786 lines
89 KiB
XML
786 lines
89 KiB
XML
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<mods:mods id="36795BC0CE0BF70E1096B7718F07E288" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4BDC98FDCDA99FD15D6BE42582F62F32">
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<mods:title id="3DE24FB8FE0EFFA0A50E99A3114DCC42">Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae)</mods:title>
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<mods:namePart id="C373F93BCDEE1305A9F0F18639407039">Goulding, Tricia C.</mods:namePart>
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<mods:namePart id="8FC9F2055EEDA4FE0FCD99757BF3E9A7">Khalil, Munawar</mods:namePart>
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<mods:namePart id="3AFD0C2099C95EA0CB557EDB9D3B0051">Tan, Shau Hwai</mods:namePart>
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<mods:namePart id="12C1FA1647B36789D4CF5F1A0DEFFA29">Dayrat, Benoit</mods:namePart>
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<mods:typeOfResource id="A6F17592E4BD88062E0042E93130903E">text</mods:typeOfResource>
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<mods:title id="94E777AFB8E4B174813B7A6E221CCE01">ZooKeys</mods:title>
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<mods:date id="DB200FAE2D551D6F593339F457E80300">2018</mods:date>
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<mods:detail id="1C9F48F876BCE77C4647215286AB2176" type="volume">
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<mods:number id="F6ECFBB6C04074CBB87F5FD0EC46973E">763</mods:number>
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<mods:url id="B445C4B0B3BD13A6B23A2E242830B4F5">http://dx.doi.org/10.3897/zookeys.763.21252</mods:url>
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<mods:classification id="DCAEE649BBB96E2A14735AEB3FF07BE7">journal article</mods:classification>
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<mods:identifier id="CC174623600FECAD7FFF394F64FD9CB1" type="DOI">http://dx.doi.org/10.3897/zookeys.763.21252</mods:identifier>
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<mods:identifier id="4F1A82D0A2D8ED3895A9C4974B33AB8A" type="Pensoft-Pub">1313-2970-763-1</mods:identifier>
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<mods:identifier id="B635364E3EF5E97076C5FA77B41620EE" type="ZooBank">90B772554C5E436CA793D924892B5B14</mods:identifier>
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</mods:mods>
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<treatment id="7B4B53B5A05304A7B261AC3213A196C5" ID-GBIF-Taxon="144234921" LSID="urn:lsid:plazi:treatment:7B4B53B5A05304A7B261AC3213A196C5" httpUri="http://treatment.plazi.org/id/7B4B53B5A05304A7B261AC3213A196C5" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="ADE39C0B87511CDEF3BF7330B1E4027F" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="55AB6D2602EE4FBF0C316F5B8A3AEE7C" pageId="0" pageNumber="1">
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<taxonomicName id="A3D19817A3B68A73581C521A8A237A6D" ID-CoL="7FV28" authority="Lesson, 1830" authorityName="Lesson" authorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater (Lesson, 1830)</taxonomicName>
|
||
<taxonomicNameLabel id="9175D4DE2AD505C8AE6B9E450D497381" pageId="0" pageNumber="1">comb. n.</taxonomicNameLabel>
|
||
Figs 34, 35, 36, 37, 38, 39, 40, 41, 42
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4EA000DD19C9A6B04D1EC033A251E1C1" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="D132027D19731C0068AA0070E6712745" pageId="0" pageNumber="1">
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||
<taxonomicName id="E2EFDB626E7B2E7B222905DB5A79682E" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Onchidium ater</taxonomicName>
|
||
Lesson, 1830: 300;
|
||
<bibRefCitation id="BA6EEE89E65EEE36D1AFBE584739EE51" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall 1919</bibRefCitation>
|
||
: 327;
|
||
<bibRefCitation id="4F853C38B38A19C34A0D30353D359ADE" author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher (Jena)" pageId="0" pageNumber="1" pagination="29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
|
||
: 84-85.
|
||
</paragraph>
|
||
<paragraph id="0EA150D7E605E3D4D7CFCAC71FE7A07E" pageId="0" pageNumber="1">
|
||
<taxonomicName id="7FBB5019DF601FD93F24EE63D2A71363" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium keiense" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="keiense">Onchidium keiense</taxonomicName>
|
||
Hoffmann, 1926: 18-24, figs 1-5;
|
||
<bibRefCitation id="50315EE7A2C23171172389E73E1E932B" author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher (Jena)" pageId="0" pageNumber="1" pagination="29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
|
||
: 81(as
|
||
<taxonomicName id="92BEB28BAC9F1F22E7CA7BA99D8CB991" class="Gastropoda" family="Onchidiidae" genus="Oncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncidium keiense" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="keiense">Oncidium keiense</taxonomicName>
|
||
). syn. n.
|
||
</paragraph>
|
||
<paragraph id="7459481876C0B2969382BF03CDED56B0" pageId="0" pageNumber="1">
|
||
<taxonomicName id="715D132733130127F105A2414F55CDF0" class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis atra" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="atra">Scaphis atra</taxonomicName>
|
||
:
|
||
<bibRefCitation id="5A0B8C70926B81AD1FCA30DE4127F140" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="9787AE920788CE96EA975BE3402D2F72" originalValue="Labbé">Labbe</normalizedToken>
|
||
1934
|
||
</bibRefCitation>
|
||
: fig. 30.
|
||
</paragraph>
|
||
<paragraph id="88A33A144F49D4212F70327BD1BA7113" pageId="0" pageNumber="1">
|
||
<taxonomicName id="AA3ACCDD701008312FE56EEC8B72AD04" class="Gastropoda" family="Onchidiidae" genus="Paraoncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraoncidium keiense" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="keiense">Paraoncidium keiense</taxonomicName>
|
||
:
|
||
<bibRefCitation id="972CF14C1200056294BAC94A3EADD1F1" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="5EAF2F86B81D0F939EC14E0A8E914D55" originalValue="Labbé">Labbe</normalizedToken>
|
||
1934
|
||
</bibRefCitation>
|
||
: 230.
