,987,1452,1473]" italics="true" pageId="14" pageNumber="209">O. nexipinnis</emphasis>
</taxonomicName>
(reported as
<taxonomicName id="A6BC4D7E4D336B30824BFA6244949B82" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[1139,1270,1452,1473]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D336B30824BFA6244949B82" box="[1139,1270,1452,1473]" italics="true" pageId="14" pageNumber="209">O. dentatus</emphasis>
</taxonomicName>
) in the lower intertidal zones (more aquatic) and smaller individuals in higher intertidal zones (more terrestrial) at their study site in Tanjung Piai,
<collectingRegion id="A378F81F4D336B308256F9C944D2985E" box="[1134,1200,1543,1565]" country="Malaysia" name="Johor" pageId="14" pageNumber="209">Johor</collectingRegion>
, along the western coast of Peninsular
<collectingCountry id="19AB766D4D336B30823AF9E8440E987F" box="[1026,1132,1574,1596]" name="Malaysia" pageId="14" pageNumber="209">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="35C366754D336B3086AAF9E340B99969" ID-DOI="http://doi.org/10.5281/zenodo.5710967" ID-Zenodo-Dep="5710967" httpUri="https://zenodo.org/record/5710967/files/figure.png" pageId="14" pageNumber="209" startId="14.[146,225,1581,1600]" targetBox="[156,752,291,1549]" targetPageId="14">
<paragraph id="610336FD4D336B3086AAF9E340B99969" blockId="14.[146,760,1581,1834]" pageId="14" pageNumber="209">
<emphasis id="53C8EAEF4D336B3086AAF9E3409E9803" bold="true" box="[146,252,1581,1600]" pageId="14" pageNumber="209">Figure 9.</emphasis>
Line drawings of several anterior vertebrae and associated elements of the first dorsal fin in (A)
<taxonomicName id="A6BC4D7E4D336B3086AAF9A641E99839" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[146,395,1640,1659]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D336B3086AAF9A641E99839" box="[146,395,1640,1659]" italics="true" pageId="14" pageNumber="209">Oxuderces nexipinnis</emphasis>
</taxonomicName>
(USNM 279359) and (B)
<taxonomicName id="A6BC4D7E4D336B3086AAF94B415F98DB" box="[146,317,1669,1688]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="53C8EAEF4D336B3086AAF94B417298DB" box="[146,272,1669,1688]" italics="true" pageId="14" pageNumber="209">Gnatholepis</emphasis>
sp.
</taxonomicName>
(USNM 287166). The neural spine of the fourth vertebra (arrow) is broad and spatulate and directed posteriorly in
<taxonomicName id="A6BC4D7E4D336B30878AF90E425D9891" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[434,575,1728,1747]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D336B30878AF90E425D9891" box="[434,575,1728,1747]" italics="true" pageId="14" pageNumber="209">O. nexipinnis</emphasis>
</taxonomicName>
and relatively thin and pointed and directed dorsally in
<taxonomicName id="A6BC4D7E4D336B30844CF913429098B3" authorityName="Bleeker" authorityYear="1874" box="[628,754,1757,1776]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="genus">
<emphasis id="53C8EAEF4D336B30844CF913429098B3" box="[628,754,1757,1776]" italics="true" pageId="14" pageNumber="209">Gnatholepis</emphasis>
</taxonomicName>
. Bone is stippled, cartilage is represented by open circles.
</paragraph>
</caption>
<subSubSection id="29A665764D336B308510F990444E98F2" pageId="14" pageNumber="209" type="etymology">
<paragraph id="610336FD4D336B308510F990444E98F2" blockId="14.[808,1423,1630,1713]" pageId="14" pageNumber="209">
<emphasis id="53C8EAEF4D336B308510F99043C49830" box="[808,934,1630,1651]" italics="true" pageId="14" pageNumber="209">Etymology:</emphasis>
Latin, ‘
<emphasis id="53C8EAEF4D336B30822CF99044389830" box="[1044,1114,1630,1651]" italics="true" pageId="14" pageNumber="209">nexus’</emphasis>
meaning ‘tied together’ and ‘
<emphasis id="53C8EAEF4D336B30854DF9B343DF98D1" box="[885,957,1661,1682]" italics="true" pageId="14" pageNumber="209">pinnis</emphasis>
’ meaning ‘fins’, referring to the continuous dorsal fins.
