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<document id="FCED3D96FBD09E662EEBED1EC1CB27B1" ID-DOI="10.1649/523" ID-ISSN="1938-4394" ID-Zenodo-Dep="5345219" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630199770588" checkinUser="tatiana" docAuthor="Arias, Elizabeth T." docDate="2004" docId="C94F87CD012AFFB31BF1FAB6FC78FA56" docLanguage="en" docName="523.pdf" docOrigin="The Coleopterists Bulletin 58 (3)" docSource="http://dx.doi.org/10.1649/523" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Alyma calafquenensis Arias 2004, new species" docType="treatment" docVersion="3" lastPageNumber="427" masterDocId="3576FFB5012DFFBB1A74FF8DFFFDFF85" masterDocTitle="A New Click Beetle Genus from Chilean Temperate Forests, ALYMA (Coleoptera: Elateridae: Pomachiliini)" masterLastPageNumber="427" masterPageNumber="413" pageNumber="413" updateTime="1699183367765" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="58528F694496EF2DD2E45F791FFE12ED">A New Click Beetle Genus from Chilean Temperate Forests, ALYMA (Coleoptera: Elateridae: Pomachiliini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="C2282BB45343720623AE97B350A1FBC9">Arias, Elizabeth T.</mods:namePart>
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<mods:title id="7C58E51E6F2B3E6A5844FD48988E1B61">The Coleopterists Bulletin</mods:title>
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<mods:date id="FC89F28AC98149ABC8103B0D59C372AD">2004</mods:date>
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<mods:number id="8C331E3F88F0CD250ABC1D6625ECEBEF">2004-09-30</mods:number>
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<mods:classification id="82147FA2F3DE5342B64C93C3424A35F8">journal article</mods:classification>
<mods:identifier id="235EBDECDCE061AE2EC2E2F50E19288B" type="DOI">10.1649/523</mods:identifier>
<mods:identifier id="01B9F5073AD72902F6CA474CA7240CB4" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="681CCC13F3A4BF41487E531CE3362E56" type="Zenodo-Dep">5345219</mods:identifier>
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<treatment id="C94F87CD012AFFB31BF1FAB6FC78FA56" LSID="urn:lsid:plazi:treatment:C94F87CD012AFFB31BF1FAB6FC78FA56" httpUri="http://treatment.plazi.org/id/C94F87CD012AFFB31BF1FAB6FC78FA56" lastPageId="8" pageId="7">
<subSubSection id="09FC6550012AFFBC1BF1FAB6FCEBFAD5" box="[389,790,1339,1360]" pageId="7" type="nomenclature">
<paragraph id="415936DB012AFFBC1BF1FAB6FCEBFAD5" blockId="7.[389,790,1339,1387]" box="[389,790,1339,1360]" pageId="7">
<heading id="1A1181B7012AFFBC1BF1FAB6FCEBFAD5" box="[389,790,1339,1360]" centered="true" fontSize="9" level="2" pageId="7" reason="2">
<taxonomicName id="86E64D58012AFFBC1BF1FAB6FD6EFAD5" authority="Arias" authorityName="Arias" authorityYear="2004" box="[389,659,1339,1360]" class="Insecta" family="Elateridae" genus="Alyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="calafquenensis" status="sp. nov.">
<emphasis id="7392EAC9012AFFBC1BF1FAB6FDABFAD5" box="[389,598,1339,1360]" italics="true" pageId="7">Alyma calafquenensis</emphasis>
Arias
</taxonomicName>
,
<emphasis id="7392EAC9012AFFBC18EBFAB6FCEBFAD5" bold="true" box="[671,790,1339,1360]" pageId="7">
<taxonomicNameLabel id="68A157B2012AFFBC18EBFAB6FCEBFAD5" box="[671,790,1339,1360]" pageId="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="09FC6550012AFFB31BADFADBFC80FB7B" lastPageId="8" pageId="7" type="description">
