155 lines
21 KiB
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155 lines
21 KiB
XML
<document id="E293FB0F03755E5D6DCA31CF87ECF248" ID-DOI="10.5281/zenodo.12649284" ID-ISSN="1407-8953" ID-Zenodo-Dep="12649284" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720074198029" checkinUser="felipe" docAuthor="Shavrin, Alexey V." docDate="2020" docId="E24FE440320DFFC41D693263FEF2FBC9" docLanguage="en" docName="ActaBiolUnivDaugavp.20.1.29-33.pdf" docOrigin="Acta Biologica Universitatis Daugavpiliensis 20 (1)" docStyle="DocumentStyle:DD7D35B4ACFEF3C20AEB77B8656EEADF.5:ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleId="DD7D35B4ACFEF3C20AEB77B8656EEADF" docStyleName="ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleVersion="5" docTitle="Deliphrosoma frischi SHAVRIN 2011" docType="treatment" docVersion="2" lastPageNumber="32" masterDocId="1E769C38320CFFC71D323634FFA7FF82" masterDocTitle="New Faunistic Data On Eight Species Of Deliphrosoma Reitter, 1909, With A Redescription Of D. Frischi Shavrin, 2011 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="33" masterPageNumber="29" pageNumber="30" updateTime="1720087858401" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E3258665E35565DA243C1162140FF5C9">New Faunistic Data On Eight Species Of Deliphrosoma Reitter, 1909, With A Redescription Of D. Frischi Shavrin, 2011 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="93432C9F46E346E20F215E636D433B47">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="597ACA05A7F94DB518C68FD341BEA762">Acta Biologica Universitatis Daugavpiliensis</mods:title>
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<mods:date id="A64EA15BC38A1D9CD470F8F90B288E0D">2020</mods:date>
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<treatmentCitation id="EB477347320DFFC61D693263FDA5FBED" author="Shavrin A. V." box="[91,514,1111,1135]" page="35" pageId="1" pageNumber="30" year="2011">
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<taxonomicName id="ADE62ED5320DFFC61D693263FDA5FBED" ID-CoL="34GMH" authority="Shavrin, 2011: 35" authorityName="SHAVRIN" authorityPageNumber="35" authorityYear="2011" box="[91,514,1111,1135]" class="Insecta" family="Staphylinidae" genus="Deliphrosoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="frischi">
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<emphasis id="58928944320DFFC61D693263FE9AFBED" box="[91,317,1111,1135]" italics="true" pageId="1" pageNumber="30">Deliphrosoma frischi</emphasis>
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<bibRefCitation id="0E7728A7320DFFC61C763263FDA5FBED" author="Shavrin A. V." box="[324,514,1111,1135]" pageId="1" pageNumber="30" pagination="35 - 37" refId="ref2430" refString="Shavrin A. V. 2011. New species and new records of Deliphrosoma Reitter, 1909 (Coleoptera: Staphylinidae: Omaliinae) from Iran with note on the taxonomic status of Geodromicus libanicus Coiffait, 1985. Caucasian Entomological Bulletin, 7 (1): 35 - 37." type="journal article" year="2011">Shavrin, 2011: 35</bibRefCitation>
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<subSubSection id="22FC06DD320DFFC61D6932A3FE0CFA8D" pageId="1" pageNumber="30" type="materials_examined">
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<paragraph id="6A595556320DFFC61D6932A3FE0CFA8D" blockId="1.[91,602,1175,1295]" pageId="1" pageNumber="30">
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<emphasis id="58928944320DFFC61D6932A3FF6CFB2D" bold="true" box="[91,203,1175,1199]" pageId="1" pageNumber="30">Material.</emphasis>
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<materialsCitation id="DA8E5F0B320DFFC61DE532A3FE00FA8D" collectingDate="2007-11-05" collectionCode="MNHUB" collectorName="S Semirom" country="Iran" elevation="2810" location="Esfahan Province" longitude="31.013056" pageId="1" pageNumber="30" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Esfahan">
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<specimenCount id="7CE09EDF320DFFC61DE532A3FE8FFB2D" box="[215,296,1175,1199]" count="1" pageId="1" pageNumber="30" type="male">1 male</specimenCount>
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,