</paragraph>
</subSubSection>
<subSubSection id="29A665764D336B318510F916415B9969" lastPageId="15" lastPageNumber="210" pageId="14" pageNumber="209" type="distribution">
<paragraph id="610336FD4D336B308510F91645ED9968" blockId="14.[808,1423,1752,1835]" pageId="14" pageNumber="209">
<emphasis id="53C8EAEF4D336B308510F91643FC994F" italics="true" pageId="14" pageNumber="209">
Distribution and ecological note:
<taxonomicName id="A6BC4D7E4D336B308322F91643FC994F" baseAuthorityName="Cantor" baseAuthorityYear="1849" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="209" phylum="Chordata" rank="species" species="nexipinnis">Oxuderces nexipinnis</taxonomicName>
</emphasis>
is distributed in the Indo-West Pacific, here reported from the southern part of the South
</paragraph>
<caption id="35C366754D326B318505F98E4401997E" ID-DOI="http://doi.org/10.5281/zenodo.5710969" ID-Zenodo-Dep="5710969" httpUri="https://zenodo.org/record/5710969/files/figure.png" pageId="15" pageNumber="210" startId="15.[829,908,1600,1619]" targetBox="[173,774,204,1740]" targetPageId="15">
<paragraph id="610336FD4D326B318505F98E4401997E" blockId="15.[829,1444,1600,1853]" pageId="15" pageNumber="210">
<emphasis id="53C8EAEF4D326B318505F98E43D69810" bold="true" box="[829,948,1600,1619]" pageId="15" pageNumber="210">Figure 10.</emphasis>
Line drawings of the last precaudal and the first two caudal vertebrae and associated elements of the anal fin in (A)
<taxonomicName id="A6BC4D7E4D326B3185E2F9B544CA98CD" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[986,1192,1659,1678]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D326B3185E2F9B544CA98CD" box="[986,1192,1659,1678]" italics="true" pageId="15" pageNumber="210">Oxuderces dentatus</emphasis>
</taxonomicName>
(USNM 86954) and (B)
<taxonomicName id="A6BC4D7E4D326B318505F956444798E9" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[829,1061,1688,1707]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D326B318505F956444798E9" box="[829,1061,1688,1707]" italics="true" pageId="15" pageNumber="210">Oxuderces nexipinnis</emphasis>
</taxonomicName>
(USNM 279359). In
<taxonomicName id="A6BC4D7E4D326B31831AF95645FC98E8" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[1314,1438,1688,1707]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D326B31831AF95645FC98E8" box="[1314,1438,1688,1707]" italics="true" pageId="15" pageNumber="210">O. dentatus</emphasis>
</taxonomicName>
, the hemal spine extends to a point at about mid-length of the first anal-fin pterygiophore; in
<taxonomicName id="A6BC4D7E4D326B3182DBF91D451298A6" baseAuthorityName="Cantor" baseAuthorityYear="1849" box="[1251,1392,1747,1766]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D326B3182DBF91D451298A6" box="[1251,1392,1747,1766]" italics="true" pageId="15" pageNumber="210">O. nexipinnis</emphasis>
</taxonomicName>
the hemal spine terminates closer to the proximal tip of the first anal-fin pterygiophore. Bone is stippled; cartilage is represented by open circles.
