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<document id="2212D84A4972F8E30E80FE769B2C74CD" ID-DOI="10.5070/P939257077" ID-ISSN="0031-0298" ID-Zenodo-Dep="13743800" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726019612735" checkinUser="felipe" docAuthor="Squires, Richard L." docDate="2022" docId="C96187CFFFBAEB45FC42FC44FBF4FC74" docLanguage="en" docName="PaleoBios.39.2.1-11.pdf" docOrigin="PaleoBios 39 (2)" docSource="http://dx.doi.org/10.5070/p939257077" docStyle="DocumentStyle:0E19E0F9495DF1C9BA1290CB4A0EE6FA.1:PaleoBios.2022.journal_article" docStyleId="0E19E0F9495DF1C9BA1290CB4A0EE6FA" docStyleName="PaleoBios.2022.journal_article" docStyleVersion="1" docTitle="Ancistrolepis carolineae Squires 1984" docType="treatment" docVersion="2" lastPageNumber="7" masterDocId="3558FFB7FFBCEB4DFFDFFF82FFBAFFE9" masterDocTitle="The earliest Ancistrolepis (Gastropoda: Buccinidae) and its geologic implications" masterLastPageNumber="11" masterPageNumber="1" pageNumber="5" updateTime="1726020633084" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="AEC057125D5BB3671F75BF89691446DB">The earliest Ancistrolepis (Gastropoda: Buccinidae) and its geologic implications</mods:title>
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<mods:namePart id="6826FBFFAB23939AEA2EBFA579D7846B">Squires, Richard L.</mods:namePart>
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<mods:date id="4EC199952647E24E70CBD4584B020B7B">2022</mods:date>
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<mods:identifier id="0F24F0C0AEC8450639C8018C34A39106" type="ISSN">0031-0298</mods:identifier>
<mods:identifier id="F85D6633E1E8EE2D46B5C9C8609F2CAC" type="Zenodo-Dep">13743800</mods:identifier>
</mods:mods>
<treatment id="C96187CFFFBAEB45FC42FC44FBF4FC74" LSID="urn:lsid:plazi:treatment:C96187CFFFBAEB45FC42FC44FBF4FC74" httpUri="http://treatment.plazi.org/id/C96187CFFFBAEB45FC42FC44FBF4FC74" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5">
<subSubSection id="09D26552FFBAEB4BFC42FC44FA2EFC37" box="[925,1428,966,990]" pageId="6" pageNumber="5" type="nomenclature">
<paragraph id="417736D9FFBAEB4BFC42FC44FA2EFC37" blockId="6.[925,1428,966,1026]" box="[925,1428,966,990]" pageId="6" pageNumber="5">
<heading id="1A3F81B5FFBAEB4BFC42FC44FA2EFC37" allCaps="true" box="[925,1428,966,990]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="5" reason="2">
<taxonomicName id="86C84D5AFFBAEB4BFC42FC44FA2EFC37" authority="SQUIRES, 1984" authorityName="Squires" authorityYear="1984" box="[925,1428,966,990]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="6" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFBAEB4BFC42FC44FB56FC34" box="[925,1260,966,989]" italics="true" pageId="6" pageNumber="5">ANCISTROLEPIS CAROLINEAE</emphasis>
<bibRefCitation id="25594B28FFBAEB4BFB2DFC44FA2EFC37" author="Squires, R. L." box="[1266,1428,966,990]" pageId="6" pageNumber="5" pagination="1 - 76" refId="ref9091" refString="Squires, R. L. 1984. Megapaleontology of the Eocene Llajas Formation, Simi Valley, California. Natural History Museum of Los Angeles County Contributions in Science 350: 1 - 76." type="journal article" year="1984">SQUIRES, 1984</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="09D26552FFBAEB4BFBBFFC6AFB68FBEB" box="[1120,1234,1000,1026]" pageId="6" pageNumber="5" type="description">
<paragraph id="417736D9FFBAEB4BFBBFFC6AFB68FBEB" blockId="6.[925,1428,966,1026]" box="[1120,1234,1000,1026]" pageId="6" pageNumber="5">
<figureCitation id="D9F32A5CFFBAEB4BFBBFFC6AFB68FBEB" box="[1120,1234,1000,1026]" captionStart="Figure 4" captionStartId="7.[112,199,1165,1191]" captionTargetBox="[163,1469,216,1118]" captionTargetId="figure-418@7.[141,1487,193,1145]" captionTargetPageId="7" captionText="Figure 4AI. Ancistrolepis carolineae Squires, 1984. “Stewart bed,” Llajas Formation, north side of Simi Valley, Ventura County, southern California. Apertural (A) and abapertural (B) views, 1.7x, length 32 mm, width 20 mm, plasto-holotype, LACMIP 10554 [=UCLA 59401], Locality LACMIP 7242.35. Apertural (C) and abapertural (D) views, x1.8x, length 28.8 mm, width 18.4 mm, hypotype, LACMIP 14904, LACMIP Locality 40444.2. Apertural (E) view, 1.8x, length 26.3, width 14.9, hypotype, LACMIP 14905, LACMIP 40374.75. Apertural (F) and abapertural (G) views, x2.3, length 22 mm, width 15.8 mm, hypotype, LACMIP 14906, LACMIP Locality 40374.76. Right-lateral apertural, with aperture end tilted up slightly (H) and oblique lower columellar (I) views, x2.5, length 28.5 mm, width 29.8 mm, hypotype, LACMIP 14907, LACMIP Locality 40374.77." figureDoi="http://doi.org/10.5281/zenodo.13743804" httpUri="https://zenodo.org/record/13743804/files/figure.png" pageId="6" pageNumber="5">FIG. 4AI</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="09D26552FFBAEB4BFC9EFBA9FB64FB81" pageId="6" pageNumber="5" type="reference_group">
<paragraph id="417736D9FFBAEB4BFC9EFBA9FB64FB81" blockId="6.[833,1521,1067,1938]" pageId="6" pageNumber="5">
<taxonomicName id="86C84D5AFFBAEB4BFC9EFBA9FA9BFBAC" authority="Squires, 1984" authorityName="Squires" authorityYear="1984" box="[833,1313,1067,1093]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="5" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFBAEB4BFC9EFBA9FC64FBAC" box="[833,990,1067,1093]" italics="true" pageId="6" pageNumber="5">Ancistrolepis</emphasis>
?