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<specimenCount id="7CE09EDF320DFFC61C0B32A3FE08FB2D" box="[313,431,1175,1199]" count="2" pageId="1" pageNumber="30" type="female">2 females</specimenCount>
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: ‘
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<collectingCountry id="12F115C6320DFFC61CF032A3FDA9FB2D" box="[450,526,1175,1199]" name="Iran" pageId="1" pageNumber="30">IRAN</collectingCountry>
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,
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<collectingRegion id="A8229BB4320DFFC61F2C32A3FF4FFB4D" country="Iran" name="Esfahan" pageId="1" pageNumber="30">Esfahan Province</collectingRegion>
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<collectorName id="C7133080320DFFC61DCE3283FEDEFB4D" box="[252,377,1207,1231]" pageId="1" pageNumber="30">S Semirom</collectorName>
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: Komeh |
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<elevation id="E1CBB265320DFFC61CD53283FD96FB4D" box="[487,561,1207,1231]" metricMagnitude="3" metricUnit="m" metricValue="2.81" pageId="1" pageNumber="30" unit="m" value="2810.0">2810m</elevation>
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</quantity>
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| N
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<geoCoordinate id="0FD23391320DFFC61D6E32E3FF40FB6C" box="[92,231,1239,1263]" degrees="31" direction="east" minutes="00" orientation="longitude" pageId="1" pageNumber="30" precision="15" seconds="47" value="31.013056">31°00’47’’ E</geoCoordinate>
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051°35’28’’ |
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<date id="1E587396320DFFC61CB832E3FDA3FB6D" box="[394,516,1239,1263]" pageId="1" pageNumber="30" value="2007-11-05">
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<collectingDate id="0E1C8A7E320DFFC61CB832E3FDA3FB6D" box="[394,516,1239,1263]" pageId="1" pageNumber="30" value="2007-11-05">11.05.2007</collectingDate>
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</date>
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, lg. [J.] Frisch & [S.] Serri’ (
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<collectionCode id="0CF7CD93320DFFC61C0432C3FE38FA8D" box="[310,415,1271,1295]" pageId="1" pageNumber="30">MNHUB</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<subSubSection id="22FC06DD320DFFC41D693303FDFDFE2E" lastPageId="3" lastPageNumber="32" pageId="1" pageNumber="30" type="description">
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<paragraph id="6A595556320DFFC61D693303FCC1FEC9" blockId="1.[89,603,1335,1679]" lastBlockId="1.[637,1149,116,332]" pageId="1" pageNumber="30">
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<emphasis id="58928944320DFFC61D693303FEA7FACD" bold="true" box="[91,256,1335,1359]" pageId="1" pageNumber="30">Redescription.</emphasis>
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Measurements (in mm): maximum width of head including eyes: 0.70–0.82; length of head (from base of labrum to neck constriction along head midline): 0.50–0.60; length of antenna (
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<typeStatus id="B55DEBF4320DFFC61C2C3383FE25FA4D" box="[286,386,1463,1487]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
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): 1.45; ocular length (longitudinal): 0.20–0.22; length of pronotum: 0.60–0.77; maximum width of pronotum: 0.95– 1.11; sutural length of elytra (length of elytra from the apex of scutellum to the posterior margin of sutural angle): 1.55–2.22; maximum width of elytra: 1.30–1.45; maximum width of abdomen: 1.00–1.37; length of metatibia (averaged): 1.02; length of metatarsus (averaged): 0.59 (metatarsomeres 1–4: 0.37; metatarsomere 5: 0.22); length of aedeagus (from base of the median lobe to apex of parameres): 0.80–0.82; total length (from anterior margin of clypeus to apex of abdomen): 3.90 (
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<typeStatus id="B55DEBF4320DFFC61F893707FCBBFEC9" box="[699,796,307,331]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
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)–4.76.