|
||
</paragraph>
|
||
<paragraph id="E947820EE23AAF73F6FB72016B0FF1BF" pageId="0" pageNumber="1">
|
||
<taxonomicName id="066DB1B3AD7123B4460E6F2D1143ADC5" class="Gastropoda" family="Onchidiidae" genus="Lessonina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lessonina ferruginea" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferruginea">Lessonina ferruginea</taxonomicName>
|
||
:
|
||
<bibRefCitation id="A4CF2FC5A89B7963F7EF67ED2B8C4B54" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="E6E23EB838389F39C16EF7C8809B6793" originalValue="Labbé">Labbe</normalizedToken>
|
||
1934
|
||
</bibRefCitation>
|
||
: fig. 50 [non
|
||
<taxonomicName id="358E358A32C2B1EDB0919ED2D2A513F9" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">Onchidium ferrugineum</taxonomicName>
|
||
Lesson, 1830].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BF18E55F0E2E042DAD4CA88267672538" pageId="0" pageNumber="1" type="type locality">
|
||
<paragraph id="91E469FB87B3DF5DE291E06505D6E84D" pageId="0" pageNumber="1">Type locality</paragraph>
|
||
<paragraph id="28A684343110B9394C85105AD6C55655" pageId="0" pageNumber="1">
|
||
(
|
||
<taxonomicName id="44EA578A08D23A00B8485B2FD4F7CFBA" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
). Havre de
|
||
<normalizedToken id="4D1EB967E35BA7F61E5CC87EF194E49D" originalValue="Doréry">Dorery</normalizedToken>
|
||
(for Dorey),
|
||
<normalizedToken id="14466AA7E21ADC2C483C9CDDDD72549A" originalValue="à">a</normalizedToken>
|
||
la Nouvelle
|
||
<normalizedToken id="94E5DEAC1092E2180FF2140013ED7937" originalValue="Guinée">Guinee</normalizedToken>
|
||
[Manokwari, West Papua, Indonesia]. No fresh material was collected from the type locality. However, fresh material was collected from Halmahera, approximately 700 kilometers from the type locality in West Papua (see below, additional material).
|
||
</paragraph>
|
||
<paragraph id="973EC6774FC25131CA00584483FCE0F6" pageId="0" pageNumber="1">
|
||
Type locality (
|
||
<taxonomicName id="D25B77793DB0E298987260F2A8E2FFC5" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
). Toeal (Kei-Inseln) [Kei Islands, Moluccan Islands, Indonesia]. Fresh specimens were collected from the type locality (see below, additional material).
|
||
</paragraph>
|
||
<paragraph id="C528E3C4FB2D9AD9BCDFD58D6CC89C05" pageId="0" pageNumber="1">
|
||
Type material (
|
||
<taxonomicName id="2826D40F187D6980B4FB321E313E1301" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
). Two syntypes: 30/17 and 28/15 mm (MNHN 22950). Both syntypes were previously dissected and some internal organs are missing. The posterior (female) reproductive parts remain in the smaller syntype and only the oviduct remains in the larger syntype. The male (anterior) parts are missing in both syntypes. The digestive system is present in both syntypes, but the intestinal loops are not preserved.
|
||
</paragraph>
|
||
<paragraph id="F50C465BB6ECACB2366C1EAA24644EC5" pageId="0" pageNumber="1">
|
||
Type material (
|
||
<taxonomicName id="6B7B219CF46409F79359BEA5A29A49FE" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
). Lectotype, 32/21 mm, designated here (ZMUC). Thirty paralectotypes (35/24 to 11/8 mm) are well preserved (ZMUC), although several specimens were dissected prior to the present study and the male parts were previously removed (ZMUC). The lectotype was dissected for this study and all organs left in vials inside the jar. The species name on the label is
|
||
<taxonomicName id="98CC2788437A6C8648B8D9DFDCF5150B" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium mortenseni" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mortenseni">Onchidium mortenseni</taxonomicName>
|
||
. However, the rest of the information on the label (specimens collected by Mortensen from Toeal and studied by Hoffmann) is identical to the information in
|
||
<normalizedToken id="3829EFE6BE4DD1C3877292C00D711BF4" originalValue="Hoffmann’s">Hoffmann's</normalizedToken>
|
||
original description of
|
||
<taxonomicName id="F24EC123F2C0796B36CAF765B2C2F5A8" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
(the collector is not mentioned in the species description, but the title of the paper indicates the species are described based on collections from
|
||
<normalizedToken id="8EED06D868884DF7C14B0E4E22F25BFD" originalValue="Mortensen’s">Mortensen's</normalizedToken>
|
||
expedition). The only other onchidiid species described by Hoffmann from the Kei Islands was
|
||
<taxonomicName id="0B577A5C5A13976FFBA8DDC8D5A0705D" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium verruculatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculatum">Onchidium verruculatum</taxonomicName>
|
||
Cuvier, 1830, which Hoffmann recognized thanks to the presence of gills on the posterior notum (
|
||
<taxonomicName id="45A32083ED298D75A01AE6EF457740C8" lsidName="O. verruculatum" pageId="0" pageNumber="1" rank="species" species="verruculatum">O. verruculatum</taxonomicName>
|
||
actually belongs to the genus
|
||
<taxonomicName id="E00D7034E55FE2A1A9961B263AD01351" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
|
||
). Gills are not visible on any of the syntypes labeled
|
||
<taxonomicName id="F54D3B4C457E1B50DB248B0C03CA811F" lsidName="O. mortenseni" pageId="0" pageNumber="1" rank="species" species="mortenseni">O. mortenseni</taxonomicName>
|
||
, which strongly suggests that those specimens are the type material of
|
||
<taxonomicName id="C2169822CDCFBB22C25EF5F730793D01" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
and that Hoffmann changed his mind for the specific name (i.e., he replaced
|
||
<taxonomicName id="223DC51CBCC80ABBC5568B05CCBF8B9C" lsidName="mortenseni" pageId="0" pageNumber="1" rank="species" species="mortenseni">mortenseni</taxonomicName>
|
||
by
|
||
<taxonomicName id="22BB689FF2B682B7001029BCBEC8C8FA" lsidName="keiense" pageId="0" pageNumber="1" rank="species" species="keiense">keiense</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BA037634516424671253D76AE2D59388" pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph id="F6A14AFED8FD585D70A77D8A5D852C61" pageId="0" pageNumber="1">Additional material examined.</paragraph>
|
||
<paragraph id="E4E122C709336A2CF4236B20B6A31C02" pageId="0" pageNumber="1">
|
||
Indonesia, North Sulawesi, Bahoi,
|
||
<geoCoordinate id="105830C074D33D06CA206E14C95E4F78" direction="north" orientation="latitude" precision="9" value="1.7226667">01°43.36'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="BD43EC66BF3ADA35659BC8813C20925F" direction="east" orientation="longitude" precision="9" value="125.0205">125°01.23'E</geoCoordinate>
|
||