<paragraph id="415936DB012AFFBC1BADFADBFD3EFAEE" blockId="7.[389,790,1339,1387]" box="[473,707,1366,1387]" pageId="7">
(
<figureCitation id="D9DD2A5E012AFFBC1B94FADBFDD9FAEE" box="[480,548,1366,1387]" captionStart="Figs" captionStartId="10.[169,208,950,969]" captionTargetBox="[211,915,196,926]" captionTargetId="figure-206@10.[208,916,194,929]" captionTargetPageId="10" captionText="Figs. 613. Antenna of Alyma species. 6) A. pallipes; 7) A. ariasvillegasai; 8) A." figureDoi="http://doi.org/10.5281/zenodo.5345233" httpUri="https://zenodo.org/record/5345233/files/figure.png" pageId="7">Figs. 8</figureCitation>
,
<figureCitation id="D9DD2A5E012AFFBC1845FADBFDB6FAEE" box="[561,587,1366,1387]" captionStart="Figs" captionStartId="11.[197,236,916,935]" captionTargetBox="[293,886,195,893]" captionTargetId="figure-202@11.[293,886,194,894]" captionTargetPageId="11" captionText="Figs. 1421. Pronotal hypomera of Alyma species. 14) A. pallipes; 15) A. ariasvillegasai; 16) A. calafquenensis; 17) A. contulmoensis; 18) A. lawlerae; 19) A. quiriquinaensis; 20) A." figureDoi="http://doi.org/10.5281/zenodo.5345235" httpUri="https://zenodo.org/record/5345235/files/figure.png" pageId="7">16</figureCitation>
,
<figureCitation id="D9DD2A5E012AFFBC1823FADBFD8CFAEE" box="[599,625,1366,1387]" captionStart="Figs" captionStartId="12.[169,208,833,852]" captionTargetBox="[148,979,195,810]" captionTargetId="figure-203@12.[148,980,194,811]" captionTargetPageId="12" captionText="Figs. 2228. Tarsi of Alyma species. 22) A. pallipes; 23) A. ariasvillegasai; 24) A." figureDoi="http://doi.org/10.5281/zenodo.5345237" httpUri="https://zenodo.org/record/5345237/files/figure.png" pageId="7">24</figureCitation>
,
<figureCitation id="D9DD2A5E012AFFBC1809FADBFD6AFAEE" box="[637,663,1366,1387]" captionStart="Figs" captionStartId="13.[197,236,602,621]" captionTargetBox="[176,1007,195,579]" captionTargetId="figure-336@13.[175,1007,194,580]" captionTargetPageId="13" captionText="Figs. 2933. Male genitalia of Alyma species. 29) Alyma calafquenensis; 30) A." figureDoi="http://doi.org/10.5281/zenodo.5345241" httpUri="https://zenodo.org/record/5345241/files/figure.png" pageId="7">29</figureCitation>
,
<figureCitation id="D9DD2A5E012AFFBC18D7FADBFD41FAEE" box="[675,700,1366,1387]" captionStart="Fig" captionStartId="14.[169,200,776,795]" captionTargetBox="[151,980,195,754]" captionTargetId="figure-235@14.[148,980,194,755]" captionTargetPageId="14" captionText="Fig. 34. Distribution map of Alyma species. Alyma pallipes (Solier) (N), A. ariasvillegasai" figureDoi="http://doi.org/10.5281/zenodo.5345243" httpUri="https://zenodo.org/record/5345243/files/figure.png" pageId="7">34</figureCitation>
)
</paragraph>
<paragraph id="415936DB012AFFBC1AB1FAF2FD8BFA2F" blockId="7.[175,1005,1407,1570]" pageId="7">
<emphasis id="7392EAC9012AFFBC1AB1FAF2FECEFA17" bold="true" box="[197,307,1407,1426]" pageId="7">Description.</emphasis>
Body: elongate; brown; integument dull; vestiture thick pale yellowish, semidecumbent; length
<quantity id="861E9B3E012AFFBC1B20FA1AFE65FA2F" box="[340,408,1431,1450]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="7" unit="mm" value="6.1">6.1 mm</quantity>
, width
<quantity id="861E9B3E012AFFBC1BADFA1AFDE3FA2F" box="[473,542,1431,1450]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="7" unit="mm" value="2.1">2.1 mm</quantity>
; PEI: 2.6.