</paragraph>
</caption>
<paragraph id="610336FD4D326B3186D0F8C2415B9969" blockId="15.[232,313,1804,1834]" box="[232,313,1804,1834]" pageId="15" pageNumber="210">
<quantity id="A6449B184D326B3186D0F8C2415B9969" box="[232,313,1804,1834]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="210" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
</subSubSection>
<subSubSection id="29A665764D326B318505FF0944B69B04" pageId="15" pageNumber="210" type="discussion">
<paragraph id="610336FD4D326B318505FF0944C09CCB" blockId="15.[829,1444,199,648]" pageId="15" pageNumber="210">
<collectingCountry id="19AB766D4D326B318505FF0943E09E9E" box="[829,898,199,221]" name="China" pageId="15" pageNumber="210">China</collectingCountry>
Sea, Gulf of
<collectingCountry id="19AB766D4D326B31820CFF0944FC9E9E" box="[1076,1182,199,221]" name="Thailand" pageId="15" pageNumber="210">Thailand</collectingCountry>
, Java Sea, Andaman Sea and Bay of Bengal (
<figureCitation id="F9872A784D326B318250FF2B44D99EB8" box="[1128,1211,229,251]" captionStart="Figure 13" captionStartId="17.[166,245,1183,1202]" captionTargetBox="[174,1440,198,1150]" captionTargetId="figure-335@17.[174,1441,198,1150]" captionTargetPageId="17" captionText="Figure 13. Distribution map of Oxuderces dentatus (solid circles) and Oxuderces nexipinnis (solid triangles). Each symbol represents one catalogued museum lot. Distributional limits are approximated by a solid line." figureDoi="http://doi.org/10.5281/zenodo.5710975" httpUri="https://zenodo.org/record/5710975/files/figure.png" pageId="15" pageNumber="210">Fig. 13</figureCitation>
). This species lives on intertidal soft-bottom habitats, predominantly mudflats. Adults have been observed to swim through the muddy substrate with only a thin film of water above them, through which they would occasionally poke their heads (
<bibRefCitation id="052D4B0C4D326B318298FEB045549FD0" author="Murdy EO" box="[1184,1334,382,404]" pageId="15" pageNumber="210" pagination="1 - 93" refId="ref12918" refString="Murdy EO. 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Records of the Australian Museum 11 (Supplement): 1 - 93." type="journal article" year="1989">Murdy, 1989</bibRefCitation>
, reported as
<taxonomicName id="A6BC4D7E4D326B31855CFE5343889FF1" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[868,1002,413,434]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D326B31855CFE5343889FF1" box="[868,1002,413,434]" italics="true" pageId="15" pageNumber="210">O. dentatus</emphasis>
</taxonomicName>
). This species has been reported to store air in its burrows, especially in the spawning chambers (
<bibRefCitation id="052D4B0C4D326B3185F5FE1444929FAC" author="Ishimatsu A &amp; Hishida Y &amp; Takita T &amp; Kanda T &amp; Oikawa S &amp; Takeda T &amp; Khoo KH" box="[973,1264,474,496]" pageId="15" pageNumber="210" pagination="237 - 238" refId="ref12691" refString="Ishimatsu A, Hishida Y, Takita T, Kanda T, Oikawa S, Takeda T, Khoo KH. 1998. Mudskippers store air in their burrows. Nature 391: 237 - 238." type="journal article" year="1998">
Ishimatsu
<emphasis id="53C8EAEF4D326B318264FE1444F49FAC" box="[1116,1174,474,495]" italics="true" pageId="15" pageNumber="210">et al.</emphasis>
, 1998
</bibRefCitation>
, reported as
<taxonomicName id="A6BC4D7E4D326B318505FE3743A19C4E" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[829,963,504,526]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D326B318505FE3743A19C4E" box="[829,963,504,526]" italics="true" pageId="15" pageNumber="210">O. dentatus</emphasis>
</taxonomicName>
). In
<collectingCountry id="19AB766D4D326B318230FE3644119C4D" box="[1032,1139,504,526]" name="Malaysia" pageId="15" pageNumber="210">Malaysia</collectingCountry>
, it is a common prey of homalopsine snakes such as
<taxonomicName id="A6BC4D7E4D326B318295FDD945049C6F" box="[1197,1382,535,556]" class="Reptilia" family="Homalopsidae" genus="Bitia" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="hydroides">
<emphasis id="53C8EAEF4D326B318295FDD945049C6F" box="[1197,1382,535,556]" italics="true" pageId="15" pageNumber="210">Bitia hydroides</emphasis>