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</paragraph>
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<paragraph id="6A595556320DFFC61F4D3740FC47FA0E" blockId="1.[637,1152,372,1420]" pageId="1" pageNumber="30">
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Body, antenna and femora yellow-brown to brown (
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<typeStatus id="B55DEBF4320DFFC61FFF37A0FC8AFE2E" box="[717,813,404,428]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
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), some specimens with distinctly paler basal margins of pronotum and entire elytra (Figs. 1–2); mouthparts, tibiae and tarsi yellow. Body shiny; head with distinct transverse microsculpture on apical portion, with very dense isodiametric meshes between base of antenna and apical margins of eyes, without (
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<typeStatus id="B55DEBF4320DFFC61EB13460FC46FDEE" box="[899,993,596,620]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
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) or with sparse microreticulation in middle and basal portions; neck with distinct, transverse meshes; pronotum without or with indistinct transverse microreticulation; scutellum with transverse microsculpture; abdomen with distinct isodiametric sculpture. Head with irregular and very sparse punctation, larger and denser in middle portion; neck with fine, sparse punctation; pronotum with irregular punctation, sparser and finer in middle portion (
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<typeStatus id="B55DEBF4320DFFC61FB435A0FD4EFC2E" box="[646,745,916,940]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
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with larger and denser punctures); scutellum without or with several fine punctures in apical portion; elytra with moderately sparse and large punctation, somewhat deeper than that on pronotum, denser and deeper in prescutellar and finer in apical portions, forming unclear six very vague and tangled longitudinal rows of punctures; abdomen without visible punctation. Body glabrous, apical and infraorbital portions of head with long semierect setae, lateral and middle parts of pronotum with three-four long setae; elytra with very sparse and moderately long setae in lateral and middle portions; surface of abdominal tergites with sparse, short setation, and long one-three additional setae on each paratergite. Habitus as in Figs. 1–2.
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</paragraph>
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<paragraph id="6A595556320DFFC51F4D3380FE62FC8E" blockId="1.[637,1149,1460,1676]" lastBlockId="2.[134,647,116,780]" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30">
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Head 1.3–1.4 times as broad as long; middle portion slightly and widely elevated, with moderately deep depressions between supra-antennal protuberances and anterior margins of eyes, middle portion between anteocellar foveae with narrow superficial impression, indistinct in
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<typeStatus id="B55DEBF4320DFFC619273060FBDFF9EE" box="[1045,1144,1620,1644]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
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; anteocellar foveae deep and moderately long, reaching level of middle length of eyes; postocular ridge indistinct, obtuse, with surface between postocular ridge and posterior margin of eyes as long as two nearest ommatidia; basal portion behind ocelli with distinct and moderately deep impression; anterior portion between antennal insertion and anterior margin of eye with semicircular, deep notch. Eyes large, convex. Ocelli large, situated behind level of postocular ridges; distance between ocelli about as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palpus about twice as long as preceding segment. Antenna reaching middle of elytra when reclined; basal antennomere suboval, about three times as long as broad, antennomere 2 distinctly narrower than basal antennomere, 3 about 1.3 times as long as and slightly narrower than 2, 4–7 distinctly shorter than 3, 8 slightly shorter than 7, 9–10 slightly shorter and broader than 8, apical antennomere about 1.3 times as long as 10.