</paragraph>
<paragraph id="415936DB012AFFBC1AB3FA22FC4CFA77" blockId="7.[175,1005,1407,1570]" pageId="7">
<emphasis id="7392EAC9012AFFBC1AB3FA22FF08FA47" box="[199,245,1455,1474]" italics="true" pageId="7">Head</emphasis>
: convex; frontoclypeal carina region sloping downward; clypeus and labrum with long setae, decumbent; labrum slightly convex, 2.0
<emphasis id="7392EAC9012AFFBC1842FA4BFDB5FA5C" bold="true" box="[566,584,1478,1497]" pageId="7">3</emphasis>
as long as wide; antennomera 2 through 8 serrate, antennomere 9 exceeding pronotal posterior angles, [AP: 7-6-9-9-12-9-9-9-9-12] (
<figureCitation id="D9DD2A5E012AFFBC1904FA52FC58FA77" box="[880,933,1503,1522]" captionStart="Figs" captionStartId="10.[169,208,950,969]" captionTargetBox="[211,915,196,926]" captionTargetId="figure-206@10.[208,916,194,929]" captionTargetPageId="10" captionText="Figs. 613. Antenna of Alyma species. 6) A. pallipes; 7) A. ariasvillegasai; 8) A." figureDoi="http://doi.org/10.5281/zenodo.5345233" httpUri="https://zenodo.org/record/5345233/files/figure.png" pageId="7">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="415936DB012AFFBC1AB3FA7AFC11F9A7" blockId="7.[175,1005,1407,1570]" pageId="7">
<emphasis id="7392EAC9012AFFBC1AB3FA7AFEE0F98F" box="[199,285,1527,1546]" italics="true" pageId="7">Prothorax</emphasis>
: strongly convex medially; areolate, sides parallel; base curved; pronotal posterior angles straight; prosternal lobe strongly bent; prosternum strongly convex; pronotosternal lobe
</paragraph>
<paragraph id="415936DB012AFFBC1A6FF901FD05F91B" blockId="7.[27,784,1676,1713]" box="[27,760,1676,1694]" pageId="7">
Downloaded From: https://bioone.org/journals/The-Coleopterists-Bulletin on
<date id="3558101B012AFFBC18FBF901FD05F91B" box="[655,760,1676,1694]" pageId="7" value="2021-08-26">26 Aug 2021</date>
</paragraph>
<paragraph id="415936DB012AFFBC1A6FF92DFCEDF935" blockId="7.[27,784,1676,1713]" box="[27,784,1695,1713]" pageId="7">Terms of Use: https://bioone.org/terms-of-use Access provided by The Coleopterists Society</paragraph>
<paragraph id="415936DB0125FFB31AE0FF0DFEC9FBD2" blockId="8.[148,180,128,147]" lastBlockId="8.[148,979,1020,1278]" pageId="8">
420 marginate broadly curved; pronotosternal suture sinuate; hypomeron rugulose, uni-excavate, apically not marginate, antennal groove present (
<figureCitation id="D9DD2A5E0125FFB31840FB99FD8EFBA2" box="[564,627,1044,1063]" captionStart="Figs" captionStartId="11.[197,236,916,935]" captionTargetBox="[293,886,195,893]" captionTargetId="figure-202@11.[293,886,194,894]" captionTargetPageId="11" captionText="Figs. 1421. Pronotal hypomera of Alyma species. 14) A. pallipes; 15) A. ariasvillegasai; 16) A. calafquenensis; 17) A. contulmoensis; 18) A. lawlerae; 19) A. quiriquinaensis; 20) A." figureDoi="http://doi.org/10.5281/zenodo.5345235" httpUri="https://zenodo.org/record/5345235/files/figure.png" pageId="8">Fig. 16</figureCitation>
); prosternum at procoxae marginate, and slightly elevated; procoxal distance 2.0
<emphasis id="7392EAC90125FFB31B9AFBA6FDFDFBBB" bold="true" box="[494,512,1067,1086]" pageId="8">3</emphasis>
procoxal diameter, pronotosternal spine length 0.7
<emphasis id="7392EAC90125FFB319B5FBA6FC2EFBBB" bold="true" box="[961,979,1067,1086]" pageId="8">3</emphasis>
procoxal diameter.
</paragraph>
<caption id="159966530125FFB31ADDFC29FF26FC55" ID-DOI="http://doi.org/10.5281/zenodo.5345229" ID-Zenodo-Dep="5345229" httpUri="https://zenodo.org/record/5345229/files/figure.png" pageId="8" startId="8.[169,200,932,951]" targetBox="[336,788,195,901]" targetPageId="8">
<paragraph id="415936DB0125FFB31ADDFC29FF26FC55" blockId="8.[148,978,932,976]" pageId="8">
<emphasis id="7392EAC90125FFB31ADDFC29FF18FC32" bold="true" box="[169,229,932,951]" pageId="8">Fig. 4.</emphasis>
Female genitalia of
<taxonomicName id="86E64D580125FFB31BD3FC28FC99FC3D" authority="(Illustration Nancy V. Arias)" box="[423,868,933,952]" class="Insecta" family="Elateridae" genus="Alyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="ariasvillegasai">
<emphasis id="7392EAC90125FFB31BD3FC28FD9FFC3D" box="[423,610,933,952]" italics="true" pageId="8">Alyma ariasvillegasai</emphasis>
(Illustration Nancy V. Arias)
</taxonomicName>
. Scale bar
<emphasis id="7392EAC90125FFB319B6FC28FC2FFC32" bold="true" box="[962,978,933,951]" italics="true" pageId="8">¼</emphasis>
1.0 mm.