</taxonomicName>
(see
<bibRefCitation id="052D4B0C4D326B318505FDF8441E9C08" author="Jayne BC &amp; Ward TJ &amp; Voris HK" box="[829,1148,565,587]" pageId="15" pageNumber="210" pagination="800 - 808" refId="ref12768" refString="Jayne BC, Ward TJ, Voris HK. 1995. Morphology, reproduction and diet of the marine homalopsine snake Bitia hydroides in Peninsula Malaysia. Copeia 1995: 800 - 808." type="journal article" year="1995">Jayne, Ward &amp; Voris, 1995</bibRefCitation>
, reported as
<taxonomicName id="A6BC4D7E4D326B31832EFDF845FE9C09" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[1302,1436,565,587]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D326B31832EFDF845FE9C09" box="[1302,1436,565,587]" italics="true" pageId="15" pageNumber="210">O. dentatus</emphasis>
</taxonomicName>
) and
<taxonomicName id="A6BC4D7E4D326B31854DFD9A442F9C2A" baseAuthorityName="Schneider" baseAuthorityYear="1799" box="[885,1101,596,617]" class="Reptilia" family="Homalopsidae" genus="Cerberus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="rynchops">
<emphasis id="53C8EAEF4D326B31854DFD9A442F9C2A" box="[885,1101,596,617]" italics="true" pageId="15" pageNumber="210">Cerberus rynchops</emphasis>
</taxonomicName>
(see
<bibRefCitation id="052D4B0C4D326B3182AAFD9A43149CCB" author="Jayne BC &amp; Voris HK &amp; Heang KB" pageId="15" pageNumber="210" pagination="1 - 15" refId="ref12728" refString="Jayne BC, Voris HK, Heang KB. 1988. Diet, feeding behavior, growth and numbers of a population of Cerberus rynchops (Serpentes: Homalopsinae) in Malaysia. Fieldiana 50: 1 - 15." type="journal article" year="1988">Jayne, Voris &amp; Heang, 1988</bibRefCitation>
, reported as
<taxonomicName id="A6BC4D7E4D326B318234FDBD44F09CC4" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[1036,1170,626,648]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D326B318234FDBD44F09CC4" box="[1036,1170,626,648]" italics="true" pageId="15" pageNumber="210">O. dentatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="610336FD4D326B318505FD6144B69B04" blockId="15.[829,1445,687,1351]" pageId="15" pageNumber="210">
<emphasis id="53C8EAEF4D326B318505FD6144D29C86" box="[829,1200,687,709]" italics="true" pageId="15" pageNumber="210">
Remarks:
<taxonomicName id="A6BC4D7E4D326B31858FFD7E44D29C86" authorityName="Cantor, 1849:" authorityYear="1170" box="[951,1200,688,709]" class="Actinopterygii" family="Gobiidae" genus="Apocryptes" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="nexipinnis">Apocryptes nexipinnis</taxonomicName>
</emphasis>
, described by
<bibRefCitation id="052D4B0C4D326B31836DFD7E43E59CA7" author="Cantor T" pageId="15" pageNumber="210" pagination="983 - 1443" refId="ref12023" refString="Cantor T. 1849. Catalog of Malayan fishes. Journal of the Asiatic Society of Bengal 18: 983 - 1443." type="journal article" year="1849">Cantor (1849)</bibRefCitation>
, was classified in the genus
<taxonomicName id="A6BC4D7E4D326B3182C8FD0045079CA0" authorityName="NEXIPINNIS (CANTOR" authorityYear="1849" box="[1264,1381,718,739]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="53C8EAEF4D326B3182C8FD0045079CA0" box="[1264,1381,718,739]" italics="true" pageId="15" pageNumber="210">Oxuderces</emphasis>
</taxonomicName>
, and synonymized with
<taxonomicName id="A6BC4D7E4D326B318222FD2344C29D41" authorityName="Eydoux &amp; Souleyet" authorityYear="1848" box="[1050,1184,749,770]" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="dentatus">
<emphasis id="53C8EAEF4D326B318222FD2344C29D41" box="[1050,1184,749,770]" italics="true" pageId="15" pageNumber="210">O. dentatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="052D4B0C4D326B3182E8FD2345139D40" author="Murdy EO" box="[1232,1393,749,771]" pageId="15" pageNumber="210" pagination="1 - 93" refId="ref12918" refString="Murdy EO. 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Records of the Australian Museum 11 (Supplement): 1 - 93." type="journal article" year="1989">Murdy (1989)</bibRefCitation>
. We resurrect this name from synonymy as
