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<mods:title id="08AC720A73631F8BF0252BCF4C05135F">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part IX. Lanzatus, Orthochirus, and Somalicharmus (Buthidae), with description of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n.</mods:title>
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<paragraph id="533E36BDFFFDE165ED4BFD65FD88A60A" blockId="3.[303,690,690,805]" box="[366,627,751,774]" pageId="3" pageNumber="2">
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<taxonomicName id="94814D3EFFFDE165ED4BFD65FD88A60A" ID-CoL="62XN3" authority="Kovarik, 2001" authorityName="Kovarik" authorityYear="2001" box="[366,627,751,774]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="61F5EAAFFFFDE165ED4BFD65FE28A60A" bold="true" box="[366,467,751,774]" italics="true" pageId="3" pageNumber="2">Lanzatus</emphasis>
Kovařík, 2001
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<subSubSection id="1B9B6536FFFDE165ED15FC84FD4AA629" box="[304,689,782,805]" pageId="3" pageNumber="2" type="description">
<paragraph id="533E36BDFFFDE165ED15FC84FD4AA629" blockId="3.[303,690,690,805]" box="[304,689,782,805]" pageId="3" pageNumber="2">
(
<figureCitation id="CBBA2A38FFFDE165ED1DFC84FE56A629" box="[312,429,782,805]" captionStart-0="Figures 16" captionStart-1="Figures 712" captionStart-2="Figures 1314" captionStart-3="Figures 1529" captionStart-4="Figures 3037" captionStart-5="Figures 3840" captionStartId-0="4.[192,278,1758,1781]" captionStartId-1="5.[192,278,1446,1469]" captionStartId-2="6.[192,278,1016,1039]" captionStartId-3="7.[192,278,1567,1590]" captionStartId-4="8.[192,278,1273,1296]" captionStartId-5="9.[192,278,1830,1853]" captionTargetBox-0="[198,1430,200,1696]" captionTargetBox-1="[191,1437,191,1417]" captionTargetBox-2="[191,1437,191,986]" captionTargetBox-3="[192,1439,175,1532]" captionTargetBox-4="[198,1431,175,1233]" captionTargetBox-5="[191,1440,204,1801]" captionTargetId-0="figure-16@4.[192,1439,195,1728]" captionTargetId-1="figure-6@5.[192,1437,192,1416]" captionTargetId-2="figure-16@6.[192,1437,192,986]" captionTargetId-3="figure-6@7.[192,1439,168,1538]" captionTargetId-4="figure-38@8.[192,1437,168,1243]" captionTargetId-5="figure-28@9.[192,1439,205,1800]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="Figures 16: Lanzatus. Figures 14. L. somalilandus sp. n. Figures 12. Female paratype in dorsal (1) and ventral (2) views. Figures 34. Male holotype in dorsal (3) and ventral (4) views. Figures 56. L. somalicus, male paratype in dorsal (5) and ventral (6) views. Scale bar: 10 mm." captionText-1="Figures 712: Lanzatus, carapace and tergites IIII (7, 9, 11), and coxosternal area and sternites IIIIV (8, 10, 12). Figures 7 10. L. somalilandus sp. n., female paratype (78) and male holotype (910). Figures 1112. L. somalicus, male paratype." captionText-2="Figures 1314: L. somalilandus sp. n., right chelicera of female paratype, dorsal (13) and ventral (14). Composite white light/ UV fluorescence images. Scale bar: 200 µm." captionText-3="Figures 1529: Lanzatus. Figures 1528. L. somalilandus sp. n. Figures 1527. Female paratype. Figures 1522. Pedipalp segments, movable finger dentition (15), chela dorsal (16), external (17) and ventral (18), patella dorsal (19), external (20) and ventral (21), trochanter and femur dorsal (22). Trichobothrial