<emphasis id="73BCEACBFFBAEB4BFC2FFBA9FBCAFBAC" box="[1008,1136,1067,1093]" italics="true" pageId="6" pageNumber="5">carolineae</emphasis>
<bibRefCitation id="25594B28FFBAEB4BFBA9FBA9FA9BFBAC" author="Squires, R. L." box="[1142,1313,1067,1093]" pageId="6" pageNumber="5" pagination="1 - 76" refId="ref9091" refString="Squires, R. L. 1984. Megapaleontology of the Eocene Llajas Formation, Simi Valley, California. Natural History Museum of Los Angeles County Contributions in Science 350: 1 - 76." type="journal article" year="1984">Squires, 1984</bibRefCitation>
</taxonomicName>
. p. 30, figs. 8de. Groves and Squires. 2021. p. 191.
</paragraph>
</subSubSection>
<subSubSection id="09D26552FFBAEB4BFCBEFBF3FA4AFA21" pageId="6" pageNumber="5" type="materials_examined">
<paragraph id="417736D9FFBAEB4BFCBEFBF3FA4AFA21" blockId="6.[833,1521,1067,1938]" pageId="6" pageNumber="5">
<emphasis id="73BCEACBFFBAEB4BFCBEFBF3FB09FB62" bold="true" box="[865,1203,1137,1163]" pageId="6" pageNumber="5">Primary type material—</emphasis>
<materialsCitation id="F1A03C84FFBAEB4BFB6CFBF3FA1DFB46" collectionCode="LACMIP" pageId="6" pageNumber="5" specimenCode="LACMIP 10554, UCLA 59401" specimenCount="1" typeStatus="holotype">
<typeStatus id="9E73887BFFBAEB4BFB6CFBF3FA9CFB62" box="[1203,1318,1137,1163]" pageId="6" pageNumber="5" type="holotype">Holotype</typeStatus>
<specimenCode id="116E9EA2FFBAEB4BFAEFFBF3FA4AFB65" box="[1328,1520,1137,1164]" collectionCode="LACMIP" pageId="6" pageNumber="5">LACMIP 10554</specimenCode>
[originally
<specimenCode id="116E9EA2FFBAEB4BFC1AFB16FBD8FB46" box="[965,1122,1172,1199]" collectionCode="UCLA" country="USA" httpUri="http://grbio.org/cool/irpn-5wg7" name="University of California at Los Angeles" pageId="6" pageNumber="5">UCLA 59401</specimenCode>
],
<collectionCode id="27D9AE1CFFBAEB4BFBADFB16FB6FFB47" box="[1138,1237,1172,1198]" pageId="6" pageNumber="5">LACMIP</collectionCode>
Locality 7242.35
</materialsCitation>
.
<materialsCitation id="F1A03C84FFBAEB4BFA70FB17FB8DFB3B" collectionCode="LACMIP" pageId="6" pageNumber="5" specimenCode="LACMIP 10555" specimenCount="1" typeStatus="paratype">
<typeStatus id="9E73887BFFBAEB4BFA70FB17FCC2FB3B" pageId="6" pageNumber="5" type="paratype">Paratype</typeStatus>
<specimenCode id="116E9EA2FFBAEB4BFCA2FB3AFB8DFB3B" box="[893,1079,1207,1234]" collectionCode="LACMIP" pageId="6" pageNumber="5">LACMIP 10555</specimenCode>
</materialsCitation>
,
<materialsCitation id="F1A03C84FFBAEB4BFB9FFB3AFA4AFB3B" box="[1088,1520,1207,1234]" collectionCode="LACMIP" collectorName="Locality" pageId="6" pageNumber="5" specimenCount="1" typeStatus="paratype">
<collectionCode id="27D9AE1CFFBAEB4BFB9FFB3AFB19FB3B" box="[1088,1187,1208,1234]" pageId="6" pageNumber="5">LACMIP</collectionCode>
<collectorName id="EC3D530FFFBAEB4BFB77FB35FAB0FB38" box="[1192,1290,1207,1233]" pageId="6" pageNumber="5">Locality</collectorName>
7242.36;
<typeStatus id="9E73887BFFBAEB4BFA5EFB3AFA4AFB3B" box="[1409,1520,1208,1234]" pageId="6" pageNumber="5" type="paratype">paratype</typeStatus>
</materialsCitation>
<materialsCitation id="F1A03C84FFBAEB4BFC9EFB59FC46FB1C" box="[833,1020,1242,1269]" collectionCode="LACMIP" pageId="6" pageNumber="5" specimenCode="LACMIP 10556" specimenCount="1" typeStatus="paratype">
<specimenCode id="116E9EA2FFBAEB4BFC9EFB59FC46FB1C" box="[833,1020,1242,1269]" collectionCode="LACMIP" pageId="6" pageNumber="5">LACMIP 10556</specimenCode>
</materialsCitation>
,
<materialsCitation id="F1A03C84FFBAEB4BFBD8FB59FA4AFB1C" box="[1031,1520,1243,1269]" collectionCode="LACMIP" collectorName="Locality" pageId="6" pageNumber="5" specimenCount="1" typeStatus="paratype">
<collectionCode id="27D9AE1CFFBAEB4BFBD8FB59FBD1FB1C" box="[1031,1131,1243,1269]" pageId="6" pageNumber="5">LACMIP</collectionCode>
<collectorName id="EC3D530FFFBAEB4BFBAFFB59FB69FB1C" box="[1136,1235,1243,1269]" pageId="6" pageNumber="5">Locality</collectorName>
7242.37; and
<typeStatus id="9E73887BFFBAEB4BFA5FFB59FA4AFB1C" box="[1408,1520,1243,1269]" pageId="6" pageNumber="5" type="paratype">paratype</typeStatus>
</materialsCitation>
<materialsCitation id="F1A03C84FFBAEB4BFC9EFB7CFC47FAF0" box="[833,1021,1278,1305]" collectionCode="LACMIP" pageId="6" pageNumber="5" specimenCode="LACMIP 10557" specimenCount="1" typeStatus="paratype">
<specimenCode id="116E9EA2FFBAEB4BFC9EFB7CFC47FAF0" box="[833,1021,1278,1305]" collectionCode="LACMIP" pageId="6" pageNumber="5">LACMIP 10557</specimenCode>
</materialsCitation>
,
<materialsCitation id="F1A03C84FFBAEB4BFBD7FB7CFAF0FAD2" collectionCode="LACMIP" collectorName="Locality &amp; These" pageId="6" pageNumber="5" specimenCount="3" typeStatus="paratype">
<collectionCode id="27D9AE1CFFBAEB4BFBD7FB7CFBD6FAF1" box="[1032,1132,1278,1304]" pageId="6" pageNumber="5">LACMIP</collectionCode>
<collectorName id="EC3D530FFFBAEB4BFBAEFB7CFB6EFAF1" box="[1137,1236,1278,1304]" pageId="6" pageNumber="5">Locality</collectorName>
22312.14.