, 5 specimens 37/15 mm [2170], 30/15 mm [2145], 26/16 mm [2164], 25/14 mm [2157] and 25/11 mm [2177], st 85, sand and small rocks outside a mangrove (UMIZ 00049); North Sulawesi, Bahoi,
|
||
<geoCoordinate id="2FBC0FE756EE92118E1B2CA33EA126DE" direction="north" orientation="latitude" precision="9" value="1.7226667">01°43.36'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3D250C2B02E096290C12C1FEEDF0CF14" direction="east" orientation="longitude" precision="9" value="125.0205">125°01.23'E</geoCoordinate>
|
||
, 3 specimens 30/18 mm [2221], 29/16 mm [2220] and 26/17 mm [2228], st 88, sand and small rocks outside a mangrove (UMIZ 00050); North Sulawesi, Wori,
|
||
<geoCoordinate id="B66726BF7A3885EA18777CDE4FBC0143" direction="north" orientation="latitude" precision="9" value="1.6010001">01°36.06'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4F59742C2027F54A2322FDE6AA4925DC" direction="east" orientation="longitude" precision="9" value="124.86217">124°51.73'E</geoCoordinate>
|
||
, 1 specimen 33/22 mm [2283], st 90, old
|
||
<taxonomicName id="01303EF53ECAC713BADCDBC3EA6FC729" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
,
|
||
<taxonomicName id="D124AF83A10828A71333C729C53D2FA3" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
,
|
||
<taxonomicName id="6435E114BC154198E0F9362C70B14888" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
mangrove (UMIZ 00038); North Sulawesi, Mantehage Island,
|
||
<geoCoordinate id="70534304A82E2C7323CFB6E9ED36AB0A" direction="north" orientation="latitude" precision="9" value="1.698">01°41.88'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C2D4B0BA83B8697266B72DFD2C3656C5" direction="east" orientation="longitude" precision="9" value="124.779">124°46.74'E</geoCoordinate>
|
||
, 1 specimen 25/16 mm [2330], st 91, rocks behind a
|
||
<taxonomicName id="D781154F3FD5354C7B3CE0C2F23C4A07" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
and
|
||
<taxonomicName id="84E74F98FED6CF2611944EA4EEF9CFEB" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
mangrove (UMIZ 00039); Ambon, Haruku Island,
|
||
<geoCoordinate id="8A60613D3A202CEC05DD77DC6CA48068" direction="south" orientation="latitude" precision="9" value="-3.6086667">03°36.52'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1A7192560C1C5AD37FF8428547011AC8" direction="east" orientation="longitude" precision="9" value="128.41783">128°25.07'E</geoCoordinate>
|
||
, 1 specimen 9/8 mm [2727], st 127, rocky
|
||
<taxonomicName id="C4F1070CEAAC23CA8E531A9A9759185E" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
mangrove with coral rubble (UMIZ 00040); Maluku, Kei Islands, Tual City, Fiditan,
|
||
<geoCoordinate id="FA9703CA838938251DFC091A1D4FB7D6" direction="south" orientation="latitude" precision="9" value="-5.5993333">5°35.96'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="89B46D44D09E8EA818504A11E758EEBC" direction="east" orientation="longitude" precision="9" value="132.75183">132°45.11'E</geoCoordinate>
|
||
, 1 specimen 17/9 mm [2939], st 144, rocks behind muddy mangrove of
|
||
<taxonomicName id="81F6242A1836EC76AD68D85D4F0EF4FA" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
trees (UMIZ 00041); Lombok, Seriwe Bay,
|
||
<geoCoordinate id="31657AFB157D7FAFACA4D89CD5D2CD85" direction="south" orientation="latitude" precision="9" value="-8.861667">08°51.70'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="7A898A23012FCEE5956AA25A4D6D3F47" direction="east" orientation="longitude" precision="9" value="116.54784">116°32.87'E</geoCoordinate>
|
||
, 1 specimen 18/10 mm [2966], st 147, small beach of coral rubble and rocks by bay (UMIZ 00042); Lombok, Seriwe Bay,
|
||
<geoCoordinate id="93E89E1E5B01AE1193F4824BFABF8FCD" direction="south" orientation="latitude" precision="9" value="-8.909166">08°54.55'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1C0947765F4552EDFE37FA0B001875CF" direction="east" orientation="longitude" precision="9" value="116.37033">116°22.22'E</geoCoordinate>
|
||
, 2 specimens 43/20 mm [2970] and 35/24 mm [2974], st 148, rocks behind mangrove with
|
||
<taxonomicName id="C2836AE88A23A87DF88A71C605716A01" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
trees (UMIZ 00043); Lombok, Don Don,
|
||
<geoCoordinate id="3F2575940E74DFF58130EA4B7E3D4CA0" direction="south" orientation="latitude" precision="9" value="-8.909">08°54.54'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="7A3C6C3ECFBD694188EEA397F8FC666F" direction="east" orientation="longitude" precision="9" value="116.35833">116°21.50'E</geoCoordinate>
|
||
, 2 specimens 35/19 mm [2986] and 25/13 mm [2978] st 149, old
|
||
<taxonomicName id="CF682B01B0E745F7EA291C59F7905203" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
forest with coral rubble (UMIZ 00051); Bali, Gilimanuk,
|
||
<geoCoordinate id="945455DF07EC1E5EFF41E85258B19FFD" direction="south" orientation="latitude" precision="9" value="-8.171">08°10.26'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9ED82B8785BFE232520DB90989A49EE6" direction="east" orientation="longitude" precision="9" value="114.4435">114°26.61'E</geoCoordinate>
|
||
, 1 specimen 18/14 mm [3591], st 155, large rocks near a patch of
|
||
<taxonomicName id="23F3A8064FA452389EFA25EA21EE3CE3" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
(UMIZ 00052); Bali, Pemuteran, Labuhan Lalang Harbor,
|
||
<geoCoordinate id="0E1CEBB9E49BD9D9F23C03F7FA576361" direction="south" orientation="latitude" precision="9" value="-8.1435">08°08.61'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3B0658FA46A1BB94BC5E90D979A9168C" direction="east" orientation="longitude" precision="9" value="114.53883">114°32.33'E</geoCoordinate>
|
||
, 3 specimens 22/12 mm [3137], 20/11 mm [3132], and 14/8 mm [3130], st 157, coral rubble, rocks and a few
|
||
<taxonomicName id="B5A3BC2904BCFCA48EA1C07734D88EDC" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
(UMIZ 00053); North Maluku, Ternate, Bastiong,
|
||
<geoCoordinate id="DED8B9220730E6AFFBE1C20535EC3E39" direction="north" orientation="latitude" precision="9" value="0.7735">00°46.41'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5A234E7346C48D72F9F703BCDDD2BF40" direction="east" orientation="longitude" precision="9" value="127.37933">127°22.76'E</geoCoordinate>
|
||
, 1 specimen 38/17 mm [5057], st 203, muddy rocks near a mangrove (UMIZ 00046); Halmahera, Akelamo,
|
||
<geoCoordinate id="D7D78375713A7A15491C02CD244A5755" direction="north" orientation="latitude" precision="9" value="1.0221666">01°01.33'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9D8102901F7B37BA3E3C03732837775F" direction="east" orientation="longitude" precision="9" value="127.6515">127°39.09'E</geoCoordinate>
|
||
, 1 specimen 35/20 mm [5078], st 207, sandy-muddy beach at margin of mangrove (UMIZ 00047); Halmahera, Foli,
|
||