pattern indicated in Figures 1720 and 22. Figures 2326. Left legs IIV, retrolateral aspect. Figures 2729. Telson lateral. Figures 2728. L. somalilandus sp. n., female paratype (27) and male holotype (28). Figure 29. L. somalicus, male paratype." captionText-4="Figures 3037: Lanzatus, metasoma and telson. Figures 3031. L. somalilandus sp. n., male holotype, ventral (30), and lateral (31) views. Figures 3234. L. somalicus, male paratype, lateral (32), ventral (33), and dorsal (34) views. Figures 3537. L. somalilandus sp. n., female paratype, lateral (35), ventral (36), and dorsal (37) views." captionText-5="Figures 3840: L. somalilandus sp. n., male (38) and female (39) in vivo habitus and the type locality (40)." pageId="3" pageNumber="2">Figs. 140</figureCitation>
,
<figureCitation id="CBBA2A38FFFDE165ED92FC84FDECA629" box="[439,535,782,805]" captionStart="Figures 123128" captionStartId="28.[192,278,1758,1781]" captionTargetBox="[209,1417,187,1729]" captionTargetId="figure-115@28.[210,1416,188,1728]" captionTargetPageId="28" captionText="Figures 123128: Figures 123126: Diagrams of cheliceral finger dentition. Figures 123124. Lanzatus somalilandus sp. n., female paratype, dorsal (123) and ventral (124) views, showing typical pattern of buthid dentition. Reconstructed from specimen with worn teeth (Figs. 1314). Figures 125125. Somalicharmus whitmanae, male from locality 16EA, dorsal (125) and ventral (126) views, showing pattern of dentition highly aberrant for a buthid. Figures 127128. Somalicharmus whitmanae, left chelicera of male from locality 16EA, dorsal (127) and ventral (128) views. Composite white light/ UV fluorescence images. Scale bar: 500 µm." pageId="3" pageNumber="2">123124</figureCitation>
,
<figureCitation id="CBBA2A38FFFDE165EE07FC84FDB7A629" box="[546,588,782,805]" captionStart="Figure 155" captionStartId="35.[192,268,1422,1445]" captionTargetBox="[196,1434,196,1389]" captionTargetId="figure-260@35.[192,1437,191,1392]" captionTargetPageId="35" captionText="Figure 155: Map showing confirmed distribution of Lanzatus, Orthochirus, and Somalicharmus in Horn of Africa. Most points indicate sites sampled during 20112016 expeditions, except for the localities of Lanzatus somalicus in Somalia, and of Orthochirus sp. in Djibouti (the map indicates with arbitrary plotted symbol for two citations: a juvenile cited by Werner (1916: 81) with imprecise coordinates, and of a specimen cited by Lourenço &amp; Leguin (2011: 4, 15, figs. 5, 39); see also comments under Orthochirus in Systematics section." pageId="3" pageNumber="2">155</figureCitation>
,
<tableCitation id="1E030306FFFDE165EE72FC84FD5CA629" box="[599,679,782,805]" captionStart="Table 1" captionStartId="19.[192,257,836,859]" captionText="Table 1: Comparative measurements of adults of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="3" pageNumber="2">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1B9B6536FFFDE165ECE5FCC2FD69A17F" pageId="3" pageNumber="2" type="reference_group">
<paragraph id="533E36BDFFFDE165ECE5FCC2FD78A6D6" blockId="3.[192,802,840,1139]" pageId="3" pageNumber="2">
<taxonomicName id="94814D3EFFFDE165ECE5FCC2FDC0A653" ID-CoL="62XN3" authority="Kovarik, 2001: 41 - 44" authorityName="Kovarik" authorityPageNumber="41 - 44" authorityYear="2001" box="[192,571,840,863]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="61F5EAAFFFFDE165ECE5FCC2FEDBA653" box="[192,288,840,863]" italics="true" pageId="3" pageNumber="2">Lanzatus</emphasis>