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</paragraph>
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<paragraph id="6A595556320EFFC51DBA3500FB0CFF6E" blockId="2.[135,647,819,908]" lastBlockId="2.[684,1195,116,236]" pageId="2" pageNumber="31">Pronotum slightly convex, 1.4–1.5 times as broad as long, 1.3 times as broad as head, widest in anterior portion, widely rounded anteriad and gradually narrowed posteriad toward rounded hind angles; apical and posterior margins somewhat straight; lateral portions narrow, each with moderately deep, oval pit slightly above middle.</paragraph>
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<paragraph id="6A595556320EFFC51F9E3720FC06FE4E" blockId="2.[684,1195,276,460]" pageId="2" pageNumber="31">Elytra relatively convex, longer than wide and significantly longer than twice as long as pronotum, slightly widened toward apical third, reaching basal margin of abdominal tergites IV or V; apical margins straight or rounded. Hind wings fully developed.</paragraph>
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<paragraph id="6A595556320EFFC51F9E37C0FB0DFDAE" blockId="2.[682,1194,500,556]" pageId="2" pageNumber="31">Legs long, slender; tibiae covered by dense, long setae; metatarsus 1.7 times as long as metatibia.</paragraph>
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<paragraph id="6A595556320EFFC51F9E3460FBE9FD4E" blockId="2.[682,1194,596,716]" pageId="2" pageNumber="31">Abdomen distinctly narrower than elytra, with two small, round tomentose spots in the middle of abdominal tergite V; apical margin of abdominal tergite VII with narrow palisade fringe.</paragraph>
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<paragraph id="6A595556320EFFC51F9E34C0FB0DFC0E" blockId="2.[682,1195,756,908]" pageId="2" pageNumber="31">Male (Fig. 1). Protarsomeres 1–4 very wide. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII widely concave.Adeagus with wide basal portion, gradually narrowing toward trunctated apex (Fig. 3);</paragraph>
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<paragraph id="6A595556320EFFC41DB83043FDF7FEAE" blockId="2.[138,1027,1655,1679]" lastBlockId="3.[91,601,116,300]" lastPageId="3" lastPageNumber="32" pageId="2" pageNumber="31">
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Figs. 1–2. Habitus of
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<taxonomicName id="ADE62ED5320EFFC51C5D3043FDF6F90D" authorityName="SHAVRIN" authorityYear="2011" box="[367,593,1655,1679]" class="Insecta" family="Staphylinidae" genus="Deliphrosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="frischi">
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<emphasis id="58928944320EFFC51C5D3043FDF6F90D" box="[367,593,1655,1679]" italics="true" pageId="2" pageNumber="31">Deliphrosoma frischi</emphasis>
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</taxonomicName>
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: 1–male, 2–female. Scale bar: 1.00 mm. parameres asymmetric (right paramere slightly longer than left), narrow, exceeding apex of median lobe, with rounded apical portions and two short apical setae; internal sac narrow and moderately long, with a field of large spines in middle. Lateral aspect of aedeagus as in Fig. 4.
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</paragraph>
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<paragraph id="6A595556320FFFC41D693760FDFDFE2E" blockId="3.[91,602,340,428]" pageId="3" pageNumber="32">Female (Fig. 2). Protarsomeres 1–4 narrow. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII round.</paragraph>
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</subSubSection>
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<subSubSection id="22FC06DD320FFFC41D6937E0FEF2FBC9" pageId="3" pageNumber="32" type="discussion">
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<paragraph id="6A595556320FFFC41D6937E0FEF2FBC9" blockId="3.[89,603,468,1100]" pageId="3" pageNumber="32">
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<emphasis id="58928944320FFFC41D6937E0FF6EFE6E" bold="true" box="[91,201,468,492]" pageId="3" pageNumber="32">Remarks.</emphasis>
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The original description of
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<taxonomicName id="ADE62ED5320FFFC41CC637E0FF4DFD8E" authority="Shavrin, 2011" authorityName="SHAVRIN" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Deliphrosoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="frischi">
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<emphasis id="58928944320FFFC41CC637E0FDA0FE69" box="[500,519,468,491]" italics="true" pageId="3" pageNumber="32">D</emphasis>
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.