<geoCoordinate id="199F3276F588A8624888F1798110A758" direction="north" orientation="latitude" precision="9" value="1.2443333">01°14.66'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="812AD0435DEB50F518E202A3F163DCB0" direction="east" orientation="longitude" precision="9" value="128.17683">128°10.61'E</geoCoordinate>
|
||
, 1 specimen 28/16 mm [5121], st 217, rocky shore near a beach (UMIZ 00048); Halmahera, Foli,
|
||
<geoCoordinate id="0D30A06A438DE43EC9F5045B003D6C2D" direction="north" orientation="latitude" precision="9" value="1.2443333">01°14.66'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="29A26F642AEDD5F879F0ADA3FA75805B" direction="east" orientation="longitude" precision="9" value="128.17683">128°10.61'E</geoCoordinate>
|
||
, 1 specimen 25/10 mm [5125], st 217, rocky shore near a beach (UMIZ 00054). Philippines, Luzon, Batangas, Calatagan,
|
||
<geoCoordinate id="2642DB224BC0FED406E38E441CD83D76" direction="north" orientation="latitude" precision="9" value="13.922">13°55.32'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3AAB9E448BEF9A91DEC1744EE842FBDB" direction="east" orientation="longitude" precision="9" value="120.621">120°37.26'E</geoCoordinate>
|
||
, 3 specimens 30/16 mm [3215], 29/16 mm [3212] and 26/12 mm [3210], st 183, rocks in
|
||
<taxonomicName id="E84B5A0BE548AF943BEDFCB2EAF1603F" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
and
|
||
<taxonomicName id="D1E819208A87D2972EF580492017873A" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
mangrove (PNM 041221); Bohol, Maribojoc, 09°44.02'N, E 123°47.45'E, 6 specimens 36/20 mm [3266], 34/21 mm [3271], 33/18 mm [3265], 33/22 mm [3272], 32/20 mm [3277], and 30/17 mm [3270], st 191, uplifted coral rubble with sand (PNM 041222); Bohol, Mabini,
|
||
<geoCoordinate id="DA078D645AF5759C3BD2FFBE245A0632" direction="north" orientation="latitude" precision="9" value="9.859834">09°51.59'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E924954D22ED4FE598CED79FE9C5AE7C" direction="east" orientation="longitude" precision="9" value="124.569336">124°34.16'E</geoCoordinate>
|
||
, 2 specimens 21/12 mm [3373] and 14/8 mm [3370], st 196, open
|
||
<taxonomicName id="D7A1F7CF54D7EAA77FB35E6AF9742F06" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
and
|
||
<taxonomicName id="134DCDA60E9B83467D6B376459B2FFEC" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
mangrove with sand, and coral rubble (PNM 041223); Bohol, Guindulman,
|
||
<geoCoordinate id="64269A03ED735C79C1E9E53BCA1B7385" direction="north" orientation="latitude" precision="9" value="9.734333">09°44.06'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="55119E9658A79F727E776BF86BEA4CA8" direction="east" orientation="longitude" precision="9" value="124.4605">124°27.63'E</geoCoordinate>
|
||
, 2 specimens 38/15 mm [3624] and 22/13 mm [3629], st 197, rocks and coral rubble near a few
|
||
<taxonomicName id="737C2CA4E01555AF8DE3022F87CD4B88" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
trees (PNM 041216); Bohol, Loay,
|
||
<geoCoordinate id="F62536B488D9BECD8FAE573C6F4368EF" direction="north" orientation="latitude" precision="9" value="9.603833">09°36.23'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B30EE3FFD2A4149E01C6D60634ECE1A2" direction="east" orientation="longitude" precision="9" value="123.9955">123°59.73'E</geoCoordinate>
|
||
, 1 specimen 33/19 mm [3634], st 198, mostly sand, and a few
|
||
<taxonomicName id="F2C61A38541C1D8BCCD965BE6D5CAA1A" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
(PNM 041217); Bohol, Maribojoc,
|
||
<geoCoordinate id="1640B6256F77EC4349C24DCA1874745C" direction="north" orientation="latitude" precision="9" value="9.733666">09°44.02'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F36466C39D7EB398A0D9FD8E9E312DA6" direction="east" orientation="longitude" precision="9" value="123.79083">123°47.45'E</geoCoordinate>
|
||
, 2 specimens 29/16 mm [3393] and 19/12 mm [3404], st 200, coral rubble with sand, at night (PNM 041218); Bohol, Maribojoc,
|
||
<geoCoordinate id="5FFE6A01CBED29384E84EA56561F2D17" direction="north" orientation="latitude" precision="9" value="9.738">09°44.28'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4B25AD00B4B4D0B6D230EDC474A1EBF0" direction="east" orientation="longitude" precision="9" value="123.823166">123°49.39'E</geoCoordinate>
|
||
, 4 specimens 37/17 mm [3420], 33/23 mm [3406], 26/16 mm [3405] and 21/11 mm [3408], st 202, uplifted coral rubble with sand and algae, near
|
||
<taxonomicName id="98B324EF3C5E1621B20E2F29D93BD9AD" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
trees (PNM 041224).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2865A63D3F1FC6ABCD2C98BA5D029C1" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="D0EEDE11A14D34F014562D516E0375EC" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="72C5247511908CCF45E262502EAB21A0" pageId="0" pageNumber="1">
|
||
Indonesia: Bali, Edam (near Jakarta), Halmahera, Kei Islands (type locality of
|
||
<taxonomicName id="B1E008B77F7DC01F404B756836A1EEB5" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
), Lombok, Seram, Sulawesi, and West Papua (type locality of
|
||
<taxonomicName id="2AF1F725F8A967ED47E8FF1DDD00882A" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
). Philippines: Bohol, Luzon, and Mindanao. All records are new except for the type localities as well as the presence of
|
||
<taxonomicName id="B8A380BE062B63B9E1436269185E0C77" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
in Edam and Mindanao (
|
||
<bibRefCitation id="67B54913CAE5D4CF0836DDCF0ABC398C" author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher (Jena)" pageId="0" pageNumber="1" pagination="29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
|
||
: 81).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="345D75D67DB1956B758B4807F49A1916" pageId="0" pageNumber="1" type="habitat">
|
||
<paragraph id="B344AD40A4768DF516648B01D3D9015F" pageId="0" pageNumber="1">Habitat</paragraph>
|
||
<paragraph id="6C717C3821D70D62DA504E3DA7BD6473" pageId="0" pageNumber="1">
|
||
(Figs 34-35, Table 3).
|
||
<taxonomicName id="93D7586E4157F969BC0AECD8597F1976" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
is found in the intertidal on coral rubble and small stones. It is often in patches of mangrove trees with coral rubble (Fig. 34), but may also be found in intertidal regions without mangrove trees (Fig. 35).
|
||
</paragraph>
|
||
<caption id="030A46A2E754BFECA9A4450B2290428C" pageId="0" pageNumber="1">
|
||
<paragraph id="DE5E9F40126E1746A11D6A698729F2EC" pageId="0" pageNumber="1">
|
||
Figure 34. Mangrove habitats,
|
||
<taxonomicName id="D9BA26E25A4E4BF614568C82999030C2" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
.