<collectorName id="EC3D530FFFBAEB4BFA82FB7CFA12FAF1" box="[1373,1448,1278,1304]" pageId="6" pageNumber="5">These</collectorName>
<specimenCount id="57CEFD50FFBAEB4BFA71FB7CFC07FAD5" count="3" pageId="6" pageNumber="5" type="generic" typeStatus="paratype">three paratypes</specimenCount>
were originally referred to as lot
</materialsCitation>
<materialsCitation id="F1A03C84FFBAEB4BFA90FAA3FA51FAD5" box="[1359,1515,1313,1340]" collectionCode="UCLA" pageId="6" pageNumber="5" specimenCode="UCLA 59402" specimenCount="1" typeStatus="paratype">
<specimenCode id="116E9EA2FFBAEB4BFA90FAA3FA51FAD5" box="[1359,1515,1313,1340]" collectionCode="UCLA" country="USA" httpUri="http://grbio.org/cool/irpn-5wg7" name="University of California at Los Angeles" pageId="6" pageNumber="5">UCLA 59402</specimenCode>
</materialsCitation>
, which was, at some point in the past, split into the three lots listed here.
<materialsCitation id="F1A03C84FFBAEB4BFBD3FAEAFA4AFA6B" box="[1036,1520,1383,1410]" collectionCode="LACMIP" collectorName="Locality" pageId="6" pageNumber="5" specimenCount="1" typeStatus="paratype">
<collectionCode id="27D9AE1CFFBAEB4BFBD3FAEAFBCAFA6B" box="[1036,1136,1384,1410]" pageId="6" pageNumber="5">LACMIP</collectionCode>
<collectorName id="EC3D530FFFBAEB4BFBA6FAE5FB66FA68" box="[1145,1244,1383,1409]" pageId="6" pageNumber="5">Locality</collectorName>
7242 is equivalent to
</materialsCitation>
<collectionCode id="27D9AE1CFFBAEB4BFC9EFA09FC3CFA4C" box="[833,902,1419,1445]" country="USA" httpUri="http://grbio.org/cool/irpn-5wg7" name="University of California at Los Angeles" pageId="6" pageNumber="5">UCLA</collectionCode>
<materialsCitation id="F1A03C84FFBAEB4BFC55FA09FA4AFA21" collectingDate="1946-04-19" collectionCode="UCLA" collectorName="Locality &amp; W. P. Popenoe" pageId="6" pageNumber="5" specimenCount="1" typeStatus="paratype">
<collectorName id="EC3D530FFFBAEB4BFC55FA09FC51FA4C" box="[906,1003,1419,1445]" pageId="6" pageNumber="5">Locality</collectorName>
22312 (originally
<collectionCode id="27D9AE1CFFBAEB4BFB19FA09FAB0FA4C" box="[1222,1290,1419,1445]" country="USA" httpUri="http://grbio.org/cool/irpn-5wg7" name="University of California at Los Angeles" pageId="6" pageNumber="5">UCLA</collectionCode>
<collectorName id="EC3D530FFFBAEB4BFAD0FA09FACAFA4C" box="[1295,1392,1419,1445]" pageId="6" pageNumber="5">Locality</collectorName>
2312 (collected by
<collectorName id="EC3D530FFFBAEB4BFC6CFA2CFBEEFA21" box="[947,1108,1454,1480]" pageId="6" pageNumber="5">W.P.Popenoe</collectorName>
classes,
<date id="35761019FFBAEB4BFB66FA2CFADCFA21" box="[1209,1382,1454,1480]" pageId="6" pageNumber="5" value="1946-04-19">
<collectingDate id="2532E9F1FFBAEB4BFB66FA2CFADCFA21" box="[1209,1382,1454,1480]" pageId="6" pageNumber="5" value="1946-04-19">April 19, 1946</collectingDate>
</date>
and 1950).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="09D26552FFBAEB45FCBEFA53FD3EFECA" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5" type="description">
<paragraph id="417736D9FFBAEB4BFCBEFA53FC2FF9D8" blockId="6.[833,1521,1067,1938]" pageId="6" pageNumber="5">
<emphasis id="73BCEACBFFBAEB4BFCBEFA53FB6AFA02" bold="true" box="[865,1232,1489,1515]" pageId="6" pageNumber="5">
Dimensions of
<typeStatus id="9E73887BFFBAEB4BFBE8FA53FB15FA02" box="[1079,1199,1489,1515]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
</emphasis>
Length
<quantity id="86309B3CFFBAEB4BFAE9FA53FA2FFA05" box="[1334,1429,1489,1516]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="5" unit="mm" value="32.0">32 mm</quantity>
, width
<quantity id="86309B3CFFBAEB4BFC9EFA76FC20F9E6" box="[833,922,1524,1551]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="5" unit="mm" value="20.0">20 mm</quantity>
(specimen is missing most its anterior siphonal canal).
</paragraph>
<paragraph id="417736D9FFBAEB4BFCBEF9B8FA90F90B" blockId="6.[833,1521,1067,1938]" pageId="6" pageNumber="5">
<emphasis id="73BCEACBFFBAEB4BFCBEF9B8FB07F9BD" bold="true" box="[865,1213,1594,1621]" pageId="6" pageNumber="5">
Secondary
<typeStatus id="9E73887BFFBAEB4BFC2FF9B9FB90F9BC" box="[1008,1066,1595,1621]" pageId="6" pageNumber="5">type</typeStatus>
material—
</emphasis>
LACMIP hypotype 14904, LACMIP Locality 40444.2; LACMIP hypotype 14905, LACMIP Locality 40374.75; LACMIP hypotype 14906, LACMIP Locality 40374.76; and LACMIP hypotype
<materialsCitation id="F1A03C84FFBAEB4BFA6BF926FA9CF90B" collectionCode="LACMIP" pageId="6" pageNumber="5" specimenCode="LACMIP 14907" specimenCount="1" typeStatus="holotype">
<specimenCode id="116E9EA2FFBAEB4BFA6BF926FC76F90B" collectionCode="LACMIP" pageId="6" pageNumber="5">LACMIP 14907</specimenCode>
,
<collectionCode id="27D9AE1CFFBAEB4BFC07F94AFB86F90B" box="[984,1084,1736,1762]" pageId="6" pageNumber="5">LACMIP</collectionCode>
Locality 40374.77
</materialsCitation>
.