<taxonomicName id="A6BC4D7E4D326B318317FCC543D19D7C" baseAuthorityName="Cantor" baseAuthorityYear="1849" class="Actinopterygii" family="Gobiidae" genus="Oxuderces" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="210" phylum="Chordata" rank="species" species="nexipinnis">
<emphasis id="53C8EAEF4D326B318317FCC543D19D7C" italics="true" pageId="15" pageNumber="210">Oxuderces nexipinnis</emphasis>
</taxonomicName>
. The BMNH
<typeStatus id="BE07885F4D326B318259FCE444A79D7C" box="[1121,1221,810,831]" pageId="15" pageNumber="210" type="syntype">syntypes</typeStatus>
are the left sides (heads and bodies) of two individuals that have been dried, filleted and stuffed with cotton wool (
<figureCitation id="F9872A784D326B31837CFCA945FA9D3E" box="[1348,1432,871,893]" captionStart="Figure 14" captionStartId="18.[146,225,1675,1694]" captionTargetBox="[153,1415,198,1641]" captionTargetId="figure-93@18.[153,1415,198,1642]" captionTargetPageId="18" captionText="Figure 14. Apocryptes nexipinnis Cantor, 1849, syntypes, BMNH1860.3.19.568-69 (2), dried, filleted and stuffed with cotton wool. Anterior is to the left. Top: the larger of the two syntypes, approximately 67.8 mm SL. Middle: the same specimen as the top, rotated on the horizontal axis. Bottom: the smaller of the two syntypes, approximately 65 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5710977" httpUri="https://zenodo.org/record/5710977/files/figure.png" pageId="15" pageNumber="210">Fig. 14</figureCitation>
).
<bibRefCitation id="052D4B0C4D326B318505FC4843889DDF" author="Cantor T" box="[829,1002,902,924]" pageId="15" pageNumber="210" pagination="983 - 1443" refId="ref12023" refString="Cantor T. 1849. Catalog of Malayan fishes. Journal of the Asiatic Society of Bengal 18: 983 - 1443." type="journal article" year="1849">Cantor (1849)</bibRefCitation>
did not indicate the number of specimens he had on hand and reported the total length of the specimen (or specimens) on which he based his description as 3 3/
<quantity id="A6449B184D326B3182BCFC2F44819DB4" box="[1156,1251,993,1015]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="15" pageNumber="210" unit="in" value="8.0">8 inches</quantity>
, or
<quantity id="A6449B184D326B318328FC2F45119DB4" box="[1296,1395,993,1015]" metricMagnitude="-2" metricUnit="m" metricValue="8.57" pageId="15" pageNumber="210" unit="mm" value="85.7">85.7 mm</quantity>
TL. He further indicated that the caudal fin comprised 1/5 of the TL, which would mean that the specimen (or specimens) was approximately
<quantity id="A6449B184D326B3182EAFBF345439A11" box="[1234,1313,1085,1107]" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="15" pageNumber="210" unit="mm" value="68.0">68 mm</quantity>
SL. The BMNH
<typeStatus id="BE07885F4D326B3185AFFB9243999A32" box="[919,1019,1116,1137]" pageId="15" pageNumber="210" type="syntype">syntypes</typeStatus>
measure approximately
<quantity id="A6449B184D326B318320FB95451E9A32" box="[1304,1404,1115,1137]" metricMagnitude="-2" metricUnit="m" metricValue="6.779999999999999" pageId="15" pageNumber="210" unit="mm" value="67.8">67.8 mm</quantity>
SL (possibly a male) and
<quantity id="A6449B184D326B318279FBB444F29ACC" box="[1089,1168,1146,1168]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="210" unit="mm" value="65.0">65 mm</quantity>
SL (possibly a female). Although the larger specimen is closer in size to that given by
<bibRefCitation id="052D4B0C4D326B318594FB79442D9A8E" author="Cantor T" box="[940,1103,1207,1229]" pageId="15" pageNumber="210" pagination="983 - 1443" refId="ref12023" refString="Cantor T. 1849. Catalog of Malayan fishes. Journal of the Asiatic Society of Bengal 18: 983 - 1443." type="journal article" year="1849">Cantor (1849)</bibRefCitation>
, we do not designate it as a
<typeStatus id="BE07885F4D326B318505FB1843C19AAF" box="[829,931,1238,1260]" pageId="15" pageNumber="210" type="lectotype">lectotype</typeStatus>
because both specimens are extremely distorted, as well as incomplete, and we cannot identify either with certainty as the primary specimen on which the description was based.
</paragraph>
</subSubSection>
</treatment>
</document>
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