|
||
<normalizedToken id="B85618DB0A4887B536DDDCFBB3BA7C9C" originalValue="A–B">A-B</normalizedToken>
|
||
Indonesia, Bali, Pemuteran, Labuhan Lalang Harbor, coral rubble, rocks and a few
|
||
<taxonomicName id="E3403F310103ABBE5CE88A3C6F1257E5" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
(st 157)
|
||
<normalizedToken id="60D8E4C372A60F5A3B5EA58276A4819E" originalValue="C–D">C-D</normalizedToken>
|
||
Indonesia, Lombok, Don Don, old
|
||
<taxonomicName id="D01B5A31F18B962DF7E69AFF43CBCCD7" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
forest, with coral rubble (st 149)
|
||
<normalizedToken id="2BBACB23967E36C3CB506DC298BF5D09" originalValue="E–F">E-F</normalizedToken>
|
||
Philippines, Bohol, Mabini, open
|
||
<taxonomicName id="D4B067F3E3D13274DCAAAEDE427F3DEE" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
and
|
||
<taxonomicName id="2E1A15BEF04CE04C6C3BE3EA0F161441" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
mangrove with sand and coral rubble (st 196).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="835F3980F1B6EB9725A953E81CDF4C6A" pageId="0" pageNumber="1">
|
||
<paragraph id="E9B7B2F6FB324A915E87226CDE358F54" pageId="0" pageNumber="1">
|
||
Figure 35. Coral rubble habitats,
|
||
<taxonomicName id="5D35846176F396189407C7263AAF4472" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
. A Indonesia, Sulawesi, Bahoi, sand and small rocks outside a mangrove (st 88) B Philippines, Bohol, Maribojoc, uplifted, dead, coral flat covered with sand and algae adjacent to a few mangrove trees (st 202) C Philippines, Bohol, Maribojoc, coral rubble of uplifted coral reef (st 191) D Indonesia, Halmahera, Foli, rocky shore by beach (st 217).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="2694A8E330B0C55876426CCC97E01E37" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="B75222B927899E4C2B10A91E833B4E41" pageId="0" pageNumber="1">Diagnosis</paragraph>
|
||
<paragraph id="5F7FEF550DA84AEF3F54A4691160FBFA" pageId="0" pageNumber="1">
|
||
(Table 5).
|
||
<taxonomicName id="653A15C8D464321E4216C87720828EE1" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
cannot be distinguished from other
|
||
<taxonomicName id="139FDBF2016E4D2F2676B6C56DC86C7A" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species based on external features. Grey or brown specimens cannot be distinguished from any other species, and brightly colored specimens cannot be distinguished from
|
||
<taxonomicName id="D935EBF9AE0AC2D7642BE2339A3AC529" lsidName="W. nangkauriense" pageId="0" pageNumber="1" rank="species" species="nangkauriense">W. nangkauriense</taxonomicName>
|
||
or
|
||
<taxonomicName id="69BAEEFEF4270945665A3F87623A9C6C" lsidName="W. graniferum" pageId="0" pageNumber="1" rank="species" species="graniferum">W. graniferum</taxonomicName>
|
||
. Internally, the large, coiled penis of
|
||
<taxonomicName id="0A61E99C55B6898ADA2CF0883BF1256D" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
with projections at its tip is quite peculiar and is diagnostic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="264DDB8353BA4AF03F289A1C84C228AA" pageId="0" pageNumber="1" type="color and morphology of live animals">
|
||
<paragraph id="4E5814F7AD5DD56941D28E3493CD19FB" pageId="0" pageNumber="1">Color and morphology of live animals</paragraph>
|
||
<paragraph id="69AA7498AE3DD3BED70331BC7CEB9F6F" pageId="0" pageNumber="1">(Figs 36-37). In sandy habitats, sand grains may be stuck to the dorsal notum. The dorsal coloration is highly variable, and often a mottling of two or three colors, of which the most common are brown, black, green, red, grey, and yellow. In some individuals, many small yellow dots are present on the dorsal notum. The color of the ocular tentacles may be yellow-orange, light brown or reddish brown. The hyponotum is cream colored, yellow-orange, yellow-grey or grey. The foot is yellow-grey, yellow-orange, or grey. The lateral sides of the foot may be the same color as the pedal sole (for example, Fig. 37H) or occasionally dark grey (Fig. 37D).</paragraph>
|
||
<caption id="38521BD2256E758B7F206616DD7B7276" pageId="0" pageNumber="1">
|
||
<paragraph id="21E07B265DD058D6DA8C84EA4829F8D9" pageId="0" pageNumber="1">
|
||
Figure 36. Live specimens, dorsal view,
|
||
<taxonomicName id="590C05D225558B507EBB4F8262D83BDB" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
. A 33 mm long [3265], Philippines, Bohol (PNM 041214) B Dorsal view, 34 mm long [3271], Philippines, Bohol (PNM 041222) C 26 mm long [3210], Philippines, Luzon (PNM 041213) D 26 mm long [3405], Philippines, Bohol (PNM 041219) E 26 mm long [2164], Indonesia, Sulawesi (UMIZ 00036) F 38 mm long [5057], Indonesia, Halmahera (UMIZ 00046) G Dorsal view, 22 mm long [3137], Indonesia, Bali (UMIZ 00053) H 25 mm long [2157], Indonesia, Sulawesi (UMIZ 00036).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CE1B190168BA5BDF0A3CC02F0DBD74FE" pageId="0" pageNumber="1">
|
||
<paragraph id="9FBED6223F5E851835D6D553DE619E45" pageId="0" pageNumber="1">
|
||
Figure 37. Live specimens,
|
||
<taxonomicName id="3DBE3292FA069503710780BD461D3792" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
. A Dorsal view, 43 mm long [2970], Indonesia, Lombok (UMIZ 00043) B Dorsal view, 37 mm long [2170], Indonesia, Sulawesi (UMIZ 00036) C Ventral view, 35 mm long [2974], Indonesia, Lombok (UMIZ 00043) D Ventral view, 36 mm long [3266], Philippines, Bohol (PNM 041214) E Ventral view, 26 mm long [3405], Philippines, Bohol (PNM 041219) F Ventral view, 33 mm long [3272], Philippines, Bohol (PNM 041214) G Ventral view, 32 mm long [3277], Philippines, Bohol (PNM 041222) H Ventral view, 37 mm long [3420], Philippines, Bohol (PNM 0412224).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="385855D2D28CD1D33C4AA54B9112DCF5" pageId="0" pageNumber="1" type="external morphology">
|
||
<paragraph id="1DFCAA75567A2075A23A3CFD8662C9BC" pageId="0" pageNumber="1">External morphology.</paragraph>
|
||
<paragraph id="BB6D0218121FDF9C1B90AF4C7C82B169" pageId="0" pageNumber="1">Between six and 18 papillae bear dorsal eyes (three or four per papilla). There is a retractable papilla with eyes in the center of the dorsal notum, which may be slightly raised above the dorsal surface.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B14F11E1F7EE5604A73144879FAA6B75" pageId="0" pageNumber="1" type="digestive system">
|
||
<paragraph id="D1B45D074784EC1557B3840FDF30DA6D" pageId="0" pageNumber="1">Digestive system</paragraph>
|
||
<paragraph id="C689D96AF107B94FB1BBBF3555FB41C1" pageId="0" pageNumber="1">
|
||
(Fig. 38, Table 4). Each radular row contains a rachidian tooth and two half rows of lateral teeth. Examples of radular formulae are presented in Table 4. The length of the rachidian teeth is approximately 20-25
|
||
<normalizedToken id="947F6BEF1FCDEC23E3F529F053F13D87" originalValue="µm">µm</normalizedToken>
|
||
, significantly smaller than that of the lateral teeth. The length of the hook of the lateral teeth gradually increases, from 40 to 70
|
||
<normalizedToken id="44BC6F13AC79CD5B0678EE58AD2ADC27" originalValue="µm">µm</normalizedToken>
|
||
, from the inner to the outer teeth (excluding the innermost and outermost lateral teeth which are significantly smaller). The intestinal loops are of type I.