</paragraph>
<paragraph id="417736D9FFBAEB4BFCBEF969FA4AF87B" blockId="6.[833,1521,1067,1938]" pageId="6" pageNumber="5">
<emphasis id="73BCEACBFFBAEB4BFCBEF969FA99F8ED" bold="true" box="[865,1315,1770,1797]" pageId="6" pageNumber="5">Number of specimens examined—</emphasis>
<materialsCitation id="F1A03C84FFBAEB4BFAFCF969FA4AF87B" county="Ventura County" location="Llajas Formation" municipality="All" pageId="6" pageNumber="5" specimenCount="17" stateProvince="California">
<specimenCount id="57CEFD50FFBAEB4BFAFCF969FC3DF8C1" count="17" pageId="6" pageNumber="5" type="generic">Seventeen specimens</specimenCount>
.
<collectingMunicipality id="A113ACA3FFBAEB4BFC4DF88CFC0EF8C1" box="[914,948,1806,1832]" pageId="6" pageNumber="5">All</collectingMunicipality>
are from four localities (7242, 40374, 40444, and 40479) in the “Stewart bed”of the
<location id="44176002FFBAEB4BFAC1F8B3FA4AF8A2" LSID="urn:lsid:plazi:treatment:C96187CFFFBAEB45FC42FC44FBF4FC74:44176002FFBAEB4BFAC1F8B3FA4AF8A2" box="[1310,1520,1841,1867]" county="Ventura County" municipality="All" name="Llajas Formation" pageId="6" pageNumber="5" stateProvince="California">Llajas Formation</location>
on the north side of
<location id="44176002FFBAEB4BFBE0F8D6FB72F887" LSID="urn:lsid:plazi:treatment:C96187CFFFBAEB45FC42FC44FBF4FC74:44176002FFBAEB4BFBE0F8D6FB72F887" box="[1087,1224,1876,1902]" county="Ventura County" municipality="All" name="Simi Valley" pageId="6" pageNumber="5" stateProvince="California">Simi Valley</location>
,
<collectingCounty id="A8164E55FFBAEB4BFB0DF8D7FA2FF887" box="[1234,1429,1876,1903]" pageId="6" pageNumber="5">Ventura County</collectingCounty>
, southern
<collectingRegion id="830CF83BFFBAEB4BFCAEF8F5FC54F878" box="[881,1006,1911,1937]" country="United States of America" name="California" pageId="6" pageNumber="5">California</collectingRegion>
.
<location id="44176002FFBAEB4BFC28F8F5FBF0F878" LSID="urn:lsid:plazi:treatment:C96187CFFFBAEB45FC42FC44FBF4FC74:44176002FFBAEB4BFC28F8F5FBF0F878" box="[1015,1098,1911,1937]" county="Ventura County" municipality="All" name="Nearly" pageId="6" pageNumber="5" stateProvince="California">Nearly</location>
all the specimens are from the
<location id="44176002FFBAEB4BFA1AF8FAFA4AF87B" LSID="urn:lsid:plazi:treatment:C96187CFFFBAEB45FC42FC44FBF4FC74:44176002FFBAEB4BFA1AF8FAFA4AF87B" box="[1477,1520,1912,1938]" county="Ventura County" municipality="All" name="Las" pageId="6" pageNumber="5" stateProvince="California">Las</location>
</materialsCitation>
</paragraph>
<caption id="15B76651FFBBEB4AFFAFFB0FFBAEFA8B" ID-DOI="http://doi.org/10.5281/zenodo.13743804" ID-Zenodo-Dep="13743804" httpUri="https://zenodo.org/record/13743804/files/figure.png" pageId="7" pageNumber="6" startId="7.[112,199,1165,1191]" targetBox="[163,1469,216,1118]" targetPageId="7" targetType="figure">
<paragraph id="417736D9FFBBEB4AFFAFFB0FFBAEFA8B" blockId="7.[112,1512,1165,1378]" pageId="7" pageNumber="6">
<emphasis id="73BCEACBFFBBEB4AFFAFFB0FFEABFB4E" bold="true" box="[112,273,1165,1191]" pageId="7" pageNumber="6">Figure 4AI.</emphasis>
<taxonomicName id="86C84D5AFFBBEB4AFEC8FB0FFD57FB4E" authority="Squires, 1984" authorityName="Squires" authorityYear="1984" box="[279,749,1165,1191]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="6" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFBBEB4AFEC8FB0FFD81FB4E" box="[279,571,1165,1191]" italics="true" pageId="7" pageNumber="6">Ancistrolepis carolineae</emphasis>
<bibRefCitation id="25594B28FFBBEB4AFD9DFB0FFD57FB4E" author="Squires, R. L." box="[578,749,1165,1191]" pageId="7" pageNumber="6" pagination="1 - 76" refId="ref9091" refString="Squires, R. L. 1984. Megapaleontology of the Eocene Llajas Formation, Simi Valley, California. Natural History Museum of Los Angeles County Contributions in Science 350: 1 - 76." type="journal article" year="1984">Squires, 1984</bibRefCitation>
</taxonomicName>
. “Stewart bed,” Llajas Formation, north side of Simi Valley, Ventura County, southern California. Apertural (
<emphasis id="73BCEACBFFBBEB4AFD1AFB25FD62FB28" bold="true" box="[709,728,1191,1217]" pageId="7" pageNumber="6">A</emphasis>
) and abapertural (
<emphasis id="73BCEACBFFBBEB4AFC1AFB2AFC62FB2B" bold="true" box="[965,984,1192,1218]" pageId="7" pageNumber="6">B</emphasis>
) views, 1.7x, length 32 mm, width 20 mm, plasto-holotype, LACMIP 10554 [=UCLA 59401], Locality LACMIP 7242.35. Apertural (
<emphasis id="73BCEACBFFBBEB4AFB7AFB40FB0CFB35" bold="true" box="[1189,1206,1218,1244]" pageId="7" pageNumber="6">C</emphasis>
) and abapertural (
<emphasis id="73BCEACBFFBBEB4AFA7DFB40FA0DFB35" bold="true" box="[1442,1463,1218,1244]" pageId="7" pageNumber="6">D</emphasis>
) views, x1.8x, length 28.8 mm, width 18.4 mm, hypotype, LACMIP 14904, LACMIP Locality 40444.2. Apertural (
<emphasis id="73BCEACBFFBBEB4AFA13FB5FFA67FB1E" bold="true" box="[1484,1501,1245,1271]" pageId="7" pageNumber="6">E</emphasis>
) view, 1.8x, length 26.3, width 14.9, hypotype, LACMIP 14905, LACMIP 40374.75. Apertural (