Kovařík, 2001: 4144
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, figs. 17; Fet &amp; Soleglad, 2005: 11; Fet et al., 2005: 3, 1112, 20, 2225,
<tableCitation id="1E030306FFFDE165ED62FC0FFE69A690" box="[327,402,901,924]" captionStart="Table 1" captionStartId="19.[192,257,836,859]" captionText="Table 1: Comparative measurements of adults of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="3" pageNumber="2">Tab. 1</tableCitation>
,
<figureCitation id="CBBA2A38FFFDE165ED83FC0FFDDEA690" box="[422,549,901,924]" captionStart="Figures 1529" captionStartId="7.[192,278,1567,1590]" captionTargetBox="[192,1439,175,1532]" captionTargetId="figure-6@7.[192,1439,168,1538]" captionTargetPageId="7" captionText="Figures 1529: Lanzatus. Figures 1528. L. somalilandus sp. n. Figures 1527. Female paratype. Figures 1522. Pedipalp segments, movable finger dentition (15), chela dorsal (16), external (17) and ventral (18), patella dorsal (19), external (20) and ventral (21), trochanter and femur dorsal (22). Trichobothrial pattern indicated in Figures 1720 and 22. Figures 2326. Left legs IIV, retrolateral aspect. Figures 2729. Telson lateral. Figures 2728. L. somalilandus sp. n., female paratype (27) and male holotype (28). Figure 29. L. somalicus, male paratype." pageId="3" pageNumber="2">Fig. 2325</figureCitation>
; Prendini &amp; Wheeler, 2005: 462, 481; Dupré, 2007: 7, 13, 16;
<bibRefCitation id="37104B4CFFFDE165EE8BFC2EFE7DA6D6" author="KOVARIK" pageId="3" pageNumber="2" pagination="201 - 209" refId="ref13435" refString="KOVARIK, F., M. E. SOLEGLAD &amp; V. FET. 2007. A new species of scorpion in the &quot; Charmus &quot; group from India (Scorpiones: Buthidae). Boletin Sociedad Entomologica Aragonesa, 40: 201 - 209." type="journal article" year="2007">Kovařík et al., 2007: 207</bibRefCitation>
;
<bibRefCitation id="37104B4CFFFDE165EDB4FC49FDADA6D6" author="KOVARIK" box="[401,598,963,986]" pageId="3" pageNumber="2" pagination="1 - 12" refId="ref13482" refString="KOVARIK, F., F. STAHLAVSKY, T. KORIN- KOVA, J. KRAL &amp; T. VAN DER ENDE. 2009. Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1 - 12." type="journal article" year="2009">Kovařík, 2009: 23</bibRefCitation>
, 31.
</paragraph>
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=
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<taxonomicName id="94814D3EFFFDE165ECFCFC6BFD7BA6F4" authority="Lourenco, 2001 a: 16 - 18" authorityName="Lourenco" authorityPageNumber="16 - 18" authorityYear="2001" box="[217,640,993,1016]" class="Arachnida" family="Buthidae" genus="Sabinebuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="61F5EAAFFFFDE165ECFCFC6BFE92A6F4" box="[217,361,993,1016]" italics="true" pageId="3" pageNumber="2">Sabinebuthus</emphasis>
<bibRefCitation id="37104B4CFFFDE165ED57FC6BFD7BA6F4" author="LOURENCO" box="[370,640,993,1016]" pageId="3" pageNumber="2" pagination="15 - 20" refId="ref13852" refString="LOURENCO, W. R. 2001 a. Un nouveau genre de Buthidae, probable vicariant geographique d' Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço, 2001a: 1618</bibRefCitation>
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, figs. 15; Fet &amp; Soleglad, 2005: 4, 11; Fet et al., 2005: 3, 11, 20, 2223,