|
||
</paragraph>
|
||
<caption id="742926EBD2F18CC3A7054350F5D9F318" pageId="0" pageNumber="1">
|
||
<paragraph id="6918928F333039849F108C24F6BD3C2F" pageId="0" pageNumber="1">
|
||
Figure 38. Radula,
|
||
<taxonomicName id="EBDD5E5C14BD30293E660F82CADABE32" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
, Indonesia (
|
||
<normalizedToken id="7AF77E1481F4A8ACF6BC00DC76BDE1A0" originalValue="A–B">A-B</normalizedToken>
|
||
) Bali [3137] (UMIZ 00053) (D) Sulawesi [2330] (UMIZ 00039) (C,
|
||
<normalizedToken id="743BD9ABC43943236B061355D52AB721" originalValue="E–F">E-F</normalizedToken>
|
||
) Sulawesi [2220] (UMIZ 00050). A Rachidian and innermost lateral teeth, scale bar 20
|
||
<normalizedToken id="9269939171AF440FF53EAA3A907E51BA" originalValue="μm">μm</normalizedToken>
|
||
B Rachidian and left innermost lateral teeth, scale bar 30
|
||
<normalizedToken id="F2FA0D8BE829501D8828DC6EBCDE9860" originalValue="μm">μm</normalizedToken>
|
||
C Rachidian and innermost lateral teeth, scale bar 50
|
||
<normalizedToken id="7EED9037D1357488F63ADF4CD2F29678" originalValue="μm">μm</normalizedToken>
|
||
D Rachidian and right innermost lateral teeth, scale bar 50
|
||
<normalizedToken id="04CC72A116E2EB401B1CF802F932A3E4" originalValue="μm">μm</normalizedToken>
|
||
E Outermost lateral teeth, scale bar 10
|
||
<normalizedToken id="2DE18B97C1DC8119A0CAAB217F19EB96" originalValue="μm">μm</normalizedToken>
|
||
F Transition between inner lateral teeth and outer lateral teeth, scale bar 40
|
||
<normalizedToken id="5F89E5C6E1EF2702915B4383CB3FF9A2" originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="63A8A145F5B6F76E60EB75D00DEF822E" pageId="0" pageNumber="1" type="reproductive system">
|
||
<paragraph id="504939277AC29259D7AB9908F847330D" pageId="0" pageNumber="1">Reproductive system</paragraph>
|
||
<paragraph id="F485862256EBDDA2BEDC78D38F55A02C" pageId="0" pageNumber="1">(Fig. 39). Posteriorly, the oviduct is larger than the deferent duct and is especially wide distally. The spermatheca is spherical and enters the distal end of the oviduct through a short duct. In small specimens (less than 17 mm long) female reproductive organs are small, with a narrower oviduct.</paragraph>
|
||
<caption id="FFE806CB73C1D8EB54AEF0BCB947CCE8" pageId="0" pageNumber="1">
|
||
<paragraph id="9AF3DF5B3DA23423F56FDF49A058F20A" pageId="0" pageNumber="1">
|
||
Figure 39. Reproductive system, hermaphroditic (female), posterior parts,
|
||
<taxonomicName id="CA1C4BB4806F952ABC013FC5F20BE54B" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
, Indonesia A Lombok, scale bar 2 mm [2970] (UMIZ 00043) B Sulawesi, scale bar 3 mm [2220] (UMIZ 00050). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct rs receptaculum seminis sp spermatheca.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="CECBC4D3A69744F030AF11BEDC970A8D" pageId="0" pageNumber="1" type="copulatory apparatus">
|
||
<paragraph id="1E6BC5892F327F9FF55E4ADFAD5240FF" pageId="0" pageNumber="1">Copulatory apparatus</paragraph>
|
||
<paragraph id="D0100A43217E97C16E887A4CA79D0248" pageId="0" pageNumber="1">(Figs 40-42). The penis is smooth. It does not bear any hooks. In mature specimens (> 22 mm long), it is tightly coiled (Fig. 42C), with approximately eight to 15 loops in the coil (Figs 40A, B, 41). However, the number of loops in the coil is correlated with the body length (and sexual maturity) and there are fewer (less than five) or no loops in shorter (<22 mm long) and less mature specimens (Fig. 40D). When the tip of the penis is evaginated, it is tapered distally and bears three pointed projections (Figs 40A, 41A, 42A, B). The penis is enclosed within the vestibule (Figs 40C, 41B), and the coil may form a U-shape within the vestibule (Fig. 41A). The penis is sometimes longitudinally folded and it appears that its loops could expand in diameter (cross-section of penis, Fig. 42D). The vestibule is exceptionally large (it can be nearly half the length of the body cavity). The penial sheath is long and usually forms a loop within the body cavity. The deferent duct is exceptionally narrow and convoluted, and only becomes wider distally in the anterior region of the body cavity (Fig. 40). In small specimens, the deferent duct is less convoluted than in larger (and more mature) specimens. The retractor muscle is short (approximately 2 mm) and inserts on the posterior body wall, on the right side of the rectum.</paragraph>
|
||
<caption id="E4750FEBD466B36BE952F9E08CC851DC" pageId="0" pageNumber="1">
|
||
<paragraph id="2D1F229555FC964D22624C0D991802F6" pageId="0" pageNumber="1">
|
||
Figure 40. Reproductive system,
|
||
<taxonomicName id="447739A30F4CD375FF6FC3BA4E26703E" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
, Indonesia. A Anterior, male copulatory parts (vestibule removed, showing the coiled and U-shaped penis), Halmahera, scale bar 2 mm [5078] (UMIZ 00047) B Penis (coil of loops removed from vestibule), Sulawesi, scale bar 2 mm [2283] (UMIZ 00038) C Anterior, male copulatory parts, scale bar 5 mm, same as B D Anterior, male copulatory parts, Kei Islands, scale bar 1 mm [2939] (UMIZ 00041). Abbreviations: dd deferent duct dpp distal penial projections p penis ps penial sheath rm retractor muscle v vestibule.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3A3FE4B527395220C6288D06C850F938" pageId="0" pageNumber="1">
|
||
<paragraph id="DDEF70B8E410036607D5503C2BA79963" pageId="0" pageNumber="1">
|
||
Figure 41. Reproductive system, anterior, male copulatory parts,
|
||
<taxonomicName id="816933D700B23E334F441955AEECACCE" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
. A Vestibule removed, showing the coiled and U-shaped penis, Indonesia, Sulawesi, scale bar 3 mm [2220] (UMIZ 00050) B Vestibule with coiled penis inside, Philippines, Bohol, scale bar 3 mm [3270] (PNM 041220). Abbreviations: dd deferent duct dpp distal penial projections p penis ps penial sheath rm retractor muscle v vestibule.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EF347B75A823696A6E694D0A14515D4D" pageId="0" pageNumber="1">
|
||
<paragraph id="CA880DD4EE61ED0496F4A0E8BD7EC1BA" pageId="0" pageNumber="1">
|
||
Figure 42. Penis,
|
||
<taxonomicName id="6E630B056362BF153B326C6058BB1593" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