<emphasis id="73BCEACBFFBBEB4AFB38FB7AFB4DFAFB" bold="true" box="[1255,1271,1272,1298]" pageId="7" pageNumber="6">F</emphasis>
) and abapertural (
<emphasis id="73BCEACBFFBBEB4AFFA4FA90FF34FAC5" bold="true" box="[123,142,1298,1324]" pageId="7" pageNumber="6">G</emphasis>
) views, x2.3, length 22 mm, width 15.8 mm, hypotype, LACMIP 14906, LACMIP Locality 40374.76. Right-lateral apertural, with aperture end tilted up slightly (
<emphasis id="73BCEACBFFBBEB4AFD32FAAFFCB8FAAE" bold="true" box="[749,770,1325,1351]" pageId="7" pageNumber="6">H</emphasis>
) and oblique lower columellar (
<emphasis id="73BCEACBFFBBEB4AFB4AFAAFFB25FAAE" bold="true" box="[1173,1183,1325,1351]" pageId="7" pageNumber="6">I</emphasis>
) views, x2.5, length 28.5 mm, width 29.8 mm, hypotype, LACMIP 14907, LACMIP Locality 40374.77.
</paragraph>
</caption>
<paragraph id="417736D9FFBBEB4AFFAFFA02FA4AF9A3" blockId="7.[112,1520,1408,1962]" pageId="7" pageNumber="6">
Llajas Canyon area: LACMIP Localities 7242 and 40374, ribs on ante-penultimate whorl, 3 ribs on penultimate which are in the immediate vicinity of each another, and whorl, and 6 (on most specimens) to 8 (rarely) ribs on Locality 40444, which is a short distance northwest- last whorl. Several widely spaced secondary spiral ribs wardly. The one exception is a specimen from LACMIP can be present inside of surface of apertural area and Locality 40479, approximately
<quantity id="86309B3CFFBBEB4AFDDDF98FFDFEF9CF" box="[514,580,1548,1575]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="6" unit="km" value="3.0">3 km</quantity>
westardly of the represent continuations of primary ribs on main surface localities in Las Llajas Canyon. of last whorl. Spacing of spirals usually consistent at 3
</paragraph>
<paragraph id="417736D9FFBBEB45FF4FF9D1FD3EFECA" blockId="7.[112,1520,1408,1962]" lastBlockId="8.[112,801,195,1911]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<emphasis id="73BCEACBFFBBEB4AFF4FF9D1FEF9F984" bold="true" box="[144,323,1619,1645]" pageId="7" pageNumber="6">Description</emphasis>
Shell ovate fusiform, shell very thin mm apart but can be as much as
<quantity id="86309B3CFFBBEB4AFB05F9D1FA98F984" box="[1242,1314,1619,1645]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="6" unit="mm" value="4.0">4 mm</quantity>
apart, even on a (“paper thin”), up to
<quantity id="86309B3CFFBBEB4AFE50F9F4FDBFF978" box="[399,517,1654,1681]" metricMagnitude="-2" metricUnit="m" metricValue="3.04" pageId="7" pageNumber="6" unit="mm" value="30.4">30.4 mm</quantity>
height and
<quantity id="86309B3CFFBBEB4AFD75F9F4FCA5F978" box="[682,799,1654,1681]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="7" pageNumber="6" unit="mm" value="17.7">17.7 mm</quantity>
single specimen. Spiral ribs beveled (not square-edged); width, with corresponding h/w = 1.72; average h/w of interstitally (especially on columellar inner lip) with five all specimens = 1.59. Protoconch/shell apex missing, bands of spiral microscopic threads alternating with upper spire incomplete on most specimens. Teleoconch bands of five spiral submicroscopic threads; comprising up to 5 whorls; commonly only four teleoconch whorls a grand total of 25 threads. Columella has almost none present. Suture impressed and coincident with spiral to only slight twist to left; no columellar teeth; anterior rib. Only spiral ribs (widely spaced); no radial sculpture. part of columella can have a few intermediate-strength Earliest two teleoconch whorls smooth (ribs apparently ribs, becoming obsolete anteriorward and interiorward. eroded off), 2 ribs on pre-ante penultimate whorl, 2 Siphonal canal moderately long. Outer lip “terminal varix,”
<quantity id="86309B3CFFB4EB45FF19FF41FEAAFF34" box="[198,272,195,221]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="7" unit="mm" value="3.0">3 mm</quantity>
thick, smooth, solid, and flattish but with beveled on both outermost and innermost edges; minute threads axially parallel to edge of outer lip.
</paragraph>
</subSubSection>
<subSubSection id="09D26552FFB4EB45FF4FFEAFFBF4FC74" pageId="8" pageNumber="7" type="discussion">
<paragraph id="417736D9FFB4EB45FF4FFEAFFDDBFD89" blockId="8.[112,801,195,1911]" pageId="8" pageNumber="7">
<emphasis id="73BCEACBFFB4EB45FF4FFEAFFE9FFEAE" bold="true" box="[144,293,301,327]" pageId="8" pageNumber="7">Remarks—</emphasis>
The
<typeStatus id="9E73887BFFB4EB45FE86FEAEFE7CFEAF" box="[345,454,300,326]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
has been missing for several decades. A copy of a Kodrachrome 35-mm slide of the
<typeStatus id="9E73887BFFB4EB45FFAFFEF1FF5AFE64" box="[112,224,371,397]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
, taken by the author 30 years ago, is shown on the “Cover” of this present paper. The specimens of
<taxonomicName id="86C84D5AFFB4EB45FCD6FE14FF55FE3A" authorityName="Squires" authorityYear="1984" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFB4EB45FCD6FE14FCA0FE59" box="[777,794,406,432]" italics="true" pageId="8" pageNumber="7">A</emphasis>
.
<emphasis id="73BCEACBFFB4EB45FFAFFE3BFF55FE3A" box="[112,239,441,467]" italics="true" pageId="8" pageNumber="7">carolineae</emphasis>
</taxonomicName>
all have moderately poor to poor preservation because of weathering, which has caused some of the shells, which had thin shells to begin with, to become to steinkerns or nearly so. Weathering has probably caused the spiral ribs to be somewhat rounded.
</paragraph>
<paragraph id="417736D9FFB4EB45FF4FFDE8FE76FBC3" blockId="8.[112,801,195,1911]" pageId="8" pageNumber="7">
Four of the 17 Llajas Formation specimens of
<taxonomicName id="86C84D5AFFB4EB45FD1CFDE8FF00FD4F" authorityName="Squires" authorityYear="1984" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFB4EB45FD1CFDE8FF00FD4F" italics="true" pageId="8" pageNumber="7">A. carolineae</emphasis>
</taxonomicName>
have an obvious, relatively wide, flat, and sturdy thickening (callus) of the outer lip (terminal varix) of the aperture. A search of the available literature revealed that no other ancistrolepine (fossil or extant) has such a wide and flat outer lip callus, although a few extant species can have a very thin, narrow ridge callus. A terminal “varix” is common in warm, shallow-marine gastropods, but is rare in polar and deep-marine habitats (
<bibRefCitation id="25594B28FFB4EB45FD67FC01FEE8FC29" author="Webster, N. B. &amp; G. J. Vermeij" pageId="8" pageNumber="7" pagination="732 - 754" refId="ref9758" refString="Webster, N. B. and G. J. Vermeij. 2017. The varix: evolution, distribution, and phylogenetic clumping of a repeated gastropod innovation. Zoological Journal of the Linnean Society 180: 732 - 754." type="journal article" year="2017">Webster and Vermeij 2017</bibRefCitation>
), therefore, the presence of this feature on
<taxonomicName id="86C84D5AFFB4EB45FF10FC48FED6FC0A" authorityName="Squires" authorityYear="1984" box="[207,364,969,996]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFB4EB45FF10FC48FED6FC0A" box="[207,364,969,996]" italics="true" pageId="8" pageNumber="7">A. carolineae</emphasis>
</taxonomicName>
might be another indication that
<taxonomicName id="86C84D5AFFB4EB45FCD6FC48FF4AFBEF" authorityName="Squires" authorityYear="1984" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFB4EB45FCD6FC48FF4AFBEF" italics="true" pageId="8" pageNumber="7">A. carolineae</emphasis>
</taxonomicName>
lived in an environment quite different than the extant
<emphasis id="73BCEACBFFB4EB45FF2BFB92FE3CFBC3" box="[244,390,1040,1066]" italics="true" pageId="8" pageNumber="7">Ancistrolepi</emphasis>
s spp.
</paragraph>
<paragraph id="417736D9FFB4EB45FF4FFBB1FE57F9FF" blockId="8.[112,801,195,1911]" pageId="8" pageNumber="7">
<taxonomicName id="86C84D5AFFB4EB45FF4FFBB1FE0BFBA4" authorityName="Squires" authorityYear="1984" box="[144,433,1075,1101]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFB4EB45FF4FFBB1FE0BFBA4" box="[144,433,1075,1101]" italics="true" pageId="8" pageNumber="7">Ancistrolepis carolineae</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="86C84D5AFFB4EB45FD08FBB1FE4EFB99" authority="Titova, 1993" authorityName="Titova" authorityYear="1993" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="rategiensis">
<emphasis id="73BCEACBFFB4EB45FD08FBB1FEEFFB99" italics="true" pageId="8" pageNumber="7">Ancistrolepis rategiensis</emphasis>
<bibRefCitation id="25594B28FFB4EB45FE85FBD4FE4EFB99" author="Titova, L. V." box="[346,500,1110,1136]" pageId="8" pageNumber="7" pagination="1 - 16" refId="ref9577" refString="Titova, L. V. 1993. The early history of the North Pacific Ancistrolepidinae (Gastropoda: Buccinidae). Ruthenica 3: 1 - 16." type="book chapter" year="1993">Titova, 1993</bibRefCitation>
</taxonomicName>
(p. 12, figs. 2A2D) from the upper Eocene Rategian Formation at Podkagernaya Bay, northwestern Kamchakta. The Llajas Formation species is similar in its ovate-fusiform shell shape, approximately five rounded whorls, rounded base on last whorl, subtriangular to triangular spiral ribs, about seven spiral ribs (can be variously spaced) on last whorl, and interstitial ribs on last whorl of some specimens.
<taxonomicName id="86C84D5AFFB4EB45FD09FACFFEEEFA63" authorityName="Squires" authorityYear="1984" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFB4EB45FD09FACFFEEEFA63" italics="true" pageId="8" pageNumber="7">Ancistrolepis carolineae</emphasis>
</taxonomicName>
differs from this Kamchakta species by having a smaller size shell, fewer spiral ribs on the spire whorls, many fewer and much less closely spaced spiral ribs on the siphonal canal, and the presence of a terminal varix on the outer lip.
</paragraph>
<paragraph id="417736D9FFB4EB45FF4FF9A2FB2AFECA" blockId="8.[112,801,195,1911]" lastBlockId="8.[832,1520,195,925]" pageId="8" pageNumber="7">
<bibRefCitation id="25594B28FFB4EB45FF4FF9A2FE84F9D3" author="Titova, L. V." box="[144,318,1568,1594]" pageId="8" pageNumber="7" pagination="1 - 16" refId="ref9577" refString="Titova, L. V. 1993. The early history of the North Pacific Ancistrolepidinae (Gastropoda: Buccinidae). Ruthenica 3: 1 - 16." type="book chapter" year="1993">Titova (1993)</bibRefCitation>
reported that
<taxonomicName id="86C84D5AFFB4EB45FE29F9A2FEACF9B4" authority="Titova, 1993" authorityName="Titova" authorityYear="1993" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="rategiensis">
<emphasis id="73BCEACBFFB4EB45FE29F9A2FC9AF9D0" box="[502,800,1568,1594]" italics="true" pageId="8" pageNumber="7">Ancistrolepis rategiensis</emphasis>
<bibRefCitation id="25594B28FFB4EB45FFAFF9C1FEACF9B4" author="Titova, L. V." box="[112,278,1603,1629]" pageId="8" pageNumber="7" pagination="1 - 16" refId="ref9577" refString="Titova, L. V. 1993. The early history of the North Pacific Ancistrolepidinae (Gastropoda: Buccinidae). Ruthenica 3: 1 - 16." type="book chapter" year="1993">Titova, 1993</bibRefCitation>
</taxonomicName>
shows much similarity to
<taxonomicName id="86C84D5AFFB4EB45FDA3F9C1FEA7F969" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="matchgarense">
<emphasis id="73BCEACBFFB4EB45FDA3F9C1FEA7F969" italics="true" pageId="8" pageNumber="7">Ancistrolepis matchgarense</emphasis>
</taxonomicName>
(
<bibRefCitation id="25594B28FFB4EB45FEEDF9E4FDB9F969" author="Makiyama, J." box="[306,515,1638,1664]" pageId="8" pageNumber="7" pagination="121 - 167" refId="ref8134" refString="Makiyama, J. 1934. The Asagaian molluscs of Yotukura and Matchgar. Memoirs of the College of Science, Kyoto Imperial University, Series B, 10: 121 - 167." type="journal article" year="1934">Makiyama, 1934</bibRefCitation>
: p. 165, pl. 7, figs. 56, 57) from the upper Oligocene Asagai Sandstone near Matchgar in northern Sakhalin. The Llajas Formation species is similar, therefore, to
<taxonomicName id="86C84D5AFFB4EB45FE33F952FD0CF903" box="[492,694,1744,1770]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="matchgarense">
<emphasis id="73BCEACBFFB4EB45FE33F952FD0CF903" box="[492,694,1744,1770]" italics="true" pageId="8" pageNumber="7">A. matchgarense</emphasis>
</taxonomicName>
, but this latter species, which has a small shell (like
<taxonomicName id="86C84D5AFFB4EB45FD7FF971FF20F8D9" authorityName="Squires" authorityYear="1984" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFB4EB45FD7FF971FF20F8D9" italics="true" pageId="8" pageNumber="7">A. carolineae</emphasis>
</taxonomicName>
) has rather poor preservation (interstitial threads not visible) and its siphonal canal is missing. The Llajas Formation species differs by having a “terminal varix” on the outer lip.
<taxonomicName id="86C84D5AFFB4EB45FBD5FF41FAE3FF34" box="[1034,1369,195,221]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="matchgarense">
<emphasis id="73BCEACBFFB4EB45FBD5FF41FAE3FF34" box="[1034,1369,195,221]" italics="true" pageId="8" pageNumber="7">Ancistrolepis matchgarense</emphasis>
</taxonomicName>
occurs with calcareous concretions containing
<taxonomicName id="86C84D5AFFB4EB45FB2EFF64FADDFEE9" authorityName="Lamarck" authorityYear="1799" box="[1265,1383,230,256]" class="Gastropoda" family="Turritellidae" genus="Turritella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="73BCEACBFFB4EB45FB2EFF64FADDFEE9" box="[1265,1383,230,256]" italics="true" pageId="8" pageNumber="7">Turritella</emphasis>
</taxonomicName>
, which is a shallow-marine gastropod.
</paragraph>
<paragraph id="417736D9FFB4EB45FCBFFEAEFB24FDD4" blockId="8.[832,1520,195,925]" pageId="8" pageNumber="7">
<taxonomicName id="86C84D5AFFB4EB45FCBFFEAEFB3FFEAF" authorityName="Squires" authorityYear="1984" box="[864,1157,300,326]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="carolineae">
<emphasis id="73BCEACBFFB4EB45FCBFFEAEFB3FFEAF" box="[864,1157,300,326]" italics="true" pageId="8" pageNumber="7">Ancistrolepis carolineae</emphasis>
</taxonomicName>
,
<taxonomicName id="86C84D5AFFB4EB45FB4CFEAFFA8CFEAF" authorityName="Titova" authorityYear="1993" box="[1171,1334,301,327]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="rategiensis">
<emphasis id="73BCEACBFFB4EB45FB4CFEAFFA8CFEAF" box="[1171,1334,301,327]" italics="true" pageId="8" pageNumber="7">A. rategiensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="86C84D5AFFB4EB45FAA4FEAFFC1AFE80" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="matchgarense">
<emphasis id="73BCEACBFFB4EB45FAA4FEAFFC1AFE80" italics="true" pageId="8" pageNumber="7">A. matchgarense</emphasis>
</taxonomicName>
differ from most other species of this genus by having subtriangular spiral ribs, rounded whorls, and a rounded shell base. Four other species, all from the Kuschiro coal fields in
<collectingCountry id="39DF7649FFB4EB45FB8AFE38FB27FE3D" box="[1109,1181,442,468]" name="Japan" pageId="8" pageNumber="7">Japan</collectingCountry>
, that share these particular properties are
<taxonomicName id="86C84D5AFFB4EB45FC28FE5FFAC3FE1E" authority="(Matsui, 1958)" baseAuthorityName="Matsui" baseAuthorityYear="1958" box="[1015,1401,476,503]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="chizuzenensis">
<emphasis id="73BCEACBFFB4EB45FC28FE5FFB00FE1F" box="[1015,1210,476,503]" italics="true" pageId="8" pageNumber="7">A. chizuzenensis</emphasis>
(
<bibRefCitation id="25594B28FFB4EB45FB14FE5EFAD7FE1E" author="Matsui, M." box="[1227,1389,476,503]" pageId="8" pageNumber="7" pagination="201 - 210" refId="ref8258" refString="Matsui, M. 1958. Species of the genus Neptunea from the Paleogene formations in the Kushiro coal field, Hokkaido, Japan. Transactions and Proceeding of the Paleontological Society of Japan. New Series, 30: 201 - 210." type="journal article" year="1958">Matsui, 1958</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="86C84D5AFFB4EB45FA5AFE5FFBE4FDF3" authority="(Matsui, 1959)" baseAuthorityName="Matsui" baseAuthorityYear="1959" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="onbetsuensis">
<emphasis id="73BCEACBFFB4EB45FA5AFE5FFC2DFDF0" italics="true" pageId="8" pageNumber="7">A. onbetsuensis</emphasis>
(
<bibRefCitation id="25594B28FFB4EB45FC74FD82FBE8FDF3" author="Matsui, M." box="[939,1106,511,538]" pageId="8" pageNumber="7" pagination="289 - 304" refId="ref8302" refString="Matsui, M. 1959. On some Oligocene molluscan fossils from Hokkaido, Japan. Contributions from the Department of Geology and Mineralogy Faculty of Science Hokkaido Unversity 761: 289 - 304." type="journal article" year="1959">Matsui, 1959</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="86C84D5AFFB4EB45FBB1FD82FA53FDF3" authority="(Matsui, 1958)" baseAuthorityName="Matsui" baseAuthorityYear="1958" box="[1134,1513,511,538]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="subcarinata">
<emphasis id="73BCEACBFFB4EB45FBB1FD82FA98FDF3" box="[1134,1314,512,538]" italics="true" pageId="8" pageNumber="7">A. subcarinata</emphasis>
(
<bibRefCitation id="25594B28FFB4EB45FAE8FD82FA67FDF3" author="Matsui, M." box="[1335,1501,511,538]" pageId="8" pageNumber="7" pagination="201 - 210" refId="ref8258" refString="Matsui, M. 1958. Species of the genus Neptunea from the Paleogene formations in the Kushiro coal field, Hokkaido, Japan. Transactions and Proceeding of the Paleontological Society of Japan. New Series, 30: 201 - 210." type="journal article" year="1958">Matsui, 1958</bibRefCitation>
)
</taxonomicName>
; and
<taxonomicName id="86C84D5AFFB4EB45FCAAFDA1FB22FDD7" authority="Takeda, 1953" authorityName="Takeda" authorityYear="1953" box="[885,1176,547,574]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="ezoana">
<emphasis id="73BCEACBFFB4EB45FCAAFDA1FC3CFDD4" box="[885,902,547,573]" italics="true" pageId="8" pageNumber="7">A</emphasis>
.
<emphasis id="73BCEACBFFB4EB45FC4CFDA6FC51FDD4" box="[915,1003,548,573]" italics="true" pageId="8" pageNumber="7">ezoana</emphasis>
<bibRefCitation id="25594B28FFB4EB45FC2EFDA1FB22FDD7" author="Takeda, H." box="[1009,1176,547,574]" pageId="8" pageNumber="7" pagination="1 - 103" refId="ref9497" refString="Takeda, H. 1953. The Poronai Formation (Oligocene Tertiary) of Hokkaido and South Sakhalin and its fossil fauna. Studies on Coal Geology, the Hokkaido Association of Coal Mining Technologists 3: 1 - 103." type="journal article" year="1953">Takeda, 1953</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="417736D9FFB4EB45FCBFFDC4FBF4FC74" blockId="8.[832,1520,195,925]" pageId="8" pageNumber="7">
Previous workers (
<bibRefCitation id="25594B28FFB4EB45FB95FDC4FB64FD89" author="Titova, L. V." box="[1098,1246,582,608]" pageId="8" pageNumber="7" pagination="1 - 16" refId="ref9577" refString="Titova, L. V. 1993. The early history of the North Pacific Ancistrolepidinae (Gastropoda: Buccinidae). Ruthenica 3: 1 - 16." type="book chapter" year="1993">Titova 1993</bibRefCitation>
,
<bibRefCitation id="25594B28FFB4EB45FB36FDC5FC3BFD6A" author="Egorov, R. V. &amp; S. L. Barsukov" pageId="8" pageNumber="7" refId="ref7215" refString="Egorov, R. V. and S. L. Barsukov. 1994. Recent Ancistrolepidinae (Buccinidae). Colus-Doverie Ltd., Moscow. 47 pp. (Title and introduction in Russian)." type="book" year="1994">Egorov and Barsukov 1994</bibRefCitation>
) have placed various species of
<taxonomicName id="86C84D5AFFB4EB45FAC6FDEBFA0CFD6A" authorityName="Dall" authorityYear="1895" box="[1305,1462,617,643]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="73BCEACBFFB4EB45FAC6FDEBFA0CFD6A" box="[1305,1462,617,643]" italics="true" pageId="8" pageNumber="7">Ancistrolepis</emphasis>
</taxonomicName>
into two informal groups, referred commonly to as the
<taxonomicName id="86C84D5AFFB4EB45FA7CFD0FFCC0FD20" baseAuthorityName="Dall" baseAuthorityYear="1891" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="eucosmius">
<emphasis id="73BCEACBFFB4EB45FA7CFD0FFCC0FD20" italics="true" pageId="8" pageNumber="7">eucosmius</emphasis>
</taxonomicName>
“stock” and the
<emphasis id="73BCEACBFFB4EB45FB9AFD32FB69FD20" box="[1093,1235,688,713]" italics="true" pageId="8" pageNumber="7">grammatus</emphasis>
“stock.” The
<taxonomicName id="86C84D5AFFB4EB45FAADFD32FA4AFD20" baseAuthorityName="Dall" baseAuthorityYear="1891" box="[1394,1520,688,713]" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="species" species="eucosmius">
<emphasis id="73BCEACBFFB4EB45FAADFD32FA4AFD20" box="[1394,1520,688,713]" italics="true" pageId="8" pageNumber="7">eucosmius</emphasis>
</taxonomicName>
“stock,” bearing the name of the
<typeStatus id="9E73887BFFB4EB45FB3FFD51FAADFD04" box="[1248,1303,723,749]" pageId="8" pageNumber="7">type</typeStatus>
species of
<taxonomicName id="86C84D5AFFB4EB45FA79FD51FC27FCF9" authorityName="Dall" authorityYear="1895" class="Gastropoda" family="Buccinidae" genus="Ancistrolepis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="73BCEACBFFB4EB45FA79FD51FC27FCF9" italics="true" pageId="8" pageNumber="7">Ancistrolepis</emphasis>
</taxonomicName>
, is supposedly characterized by relatively small shells with relatively few spiral ribs, which are narrow and have a square shape. The
<emphasis id="73BCEACBFFB4EB45FB18FCBFFAEFFCBF" box="[1223,1365,829,854]" italics="true" pageId="8" pageNumber="7">grammatus</emphasis>
“stock,” has larger shells with much stronger spiral ribs, commonly referred to as “T” ribs
</paragraph>
</subSubSection>
</treatment>
</document>