</paragraph>
</caption>
<paragraph id="415936DB0125FFB31AD8FBD1FF19FB1A" blockId="8.[148,979,1020,1278]" pageId="8">
<taxonomicName id="86E64D580125FFB31AD8FBD1FF02FBEA" box="[172,255,1116,1135]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="7392EAC90125FFB31AD8FBD1FF02FBEA" box="[172,255,1116,1135]" italics="true" pageId="8">Scutellum</emphasis>
</taxonomicName>
: U-shaped, vestiture semi-decumbent, long, yellow; mesocoxae separated by 0.3
<emphasis id="7392EAC90125FFB319B5FBD6FC2EFBEB" bold="true" box="[961,979,1115,1134]" pageId="8">3</emphasis>
mesocoxal diameter; posterior margin of mesosternal cavity extending posteriorly 0.4
<emphasis id="7392EAC90125FFB31916FBFEFC89FB03" bold="true" box="[866,884,1139,1158]" pageId="8">3</emphasis>
mesocoxal diameter.
</paragraph>
<paragraph id="415936DB0125FFB31AD8FB29FE83FB63" blockId="8.[148,979,1020,1278]" pageId="8">
<emphasis id="7392EAC90125FFB31AD8FB29FF1DFB32" box="[172,224,1188,1207]" italics="true" pageId="8">Elytra</emphasis>
: interstria rugulose, slightly convex; striae pitted, pits irregular; interstria flat; elytral apex rounded and crenulate.
<emphasis id="7392EAC90125FFB31B2AFB31FE83FB4A" box="[350,382,1212,1231]" italics="true" pageId="8">Leg</emphasis>
: yellowish brown; tarsomere 4 similar in length to tarsomere 2, [TP: 25-19-18-23-15], (
<figureCitation id="D9DD2A5E0125FFB31B46FB5EFE8EFB63" box="[306,371,1235,1254]" captionStart="Figs" captionStartId="12.[169,208,833,852]" captionTargetBox="[148,979,195,810]" captionTargetId="figure-203@12.[148,980,194,811]" captionTargetPageId="12" captionText="Figs. 2228. Tarsi of Alyma species. 22) A. pallipes; 23) A. ariasvillegasai; 24) A." figureDoi="http://doi.org/10.5281/zenodo.5345237" httpUri="https://zenodo.org/record/5345237/files/figure.png" pageId="8">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="415936DB0125FFB31AD8FB66FC80FB7B" blockId="8.[148,979,1020,1278]" box="[172,893,1259,1278]" pageId="8">
<emphasis id="7392EAC90125FFB31AD8FB66FEDAFB7B" box="[172,295,1259,1278]" italics="true" pageId="8">Male genitalia</emphasis>
: aedeagus 2.5
<emphasis id="7392EAC90125FFB31BD6FB66FE49FB7B" bold="true" box="[418,436,1259,1278]" pageId="8">3</emphasis>
as long as wide; median lobe spoon shaped; (
<figureCitation id="D9DD2A5E0125FFB31947FB66FC8FFB7B" box="[819,882,1259,1278]" captionStart="Figs" captionStartId="13.[197,236,602,621]" captionTargetBox="[176,1007,195,579]" captionTargetId="figure-336@13.[175,1007,194,580]" captionTargetPageId="13" captionText="Figs. 2933. Male genitalia of Alyma species. 29) Alyma calafquenensis; 30) A." figureDoi="http://doi.org/10.5281/zenodo.5345241" httpUri="https://zenodo.org/record/5345241/files/figure.png" pageId="8">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="09FC65500125FFB31AD8FA93FEC1FACB" pageId="8" type="materials_examined">
<paragraph id="415936DB0125FFB31AD8FA93FEC1FACB" blockId="8.[148,977,1310,1491]" pageId="8">
<emphasis id="7392EAC90125FFB31AD8FA93FEBBFAB6" bold="true" box="[172,326,1310,1331]" pageId="8">Type Material.</emphasis>
<materialsCitation id="F18E3C860125FFB31B26FA92FEC1FACB" collectingDate="1986-01-25" collectionCode="ETA" collectorName="L. E. Pena" country="Chile" location="Calafquen" pageId="8" specimenCount="1" typeStatus="holotype">
<typeStatus id="9E5D88790125FFB31B26FA92FE53FAB1" box="[338,430,1311,1332]" pageId="8" type="holotype">Holotype</typeStatus>
: male
<collectingCountry id="39F1764B0125FFB31B9BFA92FDCAFAB1" box="[495,567,1311,1332]" name="Chile" pageId="8">CHILE</collectingCountry>
,
<location id="443960000125FFB31836FA92FD5BFAB1" LSID="urn:lsid:plazi:treatment:C94F87CD012AFFB31BF1FAB6FC78FA56:443960000125FFB31836FA92FD5BFAB1" box="[578,678,1311,1332]" country="Chile" name="Calafquen" pageId="8">Calafquén</location>
,
<date id="3558101B0125FFB318C5FA92FCF2FAB1" box="[689,783,1311,1332]" pageId="8" value="1986-01-25">
<collectingDate id="251CE9F30125FFB318C5FA92FCF2FAB1" box="[689,783,1311,1332]" pageId="8" value="1986-01-25">25.I.1986</collectingDate>
</date>
,
<collectorName id="EC13530D0125FFB3196EFA92FC86FAB1" box="[794,891,1311,1332]" pageId="8">L.E. Peña</collectorName>
,
<quantity id="861E9B3E0125FFB319F2FA92FC2CFAB1" box="[902,977,1311,1332]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="8" unit="mm" value="6.1">6.1 mm</quantity>
in length, [
<collectionCode id="27F7AE1E0125FFB31A89FAB4FECFFACB" box="[253,306,1337,1358]" pageId="8">ETA</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="09FC65500125FFB31AD8FAD9FD47FAEC" box="[172,698,1364,1385]" pageId="8" type="distribution">
<paragraph id="415936DB0125FFB31AD8FAD9FD47FAEC" blockId="8.[148,977,1310,1491]" box="[172,698,1364,1385]" pageId="8">
<emphasis id="7392EAC90125FFB31AD8FAD9FED3FAEC" bold="true" box="[172,302,1364,1385]" pageId="8">Distribution.</emphasis>
Calafquén, IX Region,
<collectingCountry id="39F1764B0125FFB31869FAD9FDAEFAEC" box="[541,595,1364,1385]" name="Chile" pageId="8">Chile</collectingCountry>
, (
<figureCitation id="D9DD2A5E0125FFB31811FAD9FD53FAEC" box="[613,686,1364,1385]" captionStart="Fig" captionStartId="14.[169,200,776,795]" captionTargetBox="[151,980,195,754]" captionTargetId="figure-235@14.[148,980,194,755]" captionTargetPageId="14" captionText="Fig. 34. Distribution map of Alyma species. Alyma pallipes (Solier) (N), A. ariasvillegasai" figureDoi="http://doi.org/10.5281/zenodo.5345243" httpUri="https://zenodo.org/record/5345243/files/figure.png" pageId="8">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="09FC65500125FFB31AD8FAE3FED3FA3C" pageId="8" type="discussion">
<paragraph id="415936DB0125FFB31AD8FAE3FED3FA3C" blockId="8.[148,977,1310,1491]" pageId="8">
<emphasis id="7392EAC90125FFB31AD8FAE3FEF3FA06" bold="true" box="[172,270,1390,1411]" pageId="8">Remarks.</emphasis>
<taxonomicName id="86E64D580125FFB31B6EFAE2FE11FA01" authorityName="Arias" authorityYear="2004" box="[282,492,1391,1412]" class="Insecta" family="Elateridae" genus="Alyma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="calafquenensis">
<emphasis id="7392EAC90125FFB31B6EFAE2FE11FA01" box="[282,492,1391,1412]" italics="true" pageId="8">Alyma calafquenensis</emphasis>
</taxonomicName>
can be recognized by its dull integument, brown color, yellowish pale and thick vestiture semi-decumbent, and straight posterior pronotal angles.
</paragraph>
</subSubSection>
<subSubSection id="09FC65500125FFB31AD8FA33FC78FA56" box="[172,901,1470,1491]" pageId="8" type="etymology">
<paragraph id="415936DB0125FFB31AD8FA33FC78FA56" blockId="8.[148,977,1310,1491]" box="[172,901,1470,1491]" pageId="8">
<emphasis id="7392EAC90125FFB31AD8FA33FEDDFA56" bold="true" box="[172,288,1470,1491]" pageId="8">Etymology.</emphasis>
This species is named after the placed where it was collected.
</paragraph>
</subSubSection>
</treatment>
</document>