, Indonesia. A Detail of distal portion when fully relaxed (with distal penial projections) and partial view of the coil, Indonesia, Halmahera, scale bar 500
|
||
<normalizedToken id="A3509FCDE82A378B871FC2CDC4F38FF2" originalValue="μm">μm</normalizedToken>
|
||
[3393] (PNM 041218) B Detail of distal portion when fully relaxed (with distal penial projections) and partial view of the coil, Halmahera, scale bar 500
|
||
<normalizedToken id="167147F923C3218C65EF56362E7E720E" originalValue="μm">μm</normalizedToken>
|
||
[5125] (UMIZ 00054) C Partial view of the coil (with six loops), Philippines, Bohol, scale bar 500
|
||
<normalizedToken id="1BCB0D97292179E2AEE2A87666AC2F00" originalValue="μm">μm</normalizedToken>
|
||
[3266] (PNM 041214) D Cross-section showing the longitudinal fold, Bali, scale bar 300
|
||
<normalizedToken id="67EE4302BD383F8AB7980EF5362348E1" originalValue="μm">μm</normalizedToken>
|
||
[3137] (UMIZ 00053).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="58B7156DE69BB045AFF67CAA97FD41B9" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="2C13461EC36C17F8E175A0F4EB97C137" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="C2374489437663EBF9D74257ACB04A68" pageId="0" pageNumber="1">
|
||
<normalizedToken id="4A6B1926B175FC23A1E3EF1F35E10D45" originalValue="Lesson’s">Lesson's</normalizedToken>
|
||
original description of
|
||
<taxonomicName id="897EE3FBA6E84931920CF961EF5552FD" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Onchidium ater</taxonomicName>
|
||
was brief, mostly based on the external morphology, and lacked illustrations of the internal anatomy. Fortunately, some organs remain in the syntypes which display the diagnostic characters of
|
||
<taxonomicName id="35B17EB07314F3BFC0D6379434E5BC66" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
(intestinal loops of type I and no rectal gland).
|
||
</paragraph>
|
||
<paragraph id="D6024E188F731E06A7EAE40AC030884A" pageId="0" pageNumber="1">
|
||
The male parts were removed from both syntypes by
|
||
<normalizedToken id="E02162F7B0FA7E3DE2076419FD8FD3EA" originalValue="Labbé">Labbe</normalizedToken>
|
||
when he re-described them for his 1934 monograph. Therefore, only
|
||
<normalizedToken id="B9B263AC204306AF19A403C6CCBE8CF0" originalValue="Lesson’s">Lesson's</normalizedToken>
|
||
original description of the male parts and their re-description by
|
||
<normalizedToken id="4719E0BA7032B1EBB8401C002FF88791" originalValue="Labbé">Labbe</normalizedToken>
|
||
can be used to determine the application of the name
|
||
<taxonomicName id="976B211300C490B26D6088BFD3C92D4D" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
.
|
||
<bibRefCitation id="BEFABAEF2D89079BE8A7B2B9E7C4A4DD" pageId="0" pageNumber="1">Lesson (1830</bibRefCitation>
|
||
: 300, our translation from French) describes the penis as a "very elongated, cylindrical, twisted upon itself, and very contractile tube in the anterior part of the body." Also,
|
||
<bibRefCitation id="4651E39E91FFE8E8D97A0ED3F6F9102D" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="A3FCC1C7194D37B656E0FBD44E872B8D" originalValue="Labbé’s">Labbe's</normalizedToken>
|
||
(1934
|
||
</bibRefCitation>
|
||
: fig. 30) illustration of the penis of the syntypes of
|
||
<taxonomicName id="723265B1785201305D546B591251B28C" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
reveals that it consists of a coil of tight loops, which perfectly matches the penis of the species described here. In addition, the wide oviduct in the syntypes is consistent with this species. As a result, we apply the name
|
||
<taxonomicName id="7C1F110F6BE1051E411EEFFA95D5D7E2" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
to the species described here.
|
||
</paragraph>
|
||
<paragraph id="9FC6B50D32921DDFFD36E59A69B33734" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="EDB84EBAF93C14A9449EB0920A3D2AFE" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="3D6810A1B8A42EBD453E42F683ECAF2A" originalValue="Labbé">Labbe</normalizedToken>
|
||
(1934
|
||
</bibRefCitation>
|
||
: 206) also described "feathery gills" in the two syntypes of
|
||
<taxonomicName id="180EB2D6D15A50543E18177AD5F0E243" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
, which, as a result, he transferred to the genus
|
||
<taxonomicName id="D08D8ACCF8175CFDB060C89BDD495577" class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Scaphis</taxonomicName>
|
||
.
|
||
<normalizedToken id="36786F05B1EC79E418685F1FF4D5C4D5" originalValue="Labbé">Labbe</normalizedToken>
|
||
commonly made surprising mistakes, describing features in specimens in which they were clearly not present. In this case, there are no gills on the dorsal notum of the syntypes of
|
||
<taxonomicName id="98B0A2A1A35B495B4EE9358DB5EA2D0F" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
.
|
||
<normalizedToken id="62358E84F10FF88CFA1406F9A5E55C5D" originalValue="Labbé">Labbe</normalizedToken>
|
||
also described a penial accessory gland in the syntypes of
|
||
<taxonomicName id="BC401ADCCB87F6961AC17BC14FA52888" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
. Because the male parts are missing (
|
||
<normalizedToken id="F54AF0FE40AE64D3D7AB7001240731FF" originalValue="Labbé">Labbe</normalizedToken>
|
||
removed them), it is something that we cannot verify. However, given that there is no known onchidiid species in any other genus with this type of penial anatomy,
|
||
<normalizedToken id="5CDE3F92F97B878F9D3AAE20B97AD24C" originalValue="Labbé’s">Labbe's</normalizedToken>
|
||
description of an accessory penial gland is regarded as one of his numerous mistakes. Because of the errors he made when he re-described the two syntypes of
|
||
<taxonomicName id="93F79089CBE2E1D3D03B3E9F495490D2" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
, only
|
||
<bibRefCitation id="9C51D73C9A6E1D38B219E5B836081281" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="1641D21A6FBAD2FF5CE6DE866470D2EC" originalValue="Labbé’s">Labbe's</normalizedToken>
|
||
(1934
|
||
</bibRefCitation>
|
||
: fig. 30) illustration of the penis is cited as a description of
|
||
<taxonomicName id="3C75A706FB192007DBD09AF9A6E844D5" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
in the list of references above.