<tableCitation id="1E030306FFFDE165ED66FB94FE72A139" box="[323,393,1054,1077]" captionStart="Table 1" captionStartId="19.[192,257,836,859]" captionText="Table 1: Comparative measurements of adults of Lanzatus somalilandus sp. n. and Orthochirus afar sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="3" pageNumber="2">Tab. 1</tableCitation>
,
<figureCitation id="CBBA2A38FFFDE165EDBFFB94FE10A139" box="[410,491,1054,1077]" captionStart="Figures 1529" captionStartId="7.[192,278,1567,1590]" captionTargetBox="[192,1439,175,1532]" captionTargetId="figure-6@7.[192,1439,168,1538]" captionTargetPageId="7" captionText="Figures 1529: Lanzatus. Figures 1528. L. somalilandus sp. n. Figures 1527. Female paratype. Figures 1522. Pedipalp segments, movable finger dentition (15), chela dorsal (16), external (17) and ventral (18), patella dorsal (19), external (20) and ventral (21), trochanter and femur dorsal (22). Trichobothrial pattern indicated in Figures 1720 and 22. Figures 2326. Left legs IIV, retrolateral aspect. Figures 2729. Telson lateral. Figures 2728. L. somalilandus sp. n., female paratype (27) and male holotype (28). Figure 29. L. somalicus, male paratype." pageId="3" pageNumber="2">Fig. 23</figureCitation>
; Prendini &amp; Wheeler, 2005: 462, 481; Dupré, 2007: 10, 13, 17;
<treatmentCitation id="D22010ACFFFDE165EE58FBB7FEF6A17F" author="KOVARIK" page="23" pageId="3" pageNumber="2" year="2009">
<bibRefCitation id="37104B4CFFFDE165EE58FBB7FEF6A17F" author="KOVARIK" pageId="3" pageNumber="2" pagination="1 - 12" refId="ref13482" refString="KOVARIK, F., F. STAHLAVSKY, T. KORIN- KOVA, J. KRAL &amp; T. VAN DER ENDE. 2009. Tityus ythieri Lourenco, 2007 is a synonym of Tityus magnimanus Pocock, 1897 (Scorpiones: Buthidae): a combined approach using morphology, hybridization experiments, chromosomes, and mitochondrial DNA. Euscorpius, 77: 1 - 12." type="journal article" year="2009">Kovařík, 2009: 23</bibRefCitation>
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, 31 (syn. by
<treatmentCitation id="D22010ACFFFDE165EDB3FBD6FD7DA17F" author="LOURENCO" box="[406,646,1116,1139]" page="174" pageId="3" pageNumber="2" year="2001">
<bibRefCitation id="37104B4CFFFDE165EDB3FBD6FD7DA17F" author="LOURENCO" box="[406,646,1116,1139]" pageId="3" pageNumber="2" pagination="173 - 175" refId="ref13890" refString="LOURENCO, W. R. 2001 b. A propos de quelques decisions taxonomiques concernant des scorpions des families Buthidae, Chaerilidae et Scorpiopidae. Biogeographica, 77 (4): 173 - 175." type="journal article" year="2001">Lourenço, 2001b: 174</bibRefCitation>
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).
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</subSubSection>
<subSubSection id="1B9B6536FFFDE165ECE5FB1CFEBDA267" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="533E36BDFFFDE165ECE5FB1CFD23A1A1" blockId="3.[192,728,1174,1197]" box="[192,728,1174,1197]" pageId="3" pageNumber="2">
<materialsCitation id="E3E93CE0FFFDE165ECE5FB1CFD2FA1A1" box="[192,724,1174,1197]" collectionCode="SPECIES" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
<typeStatus id="8C3A881FFFFDE165ECE5FB1CFF03A1A1" box="[192,248,1174,1197]" pageId="3" pageNumber="2">TYPE</typeStatus>
<collectionCode id="3590AE78FFFDE165ECDBFB10FEAAA1A1" box="[254,337,1178,1197]" pageId="3" pageNumber="2">SPECIES</collectionCode>
.