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<emphasis id="58928944320FFFC41F2737E0FDFEFE6E" box="[533,601,468,492]" italics="true" pageId="3" pageNumber="32">frischi</emphasis>
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<bibRefCitation id="0E7728A7320FFFC41D6937C0FF4DFD8E" author="Shavrin A. V." box="[91,234,500,524]" pageId="3" pageNumber="32" pagination="35 - 37" refId="ref2430" refString="Shavrin A. V. 2011. New species and new records of Deliphrosoma Reitter, 1909 (Coleoptera: Staphylinidae: Omaliinae) from Iran with note on the taxonomic status of Geodromicus libanicus Coiffait, 1985. Caucasian Entomological Bulletin, 7 (1): 35 - 37." type="journal article" year="2011">Shavrin, 2011</bibRefCitation>
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||
</taxonomicName>
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||
was based on the study of the single specimen with the brown body and dense, moderately deep punctation of the pronotum. Shapes of aedeagus and internal sac of the
|
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<typeStatus id="B55DEBF4320FFFC41CF13460FD87FDEE" box="[451,544,596,620]" pageId="3" pageNumber="32" type="holotype">holotype</typeStatus>
|
||
were dramatically deformed and flattened (see Fig.
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||
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<bibRefCitation id="0E7728A7320FFFC41D6834A0FF4DFD2E" author="Shavrin A. V." box="[90,234,660,684]" pageId="3" pageNumber="32" pagination="35 - 37" refId="ref2430" refString="Shavrin A. V. 2011. New species and new records of Deliphrosoma Reitter, 1909 (Coleoptera: Staphylinidae: Omaliinae) from Iran with note on the taxonomic status of Geodromicus libanicus Coiffait, 1985. Caucasian Entomological Bulletin, 7 (1): 35 - 37." type="journal article" year="2011">Shavrin 2011</bibRefCitation>
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). Some time after this description, I received
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<specimenCount id="7CE09EDF320FFFC41D8B3480FEFEFD4E" box="[185,345,692,716]" count="3" pageId="3" pageNumber="32" type="immature">three immature</specimenCount>
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specimens of
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<taxonomicName id="ADE62ED5320FFFC41CC23480FF34FD69" authorityName="REITTER" authorityYear="1909" class="Insecta" family="Staphylinidae" genus="Deliphrosoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="58928944320FFFC41CC23480FF34FD69" italics="true" pageId="3" pageNumber="32">Deliphrosoma</emphasis>
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</taxonomicName>
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with the same label`s data as well as the
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<typeStatus id="B55DEBF4320FFFC41D6934C0FF1CFC8E" box="[91,187,756,780]" pageId="3" pageNumber="32" type="holotype">holotype</typeStatus>
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. Despite the paler coloration of the body and finer, sparser punctation of the pronotum (Figs. 1–2), these specimens are morphologically conspecific with
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<taxonomicName id="ADE62ED5320FFFC41C2A3560FE25FCEE" authorityName="SHAVRIN" authorityYear="2011" box="[280,386,852,876]" class="Insecta" family="Staphylinidae" genus="Deliphrosoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="frischi">
|
||
<emphasis id="58928944320FFFC41C2A3560FE8CFCE9" box="[280,299,852,875]" italics="true" pageId="3" pageNumber="32">D</emphasis>
|
||
.
|
||
<emphasis id="58928944320FFFC41C0F3560FE25FCEE" box="[317,386,852,876]" italics="true" pageId="3" pageNumber="32">frischi</emphasis>
|
||
</taxonomicName>
|
||
. The male (Fig. 1) have well sclerotized aedeagus and internal sac, perfectly visible in parameral and lateral projections (Figs. 3–4). Thus, I decide to modify and correct the original description of the species. In general,
|
||
<taxonomicName id="ADE62ED5320FFFC41D8435C0FEBDFB8E" authorityName="SHAVRIN" authorityYear="2011" box="[182,282,1012,1036]" class="Insecta" family="Staphylinidae" genus="Deliphrosoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="frischi">
|
||
<emphasis id="58928944320FFFC41D8435C0FEBDFB8E" box="[182,282,1012,1036]" italics="true" pageId="3" pageNumber="32">D. frischi</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the remaining species of the genus by the shape of asymmetric parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |