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<document id="92F0EDC76E6A740B24D94A736D57BEC4" ID-DOI="10.11646/zootaxa.5352.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8407061" ID-ZooBank="E1B1ADA5-60C5-4185-B531-6708FD714A1B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1696427801038" checkinUser="plazi" docAuthor="Pantoja, Gabriel M., Viana, Ana D. L. &amp; Salles, Frederico F." docDate="2023" docId="670A4B50F9406428FF2AFF58FA7CFE8C" docLanguage="en" docName="zootaxa.5352.1.6.pdf" docOrigin="Zootaxa 5352 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5352.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Campsurus mirim Pantoja &amp; Viana &amp; Salles 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="149" masterDocId="9B333328F942642DFFBDFF87FFE1FF88" masterDocTitle="A new species of Campsurus Eaton, 1868 (Ephemeroptera: Polymitarcyidae) from the Doce River Basin, Brazil" masterLastPageNumber="150" masterPageNumber="145" pageNumber="147" updateTime="1696428173079" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="94D308F9E1EA9370EE15969B5EB41B7C">A new species of Campsurus Eaton, 1868 (Ephemeroptera: Polymitarcyidae) from the Doce River Basin, Brazil</mods:title>
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<mods:namePart id="952055CF2837A59F9BBDFFE386E9F773">Pantoja, Gabriel M.</mods:namePart>
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<mods:affiliation id="737A6809CA49C7034A1AD3A9D2522C8D">Museu de Entomologia, Depto. de Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil &amp; Programa de Pós-graduação em Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil &amp; gabriel. pantoja @ ufv. br; https: // orcid. org / 0000 - 0002 - 1769 - 6070</mods:affiliation>
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<mods:namePart id="F3487652F6649CBC4D615E26172F9A32">Viana, Ana D. L.</mods:namePart>
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<mods:affiliation id="A7F4B3EA258C6C1CECE9641F5E468B28">Museu de Entomologia, Depto. de Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil &amp; Programa de Pós-graduação em Ecologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil &amp; ana. leite @ ufv. br; https: // orcid. org / 0000 - 0001 - 5374 - 1257</mods:affiliation>
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<mods:namePart id="D78AF18CC11A621476EE7A786F210809">Salles, Frederico F.</mods:namePart>
<mods:affiliation id="09D92A667C93C4C9C4BBFF56CA5A1C3F">Museu de Entomologia, Depto. de Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil</mods:affiliation>
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<taxonomicName id="28A381C5F940642FFF2AFF58FE93FF71" authority="Pantoja &amp; Viana &amp; Salles, 2023" authorityName="Pantoja &amp; Viana &amp; Salles" authorityYear="2023" box="[151,370,223,249]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="mirim" status="sp. nov.">
<emphasis id="DDD72654F940642FFF2AFF58FE93FF71" bold="true" box="[151,370,223,249]" italics="true" pageId="2" pageNumber="147">Campsurus mirim</emphasis>
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(
<figureCitation id="7798E6C3F940642FFE5AFF58FDB8FF71" box="[487,601,223,249]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="2.[151,250,1720,1744]" captionStartId-1="3.[151,250,1889,1913]" captionStartId-2="4.[151,250,1490,1514]" captionTargetBox-0="[231,1357,417,1696]" captionTargetBox-1="[158,1430,885,1866]" captionTargetBox-2="[296,1292,181,1465]" captionTargetId-0="figure-80@2.[231,1357,417,1696]" captionTargetId-1="figure-332@3.[158,1430,885,1866]" captionTargetId-2="figure-26@4.[296,1292,181,1465]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 3. Campsurus mirim sp. nov. male imago (holotype): A, head and thorax, dorsal view; B, head and thorax, lateral view; C, abdomen, dorsal view; D, abdomen, lateral view." captionText-1="FIGURE 4. Campsurus mirim sp. nov. male imago (holotype): Fore and hind wing." captionText-2="FIGURE 5. Campsurus mirim sp. nov. genitalia of male imago (holotype): A, genitalia, ventral view; B, penes lobes, ventral view; C, genitalia, lateral view. Abbreviations: IX, sternum IX; ml, main lobe; sl, secondary lobe; p, pedestal." figureDoi-0="http://doi.org/10.5281/zenodo.8407068" figureDoi-1="http://doi.org/10.5281/zenodo.8407072" figureDoi-2="http://doi.org/10.5281/zenodo.8407074" httpUri-0="https://zenodo.org/record/8407068/files/figure.png" httpUri-1="https://zenodo.org/record/8407072/files/figure.png" httpUri-2="https://zenodo.org/record/8407074/files/figure.png" pageId="2" pageNumber="147">Figs. 3–5</figureCitation>
)
</emphasis>
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</paragraph>
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<emphasis id="DDD72654F940642FFF2AFEA0FEA4FEC9" bold="true" box="[151,325,295,321]" pageId="2" pageNumber="147">Type material.</emphasis>
<materialsCitation id="5FCBF01BF940642FFEF6FEA0FF19FE01" collectingDate="2022-09-13" collectionCode="UFVB" collectorName="Viana, A. D. L. &amp; Bonfa, P. &amp; Ataide, A." country="Brazil" elevation="32" latitude="-19.330278" location="Lagoa Juparana Mirim" longLatPrecision="1" longitude="-40.170086" municipality="Linhares" pageId="2" pageNumber="147" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
<emphasis id="DDD72654F940642FFEF6FEA0FE59FEC9" bold="true" box="[331,440,295,321]" pageId="2" pageNumber="147">
<typeStatus id="301844E4F940642FFEF6FEA0FE59FEC9" box="[331,440,295,321]" pageId="2" pageNumber="147">Holotype</typeStatus>
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: ♁ imago in alcohol, 
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, 
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, 
<collectingMunicipality id="0F78603CF940642FFC57FEA0FBADFEC9" box="[1002,1100,295,321]" pageId="2" pageNumber="147">Linhares</collectingMunicipality>
municipality, 
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(Lagoa Nova), 
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, 
<geoCoordinate id="8A979C81F940642FFDFFFECCFD08FEED" box="[578,745,331,357]" degrees="40" direction="west" minutes="10" orientation="longitude" pageId="2" pageNumber="147" precision="1" seconds="12.3" value="-40.170086">W40°10’12.3”</geoCoordinate>
, 
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<elevation id="648E1D75F940642FFD45FECCFCD4FEEE" box="[760,821,331,358]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="147" unit="m" value="32.0">32 m</elevation>
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, 
<date id="9B1DDC86F940642FFCF9FECCFC5FFEED" box="[836,958,331,357]" pageId="2" pageNumber="147" value="2022-09-13">
<collectingDate id="8B59256EF940642FFCF9FECCFC5FFEED" box="[836,958,331,357]" pageId="2" pageNumber="147" value="2022-09-13">13.ix.2022</collectingDate>
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, 
<collectorName id="42569F90F940642FFC71FECCFB8CFEED" box="[972,1133,331,357]" pageId="2" pageNumber="147">Viana, A.D.L.</collectorName>
, 
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, 
<collectorName id="42569F90F940642FFB49FECCFA88FEED" box="[1268,1385,331,357]" pageId="2" pageNumber="147">Ataide, A.</collectorName>
col. (
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)
</materialsCitation>
. 
<materialsCitation id="5FCBF01BF940642FFEBEFEE8FBBAFE01" box="[259,1115,367,396]" collectingDate="2022-09-13" collectionCode="UFVB, MNRJ" collectorName="Viana, A. D. L. &amp; Bonfa, P. &amp; Ataide, A." country="Brazil" elevation="32" latitude="-19.330278" location="Lagoa Juparana Mirim" longLatPrecision="1" longitude="-40.170086" municipality="Linhares" pageId="2" pageNumber="147" specimenCount="1" stateProvince="Espirito Santo" typeStatus="paratype">
<emphasis id="DDD72654F940642FFEBEFEE8FE9AFE01" bold="true" box="[259,379,367,393]" pageId="2" pageNumber="147">
<typeStatus id="301844E4F940642FFEBEFEE8FE9AFE01" box="[259,379,367,393]" pageId="2" pageNumber="147">Paratypes</typeStatus>
</emphasis>
: 26 ♁ imagos, same data as holotype (16 at 
<collectionCode id="89B26283F940642FFCD1FEF7FC58FE02" box="[876,953,368,394]" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="2" pageNumber="147">UFVB</collectionCode>
, 10 at 
<collectionCode id="89B26283F940642FFBBEFEE8FBB2FE01" box="[1027,1107,367,393]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="147">MNRJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<paragraph id="EF1CFA46F940642FFF2AF93FFD04F97C" blockId="2.[151,1436,1720,1780]" pageId="2" pageNumber="147">
<emphasis id="DDD72654F940642FFF2AF93FFEF4F958" bold="true" box="[151,277,1720,1744]" pageId="2" pageNumber="147">FIGURE 3.</emphasis>
<taxonomicName id="28A381C5F940642FFEA0F93EFE34F958" authority="Pantoja &amp; Viana &amp; Salles, 2023" authorityName="Pantoja &amp; Viana &amp; Salles" authorityYear="2023" box="[285,469,1721,1744]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="mirim" status="sp. nov.">
<emphasis id="DDD72654F940642FFEA0F93EFE34F958" box="[285,469,1721,1744]" italics="true" pageId="2" pageNumber="147">Campsurus mirim</emphasis>
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<taxonomicNameLabel id="C6E49B2FF940642FFE60F93FFDCEF958" box="[477,559,1720,1744]" pageId="2" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
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male imago (holotype): A, head and thorax, dorsal view; B, head and thorax, lateral view; C, abdomen, dorsal view; D, abdomen, lateral view.
</paragraph>
</caption>
<subSubSection id="A7B9A9CDF940642EFF7AF8C1FD63FCD5" lastPageId="3" lastPageNumber="148" pageId="2" pageNumber="147" type="diagnosis">
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<emphasis id="DDD72654F940642FFF7AF8C1FEA3F8E8" bold="true" box="[199,322,1862,1888]" pageId="2" pageNumber="147">Diagnosis.</emphasis>
The male imago of 
<taxonomicName id="28A381C5F940642FFD88F8C0FD7CF8E8" authority="Pantoja &amp; Viana &amp; Salles, 2023" authorityName="Pantoja &amp; Viana &amp; Salles" authorityYear="2023" box="[565,669,1863,1889]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="mirim" status="sp. nov.">
<emphasis id="DDD72654F940642FFD88F8C0FD7CF8E8" box="[565,669,1863,1889]" italics="true" pageId="2" pageNumber="147">C. mirim</emphasis>
</taxonomicName>
<emphasis id="DDD72654F940642FFD1AF8C0FCE5F8E9" bold="true" box="[679,772,1863,1889]" pageId="2" pageNumber="147">
<taxonomicNameLabel id="C6E49B2FF940642FFD1AF8C0FCE5F8E9" box="[679,772,1863,1889]" pageId="2" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from other species of the genus by the following combination of characteristics: small to medium size (body 
<quantity id="285B57A3F940642FFC56F8ECFBA4F80D" box="[1003,1093,1899,1925]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="147" unit="mm" value="7.3">7.3 mm</quantity>
, forewings 
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); blackish pigments on dorsum of head widely and strongly marked in fresh material (
<figureCitation id="7798E6C3F940642FFC55F809FBA3F821" box="[1000,1090,1934,1961]" captionStart="FIGURE 3" captionStartId="2.[151,250,1720,1744]" captionTargetBox="[231,1357,417,1696]" captionTargetId="figure-80@2.[231,1357,417,1696]" captionTargetPageId="2" captionText="FIGURE 3. Campsurus mirim sp. nov. male imago (holotype): A, head and thorax, dorsal view; B, head and thorax, lateral view; C, abdomen, dorsal view; D, abdomen, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8407068" httpUri="https://zenodo.org/record/8407068/files/figure.png" pageId="2" pageNumber="147">Fig. 3A</figureCitation>
); abdominal color pattern with pale transverse dashes on terga III–VII (
<figureCitation id="7798E6C3F940642FFDE8F835FCEEF845" box="[597,783,1970,1997]" captionStart="FIGURE 3" captionStartId="2.[151,250,1720,1744]" captionTargetBox="[231,1357,417,1696]" captionTargetId="figure-80@2.[231,1357,417,1696]" captionTargetPageId="2" captionText="FIGURE 3. Campsurus mirim sp. nov. male imago (holotype): A, head and thorax, dorsal view; B, head and thorax, lateral view; C, abdomen, dorsal view; D, abdomen, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8407068" httpUri="https://zenodo.org/record/8407068/files/figure.png" pageId="2" pageNumber="147">Figs. 3C and 3D</figureCitation>
); posterior margin of sternum IX triangular at middle (
<figureCitation id="7798E6C3F940642FFAD2F835FF5CF879" captionStart="FIGURE 5" captionStartId="4.[151,250,1490,1514]" captionTargetBox="[296,1292,181,1465]" captionTargetId="figure-26@4.[296,1292,181,1465]" captionTargetPageId="4" captionText="FIGURE 5. Campsurus mirim sp. nov. genitalia of male imago (holotype): A, genitalia, ventral view; B, penes lobes, ventral view; C, genitalia, lateral view. Abbreviations: IX, sternum IX; ml, main lobe; sl, secondary lobe; p, pedestal." figureDoi="http://doi.org/10.5281/zenodo.8407074" httpUri="https://zenodo.org/record/8407074/files/figure.png" pageId="2" pageNumber="147">Fig. 5A</figureCitation>
); pedestals with both posterior margins (inner and outer) approximately of the same length, both acute (
<figureCitation id="7798E6C3F940642FFA8BF851FA6EF879" box="[1334,1423,2006,2033]" captionStart="FIGURE 5" captionStartId="4.[151,250,1490,1514]" captionTargetBox="[296,1292,181,1465]" captionTargetId="figure-26@4.[296,1292,181,1465]" captionTargetPageId="4" captionText="FIGURE 5. Campsurus mirim sp. nov. genitalia of male imago (holotype): A, genitalia, ventral view; B, penes lobes, ventral view; C, genitalia, lateral view. Abbreviations: IX, sternum IX; ml, main lobe; sl, secondary lobe; p, pedestal." figureDoi="http://doi.org/10.5281/zenodo.8407074" httpUri="https://zenodo.org/record/8407074/files/figure.png" pageId="2" pageNumber="147">Fig. 5A</figureCitation>
); penes relatively wide and large, ventrally curved on apical ½ and with apex of main lobe slightly twisted, gonopore not visible (
<figureCitation id="7798E6C3F941642EFEA1FF3CFE26FF5D" box="[284,455,186,213]" captionStart="FIGURE 5" captionStartId="4.[151,250,1490,1514]" captionTargetBox="[296,1292,181,1465]" captionTargetId="figure-26@4.[296,1292,181,1465]" captionTargetPageId="4" captionText="FIGURE 5. Campsurus mirim sp. nov. genitalia of male imago (holotype): A, genitalia, ventral view; B, penes lobes, ventral view; C, genitalia, lateral view. Abbreviations: IX, sternum IX; ml, main lobe; sl, secondary lobe; p, pedestal." figureDoi="http://doi.org/10.5281/zenodo.8407074" httpUri="https://zenodo.org/record/8407074/files/figure.png" pageId="3" pageNumber="148">Figs. 5A and B</figureCitation>
).
</paragraph>
<paragraph id="EF1CFA46F941642EFF7AFF58FA86FCB1" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="148">
<emphasis id="DDD72654F941642EFF7AFF58FEB8FF70" bold="true" box="[199,345,222,249]" pageId="3" pageNumber="148">Male imago</emphasis>
(
<figureCitation id="7798E6C3F941642EFED0FF58FE3FFF71" box="[365,478,222,249]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="2.[151,250,1720,1744]" captionStartId-1="3.[151,250,1889,1913]" captionStartId-2="4.[151,250,1490,1514]" captionTargetBox-0="[231,1357,417,1696]" captionTargetBox-1="[158,1430,885,1866]" captionTargetBox-2="[296,1292,181,1465]" captionTargetId-0="figure-80@2.[231,1357,417,1696]" captionTargetId-1="figure-332@3.[158,1430,885,1866]" captionTargetId-2="figure-26@4.[296,1292,181,1465]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 3. Campsurus mirim sp. nov. male imago (holotype): A, head and thorax, dorsal view; B, head and thorax, lateral view; C, abdomen, dorsal view; D, abdomen, lateral view." captionText-1="FIGURE 4. Campsurus mirim sp. nov. male imago (holotype): Fore and hind wing." captionText-2="FIGURE 5. Campsurus mirim sp. nov. genitalia of male imago (holotype): A, genitalia, ventral view; B, penes lobes, ventral view; C, genitalia, lateral view. Abbreviations: IX, sternum IX; ml, main lobe; sl, secondary lobe; p, pedestal." figureDoi-0="http://doi.org/10.5281/zenodo.8407068" figureDoi-1="http://doi.org/10.5281/zenodo.8407072" figureDoi-2="http://doi.org/10.5281/zenodo.8407074" httpUri-0="https://zenodo.org/record/8407068/files/figure.png" httpUri-1="https://zenodo.org/record/8407072/files/figure.png" httpUri-2="https://zenodo.org/record/8407074/files/figure.png" pageId="3" pageNumber="148">Figs. 3–5</figureCitation>
). Length (mm): body, 7.3; forewing, 6.1; hind wing, 2.9; fore leg, 3.06. General coloration yellowish white with gray markings dorsally. Head (
<figureCitation id="7798E6C3F941642EFCE6FE84FC3FFE95" box="[859,990,259,285]" captionStart="FIGURE 3" captionStartId="2.[151,250,1720,1744]" captionTargetBox="[231,1357,417,1696]" captionTargetId="figure-80@2.[231,1357,417,1696]" captionTargetPageId="2" captionText="FIGURE 3. Campsurus mirim sp. nov. male imago (holotype): A, head and thorax, dorsal view; B, head and thorax, lateral view; C, abdomen, dorsal view; D, abdomen, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8407068" httpUri="https://zenodo.org/record/8407068/files/figure.png" pageId="3" pageNumber="148">Figs. 3A–B</figureCitation>
) heavily washed with black on dorsum, especially at base of ocelli, scape and pedicel slightly washed with black, flagellum hyaline. Thorax (
<figureCitation id="7798E6C3F941642EFAB4FEA0FA71FEC9" box="[1289,1424,295,321]" captionStart="FIGURE 3" captionStartId="2.[151,250,1720,1744]" captionTargetBox="[231,1357,417,1696]" captionTargetId="figure-80@2.[231,1357,417,1696]" captionTargetPageId="2" captionText="FIGURE 3. Campsurus mirim sp. nov. male imago (holotype): A, head and thorax, dorsal view; B, head and thorax, lateral view; C, abdomen, dorsal view; D, abdomen, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8407068" httpUri="https://zenodo.org/record/8407068/files/figure.png" pageId="3" pageNumber="148">Figs. 3A–B</figureCitation>
). Pronotum translucent shaded with black on anterior portion and on lateral and posterior margins of posterior portion; medially with blackish medial line well marked. Meso- and metanota yellowish white diffusely washed with black; black marks more heavily on anteronotal impression, longitudinal median suture, and area between posteroscutal protuberance. Pleura and sterna yellowish white except on prothorax shaded purplish grey. Forelegs shaded with purplish black, lighter on tarsi and even lighter on apex of claws; vestiges of middle and hind legs yellowish white, except for black middle coxa. Wings (
<figureCitation id="7798E6C3F941642EFDF5FE78FD6FFD91" box="[584,654,511,537]" captionStart="FIGURE 4" captionStartId="3.[151,250,1889,1913]" captionTargetBox="[158,1430,885,1866]" captionTargetId="figure-332@3.[158,1430,885,1866]" captionTargetPageId="3" captionText="FIGURE 4. Campsurus mirim sp. nov. male imago (holotype): Fore and hind wing." figureDoi="http://doi.org/10.5281/zenodo.8407072" httpUri="https://zenodo.org/record/8407072/files/figure.png" pageId="3" pageNumber="148">Fig. 4</figureCitation>
). Membrane hyaline, except ¾ of C and Sc areas tinged with purple; veins slightly tinged with purple at base, becoming lighter towards outer margin. Abdomen (
<figureCitation id="7798E6C3F941642EFB09FDA4FAD8FDB5" box="[1204,1337,547,573]" captionStart="FIGURE 3" captionStartId="2.[151,250,1720,1744]" captionTargetBox="[231,1357,417,1696]" captionTargetId="figure-80@2.[231,1357,417,1696]" captionTargetPageId="2" captionText="FIGURE 3. Campsurus mirim sp. nov. male imago (holotype): A, head and thorax, dorsal view; B, head and thorax, lateral view; C, abdomen, dorsal view; D, abdomen, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8407068" httpUri="https://zenodo.org/record/8407068/files/figure.png" pageId="3" pageNumber="148">Figs. 3C–D</figureCitation>
) whitish translucent, shaded with black on terga: terga I–III lighter, terga VIII–X darker; thin medial line on terga V–X; pale lateral transversal band present on terga III–VII. Genitalia (
<figureCitation id="7798E6C3F941642EFC98FDECFC87FD0D" box="[805,870,618,645]" captionStart="FIGURE 5" captionStartId="4.[151,250,1490,1514]" captionTargetBox="[296,1292,181,1465]" captionTargetId="figure-26@4.[296,1292,181,1465]" captionTargetPageId="4" captionText="FIGURE 5. Campsurus mirim sp. nov. genitalia of male imago (holotype): A, genitalia, ventral view; B, penes lobes, ventral view; C, genitalia, lateral view. Abbreviations: IX, sternum IX; ml, main lobe; sl, secondary lobe; p, pedestal." figureDoi="http://doi.org/10.5281/zenodo.8407074" httpUri="https://zenodo.org/record/8407074/files/figure.png" pageId="3" pageNumber="148">Fig. 5</figureCitation>
) whitish yellow, forceps translucent white, slightly tinged with purple at base, sclerotized apex of penes yellowish brown. Posterior margin of sternum IX triangular at middle. Penis curved ventrally, especially at apical ½; main lobe of penis with apex twisted outwards and with its prominent dorsal (posterior) margin very sclerotized; secondary lobe membranous and cylindrical; gonopore not visible. Pedestal short and somewhat flattened, inner-posterior margin well developed and broadly acute, outerposterior margin equal in length to outer margin, with angle more acute. Caudal filaments whitish translucent.
</paragraph>
<paragraph id="EF1CFA46F941642EFF7AFCC4FD63FCD5" blockId="3.[151,1437,151,861]" box="[199,642,834,861]" pageId="3" pageNumber="148">
<emphasis id="DDD72654F941642EFF7AFCC4FDE4FCD5" bold="true" box="[199,517,834,861]" pageId="3" pageNumber="148">Female imago and nymph.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="BBDCAACEF941642EFF2AF8E6FC0FF8F1" ID-DOI="http://doi.org/10.5281/zenodo.8407072" ID-Zenodo-Dep="8407072" box="[151,1006,1889,1913]" httpUri="https://zenodo.org/record/8407072/files/figure.png" pageId="3" pageNumber="148" startId="3.[151,250,1889,1913]" targetBox="[158,1430,885,1866]" targetPageId="3" targetType="figure">
<paragraph id="EF1CFA46F941642EFF2AF8E6FC0FF8F1" blockId="3.[151,1006,1889,1913]" box="[151,1006,1889,1913]" pageId="3" pageNumber="148">
<emphasis id="DDD72654F941642EFF2AF8E6FEF2F8F1" bold="true" box="[151,275,1889,1913]" pageId="3" pageNumber="148">FIGURE 4.</emphasis>
<taxonomicName id="28A381C5F941642EFEA4F8E5FE31F8F1" authority="Pantoja &amp; Viana &amp; Salles, 2023" authorityName="Pantoja &amp; Viana &amp; Salles" authorityYear="2023" box="[281,464,1890,1913]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="mirim" status="sp. nov.">
<emphasis id="DDD72654F941642EFEA4F8E5FE31F8F1" box="[281,464,1890,1913]" italics="true" pageId="3" pageNumber="148">Campsurus mirim</emphasis>
</taxonomicName>
<emphasis id="DDD72654F941642EFE6BF8E6FDC6F8F1" bold="true" box="[470,551,1889,1913]" pageId="3" pageNumber="148">
<taxonomicNameLabel id="C6E49B2FF941642EFE6BF8E6FDC6F8F1" box="[470,551,1889,1913]" pageId="3" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male imago (holotype): Fore and hind wing.
</paragraph>
</caption>
<caption id="BBDCAACEF9466429FF2AFA55FB0AF986" ID-DOI="http://doi.org/10.5281/zenodo.8407074" ID-Zenodo-Dep="8407074" httpUri="https://zenodo.org/record/8407074/files/figure.png" pageId="4" pageNumber="149" startId="4.[151,250,1490,1514]" targetBox="[296,1292,181,1465]" targetPageId="4" targetType="figure">
<paragraph id="EF1CFA46F9466429FF2AFA55FB0AF986" blockId="4.[151,1436,1490,1550]" pageId="4" pageNumber="149">
<emphasis id="DDD72654F9466429FF2AFA55FEF5FA62" bold="true" box="[151,276,1490,1514]" pageId="4" pageNumber="149">FIGURE 5.</emphasis>
<taxonomicName id="28A381C5F9466429FEA6FA54FE32FA62" authority="Pantoja &amp; Viana &amp; Salles, 2023" authorityName="Pantoja &amp; Viana &amp; Salles" authorityYear="2023" box="[283,467,1491,1514]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="mirim" status="sp. nov.">
<emphasis id="DDD72654F9466429FEA6FA54FE32FA62" box="[283,467,1491,1514]" italics="true" pageId="4" pageNumber="149">Campsurus mirim</emphasis>
</taxonomicName>
<emphasis id="DDD72654F9466429FE64FA55FDCAFA62" bold="true" box="[473,555,1490,1514]" pageId="4" pageNumber="149">
<taxonomicNameLabel id="C6E49B2FF9466429FE64FA55FDCAFA62" box="[473,555,1490,1514]" pageId="4" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
genitalia of male imago (holotype): A, genitalia, ventral view; B, penes lobes, ventral view; C, genitalia, lateral view. Abbreviations: IX, sternum IX; ml, main lobe; sl, secondary lobe; p, pedestal.
</paragraph>
</caption>
<subSubSection id="A7B9A9CDF9466428FF7AF9E7FA7CFE8C" lastPageId="5" lastPageNumber="150" pageId="4" pageNumber="149" type="etymology">
<paragraph id="EF1CFA46F9466429FF7AF9E7FD1AF96E" blockId="4.[151,1437,1632,1766]" pageId="4" pageNumber="149">
<emphasis id="DDD72654F9466429FF7AF9E7FEAFF9F2" bold="true" box="[199,334,1632,1658]" pageId="4" pageNumber="149">Etymology.</emphasis>
From the tupi-guarani language, meaning small. An allusion to the size of the species, especially when compared to the relative larger 
<taxonomicName id="28A381C5F9466429FD88F902FD79F916" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[565,664,1669,1695]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="major">
<emphasis id="DDD72654F9466429FD88F902FD79F916" box="[565,664,1669,1695]" italics="true" pageId="4" pageNumber="149">C. major</emphasis>
</taxonomicName>
, and also an allusion to one of the names of the type-locality, the lake Juparan„ Mirim. 
<emphasis id="DDD72654F9466429FF7AF94BFE80F96E" bold="true" box="[199,353,1740,1766]" pageId="4" pageNumber="149">Distribution.</emphasis>
<collectingCountry id="97B4BAD6F9466429FEDAF94BFE51F96E" box="[359,432,1740,1766]" name="Brazil" pageId="4" pageNumber="149">Brazil</collectingCountry>
: 
<collectingRegion id="2D6734A4F9466429FE06F94BFD78F96E" box="[443,665,1740,1766]" country="Brazil" name="Espirito Santo" pageId="4" pageNumber="149">Espírito Santo State</collectingRegion>
(
<figureCitation id="7798E6C3F9466429FD15F94BFD0BF96E" box="[680,746,1740,1766]" captionStart="FIGURE 1" captionStartId="1.[151,250,1020,1044]" captionTargetBox="[210,1385,270,994]" captionTargetId="figure-64@1.[203,1385,270,994]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of Campsurus mirim sp. nov. A, Map of South America with white rectangle highlighting the State of Espírito Santo (thick lines separating Brazil from other countries; narrow lines within Brazil delimiting states; narrow lines outside Brazil delimiting South American countries); B, Map of Espírito Santo with white circle showing the known distribution of Campsurus mirim sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8407063" httpUri="https://zenodo.org/record/8407063/files/figure.png" pageId="4" pageNumber="149">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="EF1CFA46F9466429FF2AF8BEFB86F812" blockId="4.[151,1436,1848,1947]" pageId="4" pageNumber="149">
<taxonomicName id="28A381C5F9466429FF2AF8BEFE81F8DA" authority="Pantoja &amp; Viana &amp; Salles, 2023" authorityName="Pantoja &amp; Viana &amp; Salles" authorityYear="2023" box="[151,352,1849,1875]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="mirim" status="sp. nov.">
<emphasis id="DDD72654F9466429FF2AF8BEFE81F8DA" box="[151,352,1849,1875]" italics="true" pageId="4" pageNumber="149">Campsurus mirim</emphasis>
</taxonomicName>
<emphasis id="DDD72654F9466429FEDAF8BFFE21F8DA" bold="true" box="[359,448,1848,1874]" pageId="4" pageNumber="149">
<taxonomicNameLabel id="C6E49B2FF9466429FEDAF8BFFE21F8DA" box="[359,448,1848,1874]" pageId="4" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
would key out in couplet 3(2) of the key proposed by 
<bibRefCitation id="8B3287B7F9466429FBA0F8BFFAD8F8DA" author="Molineri, C. &amp; Salles, F. F." box="[1053,1337,1848,1874]" pageId="4" pageNumber="149" pagination="301 - 354" refId="ref2691" refString="Molineri, C. &amp; Salles, F. F. (2017) Review of selected species of Campsurus Eaton 1868 (Ephemeroptera: Polymitarcyidae), with description of eleven new species and a key to male imagos of the genus. Zootaxa, 4300 (3), 301 - 354. https: // doi. org / 10.11646 / zootaxa. 4300.3.1" type="journal article" year="2017">Molineri &amp; Salles (2017)</bibRefCitation>
. In order to add the new species to that key, and also to include the recently described 
<taxonomicName id="28A381C5F9466429FC57F8DAFF33F812" authority="Cruz, Molineri &amp; Hamada, 2022" authorityName="Cruz, Molineri &amp; Hamada" authorityYear="2022" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="fortuitus">
<emphasis id="DDD72654F9466429FC57F8DAFB88F8FE" box="[1002,1129,1884,1911]" italics="true" pageId="4" pageNumber="149">C. fortuitus</emphasis>
Cruz, Molineri &amp; Hamada, 2022
</taxonomicName>
, which would also key out in the same couplet, we propose the following change:
</paragraph>
<paragraph id="EF1CFA46F9466428FF2AF842FA7CFE8C" blockId="4.[151,1437,1989,2041]" lastBlockId="5.[151,1437,151,261]" lastPageId="5" lastPageNumber="150" pageId="4" pageNumber="149">
3(2) Forewing smaller than 
<quantity id="285B57A3F9466429FE7DF841FE16F853" box="[448,503,1990,2011]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="149" unit="mm" value="7.0">7 mm</quantity>
............................................................................ 4 3’. Forewing larger than 
<quantity id="285B57A3F9466429FE0CF864FE09F870" box="[433,488,2019,2040]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="149" unit="mm" value="9.0">9 mm</quantity>
................................................ couplet 4 of 
<bibRefCitation id="8B3287B7F9466429FB17F864FA7CF870" author="Molineri, C. &amp; Salles, F. F." box="[1194,1437,2019,2041]" pageId="4" pageNumber="149" pagination="301 - 354" refId="ref2691" refString="Molineri, C. &amp; Salles, F. F. (2017) Review of selected species of Campsurus Eaton 1868 (Ephemeroptera: Polymitarcyidae), with description of eleven new species and a key to male imagos of the genus. Zootaxa, 4300 (3), 301 - 354. https: // doi. org / 10.11646 / zootaxa. 4300.3.1" type="journal article" year="2017">Molineri &amp; Salles (2017)</bibRefCitation>
4(3). Penes blade-like, not curved; posterior margin of sternum IX slightly concave..................................... 5 4’. Penes curved ventrally; posterior margin of sternum IX triangular at middle.......................... 
<taxonomicName id="28A381C5F9476428FB49FF32FAA8FF42" authority="Pantoja &amp; Viana &amp; Salles, 2023" authorityName="Pantoja &amp; Viana &amp; Salles" authorityYear="2023" box="[1268,1353,181,202]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="mirim" status="sp. nov.">
<emphasis id="DDD72654F9476428FB49FF32FAA8FF42" box="[1268,1353,181,202]" italics="true" pageId="5" pageNumber="150">C. mirim</emphasis>
</taxonomicName>
<emphasis id="DDD72654F9476428FAEDFF32FA7DFF42" bold="true" box="[1360,1436,181,202]" pageId="5" pageNumber="150">
<taxonomicNameLabel id="C6E49B2FF9476428FAEDFF32FA7DFF42" box="[1360,1436,181,202]" pageId="5" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
5(4). Penes with inner distal margin of main lobe curved outwards........................................... 
<taxonomicName id="28A381C5F9476428FA92FF55FA7DFF6F" authorityName="Cruz, Molineri &amp; Hamada" authorityYear="2022" box="[1327,1436,210,231]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="fortuitus">
<emphasis id="DDD72654F9476428FA92FF55FA7DFF6F" box="[1327,1436,210,231]" italics="true" pageId="5" pageNumber="150">C. fortuitus</emphasis>
</taxonomicName>
5’. Penes with distal margin of main lobe rounded, not curved............................................. 
<taxonomicName id="28A381C5F9476428FA83FF77FA7CFE8C" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1342,1437,239,261]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="povilla">
<emphasis id="DDD72654F9476428FA83FF77FA7CFE8C" box="[1342,1437,239,261]" italics="true" pageId="5" pageNumber="150">C. povilla</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>