<taxonomicName id="94814D3EFFFDE165ED7BFB1CFD2FA1A1" authority="Kovarik, 2001" authorityName="Kovarik" authorityYear="2001" box="[350,724,1174,1197]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="61F5EAAFFFFDE165ED7BFB1CFDCBA1A1" box="[350,560,1174,1197]" italics="true" pageId="3" pageNumber="2">Lanzatus somalicus</emphasis>
Kovařík, 2001
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.
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<paragraph id="533E36BDFFFDE165ECE5FB5AFEBDA267" blockId="3.[192,802,1232,1899]" pageId="3" pageNumber="2">
EMENDED DIAGNOSIS. Total length 17.927.6 mm. Pedipalps orthobothriotaxic
<typeStatus id="8C3A881FFFFDE165EE21FB65FDC9A00A" box="[516,562,1263,1286]" pageId="3" pageNumber="2">type</typeStatus>
A (Vachon, 1974); dorsal trichobothria of femur arranged in
<emphasis id="61F5EAAFFFFDE165EE59FA84FD72A029" box="[636,649,1294,1317]" italics="true" pageId="3" pageNumber="2">β</emphasis>
-configuration (Vachon, 1975); trichobothrium
<emphasis id="61F5EAAFFFFDE165EE02FAA6FDCFA04F" box="[551,564,1324,1347]" italics="true" pageId="3" pageNumber="2">d</emphasis>
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of femur internal to dorsointernal carina;
<emphasis id="61F5EAAFFFFDE165ED90FAC1FE39A06E" box="[437,450,1355,1378]" italics="true" pageId="3" pageNumber="2">d</emphasis>
<subScript id="CF0534F8FFFDE165EDE7FADFFE30A068" attach="left" box="[450,459,1365,1380]" fontSize="6" pageId="3" pageNumber="2">3</subScript>
of patella internal to dorsomedian carina, when carina present;
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<collectionCode id="3590AE78FFFDE165EE71FAE0FD9FA08D" box="[596,612,1386,1409]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="2" type="Museum">V</collectionCode>
</emphasis>
<subScript id="CF0534F8FFFDE165EE41FAFEFD96A08F" attach="left" box="[612,621,1396,1411]" fontSize="6" pageId="3" pageNumber="2">2</subScript>
on chela manus located behind
<emphasis id="61F5EAAFFFFDE165ED41FA02FE8FA093" box="[356,372,1416,1439]" italics="true" pageId="3" pageNumber="2">
<collectionCode id="3590AE78FFFDE165ED41FA02FE8FA093" box="[356,372,1416,1439]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="2" type="Museum">V</collectionCode>
</emphasis>
<subScript id="CF0534F8FFFDE165ED50FA19FE85A0AE" attach="right" box="[373,382,1427,1442]" fontSize="6" pageId="3" pageNumber="2">1</subScript>
;
<emphasis id="61F5EAAFFFFDE165EDA9FA02FE60A093" box="[396,411,1416,1439]" italics="true" pageId="3" pageNumber="2">it</emphasis>
on distal fixed finger. Pectines with fulcra. Pectine teeth number 1824. Basal middle lamella of pectines in females not dilated. Hemispermatophore unknown. Sternum subtriangular. Tibial spurs absent on legs I
<collectionCode id="3590AE78FFFDE165EDE6F989FE1AA316" box="[451,481,1539,1562]" pageId="3" pageNumber="2">IV</collectionCode>
. Cheliceral fixed finger with one ventral denticle, margins of fingers with standard pattern of buthid dentition (Vachon, 1963). Carapace without distinct carinae, in lateral view with entire dorsal surface horizontal, or nearly so. Lateral eyes number 5 pairs. Pedipalp chela movable finger with distinct granules divided into 7 diagonal rows. Tergites I
<collectionCode id="3590AE78FFFDE165EF23F931FCD9A3DE" box="[774,802,1723,1746]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="2">VI</collectionCode>
smooth to finely granulated with one indicated carina or acarinate. Stigmata are narrow slits. Metasomal segments all elongate, smooth, acarinate. Telson elongate, smooth, without subaculear tubercle, aculeus shorter than vesicle.
</paragraph>
</subSubSection>
<subSubSection id="1B9B6536FFFDE165EF1BFF4DFB5CA0BF" pageId="3" pageNumber="2" type="discussion">
<paragraph id="533E36BDFFFDE165EF1BFF4DFC6CA742" blockId="3.[830,1441,199,590]" pageId="3" pageNumber="2">
COMMENTS ABOUT SYNONYMY.
<bibRefCitation id="37104B4CFFFDE165E887FF4DFA5BA5D2" author="LOURENCO" box="[1186,1440,199,222]" pageId="3" pageNumber="2" pagination="173 - 175" refId="ref13890" refString="LOURENCO, W. R. 2001 b. A propos de quelques decisions taxonomiques concernant des scorpions des families Buthidae, Chaerilidae et Scorpiopidae. Biogeographica, 77 (4): 173 - 175." type="journal article" year="2001">Lourenço (2001b: 174)</bibRefCitation>
stated that
<taxonomicName id="94814D3EFFFDE165EFECFF6CFAA4A5F1" authority="Kovarik, 2001" authorityName="Kovarik" authorityYear="2001" box="[969,1375,230,253]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="61F5EAAFFFFDE165EFECFF6CFB5CA5F1" box="[969,1191,230,253]" italics="true" pageId="3" pageNumber="2">Lanzatus somalicus</emphasis>
Kovařík, 2001
</taxonomicName>
is a synonym of
<taxonomicName id="94814D3EFFFDE165EFE9FE8EFA88A417" authority="Lourenco, 2001" authorityName="Lourenco" authorityYear="2001" box="[972,1395,260,283]" class="Arachnida" family="Buthidae" genus="Sabinebuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="61F5EAAFFFFDE165EFE9FE8EFB43A417" box="[972,1208,260,283]" italics="true" pageId="3" pageNumber="2">Sabinebuthus elegans</emphasis>
Lourenço, 2001
</taxonomicName>
. He provided no evidence whatsoever that
<taxonomicName id="94814D3EFFFDE165E8C1FEA9FAA8A436" authorityName="Lourenco" authorityYear="2001" box="[1252,1363,291,314]" class="Arachnida" family="Buthidae" genus="Sabinebuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="61F5EAAFFFFDE165E8C1FEA9FAA8A436" box="[1252,1363,291,314]" italics="true" pageId="3" pageNumber="2">S. elegans</emphasis>
</taxonomicName>
and
<taxonomicName id="94814D3EFFFDE165E9AEFEA9FC52A455" authorityName="Kovarik" authorityYear="2001" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="61F5EAAFFFFDE165E9AEFEA9FC52A455" italics="true" pageId="3" pageNumber="2">L. somalicus</emphasis>
</taxonomicName>
are the same species, and instead only told a story about searching for specimens and competing, which the first author perceives very differently. However, this discussion has no place in a technical paper and we therefore refrain from commenting. The published descriptions of these two genera/species include substantial differences that would indicate that they belong to separate genera, distinguishable in a key as follows:
</paragraph>
<paragraph id="533E36BDFFFDE165EF1BFDF0FA62A664" blockId="3.[830,1441,634,872]" pageId="3" pageNumber="2">
Movable finger of pedipalp with six rows of granules which do not form diagonal rows and lack accessory granules (see fig.
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<bibRefCitation id="37104B4CFFFDE165E867FD32FAD3A7C3" author="LOURENCO" box="[1090,1320,696,719]" pageId="3" pageNumber="2" pagination="15 - 20" refId="ref13852" refString="LOURENCO, W. R. 2001 a. Un nouveau genre de Buthidae, probable vicariant geographique d' Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço, 2001a: 19</bibRefCitation>
) ……..…. .......................................................
<taxonomicName id="94814D3EFFFDE165E891FD5CFA67A7E1" authorityName="Lourenco" authorityYear="2001" box="[1204,1436,726,749]" class="Arachnida" family="Buthidae" genus="Sabinebuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="61F5EAAFFFFDE165E891FD5CFA67A7E1" box="[1204,1436,726,749]" italics="true" pageId="3" pageNumber="2">Sabinebuthus elegans</emphasis>
</taxonomicName>
Movable finger of pedipalp with seven rows of granules which form diagonal rows and have accessory granules (see
<figureCitation id="CBBA2A38FFFDE165EFFCFCB8FBD1A645" box="[985,1066,818,841]" captionStart="Figures 1529" captionStartId="7.[192,278,1567,1590]" captionTargetBox="[192,1439,175,1532]" captionTargetId="figure-6@7.[192,1439,168,1538]" captionTargetPageId="7" captionText="Figures 1529: Lanzatus. Figures 1528. L. somalilandus sp. n. Figures 1527. Female paratype. Figures 1522. Pedipalp segments, movable finger dentition (15), chela dorsal (16), external (17) and ventral (18), patella dorsal (19), external (20) and ventral (21), trochanter and femur dorsal (22). Trichobothrial pattern indicated in Figures 1720 and 22. Figures 2326. Left legs IIV, retrolateral aspect. Figures 2729. Telson lateral. Figures 2728. L. somalilandus sp. n., female paratype (27) and male holotype (28). Figure 29. L. somalicus, male paratype." pageId="3" pageNumber="2">Fig. 15</figureCitation>
and fig.
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Kovařík, 2001: 44) ...........………....................................................
<taxonomicName id="94814D3EFFFDE165E91CFCDBFA62A664" authorityName="Kovarik" authorityYear="2001" box="[1337,1433,849,872]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="61F5EAAFFFFDE165E91CFCDBFA62A664" box="[1337,1433,849,872]" italics="true" pageId="3" pageNumber="2">Lanzatus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="533E36BDFFFDE165EF1BFC19FB5CA0BF" blockId="3.[830,1441,915,1459]" pageId="3" pageNumber="2">
Since the request of the first author for a loan of the
<typeStatus id="8C3A881FFFFDE165EF1BFC38FC60A6C5" box="[830,923,946,969]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
of
<taxonomicName id="94814D3EFFFDE165EFE2FC38FB48A6C5" authorityName="Lourenco" authorityYear="2001" box="[967,1203,946,969]" class="Arachnida" family="Buthidae" genus="Sabinebuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="61F5EAAFFFFDE165EFE2FC38FB48A6C5" box="[967,1203,946,969]" italics="true" pageId="3" pageNumber="2">Sabinebuthus elegans</emphasis>
</taxonomicName>
was refused, we can only speculate about two possibilities:
<emphasis id="61F5EAAFFFFDE165E935FC5AFAE6A6EB" bold="true" box="[1296,1309,976,999]" pageId="3" pageNumber="2">a</emphasis>
) fig.
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<bibRefCitation id="37104B4CFFFDE165EF1BFC65FBD3A10A" author="LOURENCO" box="[830,1064,1007,1030]" pageId="3" pageNumber="2" pagination="15 - 20" refId="ref13852" refString="LOURENCO, W. R. 2001 a. Un nouveau genre de Buthidae, probable vicariant geographique d' Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço, 2001a: 19</bibRefCitation>
is correct and
<bibRefCitation id="37104B4CFFFDE165E8F8FC65FC9DA129" author="LOURENCO" pageId="3" pageNumber="2" pagination="173 - 175" refId="ref13890" refString="LOURENCO, W. R. 2001 b. A propos de quelques decisions taxonomiques concernant des scorpions des families Buthidae, Chaerilidae et Scorpiopidae. Biogeographica, 77 (4): 173 - 175." type="journal article" year="2001">Lourenço, 2001b: 174</bibRefCitation>
is incorrect in claiming that the two taxa are synonyms;
<emphasis id="61F5EAAFFFFDE165EF9FFBA6FC32A14F" bold="true" box="[954,969,1068,1091]" pageId="3" pageNumber="2">b</emphasis>
) fig.
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<bibRefCitation id="37104B4CFFFDE165E81FFBA6FAE5A14F" author="LOURENCO" box="[1082,1310,1068,1091]" pageId="3" pageNumber="2" pagination="15 - 20" refId="ref13852" refString="LOURENCO, W. R. 2001 a. Un nouveau genre de Buthidae, probable vicariant geographique d' Anomalobuthus Kraepelin (Chelicerata, Scorpiones). Biogeographica, 77 (1): 15 - 20." type="journal article" year="2001">Lourenço, 2001a: 19</bibRefCitation>
, and with it the entire diagnosis of
<taxonomicName id="94814D3EFFFDE165E848FBC1FA98A16E" authorityName="Lourenco" authorityYear="2001" box="[1133,1379,1099,1122]" class="Arachnida" family="Buthidae" genus="Sabinebuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="61F5EAAFFFFDE165E848FBC1FA98A16E" box="[1133,1379,1099,1122]" italics="true" pageId="3" pageNumber="2">Sabinebuthus elegans</emphasis>
</taxonomicName>
, are incorrect and these two genera/species are synonyms. To mention one additional detail,
<bibRefCitation id="37104B4CFFFDE165E8B9FB02FA5BA193" author="LOURENCO" box="[1180,1440,1160,1183]" pageId="3" pageNumber="2" pagination="173 - 175" refId="ref13890" refString="LOURENCO, W. R. 2001 b. A propos de quelques decisions taxonomiques concernant des scorpions des families Buthidae, Chaerilidae et Scorpiopidae. Biogeographica, 77 (4): 173 - 175." type="journal article" year="2001">Lourenço (2001b: 174)</bibRefCitation>
correctly states that the description of
<taxonomicName id="94814D3EFFFDE165E92AFB2DFC6AA1D1" authorityName="Kovarik" authorityYear="2001" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="61F5EAAFFFFDE165E92AFB2DFC6AA1D1" italics="true" pageId="3" pageNumber="2">Lanzatus somalicus</emphasis>
</taxonomicName>
was published on
<date id="273F107DFFFDE165E845FB4CFAF1A1D1" box="[1120,1290,1222,1245]" pageId="3" pageNumber="2" value="2001-03-30">30 March 2001</date>
. However, he is incorrect in stating that the description of
<taxonomicName id="94814D3EFFFDE165E96AFB6EFC1AA016" authorityName="Lourenco" authorityYear="2001" class="Arachnida" family="Buthidae" genus="Sabinebuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="61F5EAAFFFFDE165E96AFB6EFC1AA016" italics="true" pageId="3" pageNumber="2">Sabinebuthus elegans</emphasis>
</taxonomicName>
was published on
<date id="273F107DFFFDE165E89FFA89FAA1A016" box="[1210,1370,1283,1306]" pageId="3" pageNumber="2" value="2001-03-01">1 March 2001</date>
, since the date of the issue is given only as “
<date id="273F107DFFFDE165E921FAA8FA69A035" box="[1284,1426,1314,1337]" pageId="3" pageNumber="2" value="2001-03">March 2001</date>
”. According to the ICZN, if only the month is given then the last day of that month, not the first, is the valid publication date. Consequently,
<taxonomicName id="94814D3EFFFDE165E895FAF4FA5BA099" authorityName="Lourenco" authorityYear="2001" box="[1200,1440,1406,1429]" class="Arachnida" family="Buthidae" genus="Sabinebuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="61F5EAAFFFFDE165E895FAF4FA5BA099" box="[1200,1440,1406,1429]" italics="true" pageId="3" pageNumber="2">Sabinebuthus elegans</emphasis>
</taxonomicName>
was described on
<date id="273F107DFFFDE165EFD8FA16FB58A0BF" box="[1021,1187,1436,1459]" pageId="3" pageNumber="2" value="2001-03-31">31 March 2001</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>