|
||
</paragraph>
|
||
<paragraph id="93F9FB1AE79035F467DCD077782CFC5A" pageId="0" pageNumber="1">
|
||
The unique combination of characters observed on the lectotype of
|
||
<taxonomicName id="D682FA0BC8DF06E87F250004F4CA92FA" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium keiense" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="keiense">Onchidium keiense</taxonomicName>
|
||
(intestinal loops of type I, no rectal gland, male opening below the right tentacle, and no accessory penial gland) indicates that
|
||
<taxonomicName id="19435393C8D3BE8DE623EC94D24C2618" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
belongs to
|
||
<taxonomicName id="6226B50FD1AEF8B17CB87A631BADF6E0" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
. The re-examination of the lectotype and most paralectotypes available for
|
||
<taxonomicName id="C137BC9913CF314A6CFB84D0FE234B3D" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
revealed no difference with the two syntypes of
|
||
<taxonomicName id="E9F1F31ED026711AF509D6BEFEF3E6D5" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
for the penial anatomy and the enlarged oviduct, therefore
|
||
<taxonomicName id="3986D7F9076AD257B9209D3F18D2AB31" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
is identified here as a junior synonym of
|
||
<taxonomicName id="5A8924C7086C276922C0AE48A9313F14" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
. However, not all paralectotypes of
|
||
<taxonomicName id="E0465B7B3F2A42376D5B41F797F7059F" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
are part of
|
||
<taxonomicName id="26D2C78B83D39B91718D88EA4F1CC747" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
: at least one paralectotype of
|
||
<taxonomicName id="B1878B45CC9324FCBB5D2F3B298CEDA0" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
belongs to the genus
|
||
<taxonomicName id="C3176C45AD1D27E40676DF142447951C" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
|
||
(due to the presence of an accessory gland and a different intestinal type). The designation of the lectotype makes it clear that the name
|
||
<taxonomicName id="9832C567CD1154F2A90A9168DEA9E7F2" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
applies to the species described by Hoffmann without an accessory gland.
|
||
</paragraph>
|
||
<paragraph id="6F39C6D3028C83069A7924C29036A25A" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="7D10A9D0CDC56846492EDFA327CAF314" author="Hoffmann, H" journalOrPublisher="Videnskabelige Meddelelser Dansk Naturhistorisk Forening" pageId="0" pageNumber="1" pagination="15 - 36" title="Die Nacktschnecken der Daenischen Expedition nach den Kei-Inseln." volume="81" year="1926">Hoffmann (1926</bibRefCitation>
|
||
: 23) wrote that
|
||
<taxonomicName id="AFFDDA3D52D9AA927DE66752E18E16A7" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
was very close to
|
||
<taxonomicName id="C2F6CD36A795D506AB03A95D5C88307A" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
, but he was not convinced that the two were the same species because he thought that the cylindrical "organe excitateur" that Lesson described could have either been the penis or an accessory penial gland. If Hoffmann had been able to examine the syntypes of
|
||
<taxonomicName id="AB1FAA9540AB5F7403DB7AECDDD0E78B" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
, he would have seen that the penial morphology is identical to
|
||
<taxonomicName id="32CED25332C06DA2CE68AD6E7AD83C1E" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
.
|
||
<normalizedToken id="6DF3C7E21B1E7B42EBF5B192443A44D4" originalValue="Hoffmann’s">Hoffmann's</normalizedToken>
|
||
description of
|
||
<taxonomicName id="8BEFCBEC6E1AB0FAFE60A55CC66FAE37" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
is consistent with
|
||
<taxonomicName id="A6D622DB216E50478145E5E68633E0D9" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">Wallaconchis ater</taxonomicName>
|
||
, except for the description of dorsal eyes. The dorsal notum of
|
||
<taxonomicName id="0395AB403B8215D4516C65361B3B6C2A" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
bears many papillae with three or four dorsal eyes on each papilla, while Hoffmann described 12 large papillae on the dorsal notum of
|
||
<taxonomicName id="B8DBE1A37686C6E76B11BC616910217B" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
with one dorsal eye each, and the central papillae with three or four eyes. Although the papillae are retracted in the lectotype and many paralectotypes, the presence of three or four dorsal eyes on all visible papillae was confirmed.
|
||
</paragraph>
|
||
<paragraph id="AB4F58E19837C7AEFE6D85F345E34435" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="5986006944E88A3FB85C6EE8AAF4C644" author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher (Jena)" pageId="0" pageNumber="1" pagination="29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
|
||
: 81) examined additional specimens (he does not indicate how many) from Mindanao (Philippines) and Edam (a small Indonesian island near Jakarta) which he identified as
|
||
<taxonomicName id="49746D258515CCC703224F02A954359E" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
in his subsequent revision of the
|
||
<taxonomicName id="63C528AF4D8A73B7C4B2648EAC00E77E" family="Onchidiidae" lsidName="" pageId="0" pageNumber="1" rank="family">Onchidiidae</taxonomicName>
|
||
. Hoffmann did not comment on the anatomy of these additional specimens (which were not examined for the present study). However, given that he is the original author of
|
||
<taxonomicName id="74A84A6352AF50CC6CA143B3481D23E6" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
, it can be assumed that he recognized the distinct penial anatomy of this species. Mindanao is within the geographic range of
|
||
<taxonomicName id="B70C2DD3CC86A695F9E381F96D738641" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
. However, the presence of
|
||
<taxonomicName id="2983FAC95EFAF383718CC3BEAAFFC507" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
in Edam slightly expands to the west the geographic range delineated here based on our collections and expands the known distribution of
|
||
<taxonomicName id="2400F72C31B04C9A69646C1C19F6508F" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D4C233A183411AF3F7DAF75545535220" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="5D335BF0A58CB0354A0578FF38A4BA4D" author="Hoffmann, H" journalOrPublisher="Videnskabelige Meddelelser Dansk Naturhistorisk Forening" pageId="0" pageNumber="1" pagination="15 - 36" title="Die Nacktschnecken der Daenischen Expedition nach den Kei-Inseln." volume="81" year="1926">Hoffmann (1926</bibRefCitation>
|
||
: 24, our translation from German) considered creating a new genus for
|
||
<taxonomicName id="41CE389C48D1988425BC9567BC4B5F9F" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium keiense" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="keiense">Onchidium keiense</taxonomicName>
|
||
: "The peculiarity of the penis could almost suggest the idea that we are dealing here with a whole new genus. In the genus
|
||
<taxonomicName id="C783212AFF8BDFE3FC7FECF4FD980001" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidium</taxonomicName>
|
||
[spelled
|
||
<taxonomicName id="709A2239CC1881858B04E70267B266DB" class="Gastropoda" family="Onchidiidae" genus="Oncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Oncidium</taxonomicName>
|
||
], however, there are already species which differ quite considerably from each other, so that I have no reason to create a new genus for my species." Hoffmann recognized the similarity between some
|
||
<taxonomicName id="6696C0BD6C8F21B5BF3D1C017194738D" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidium</taxonomicName>
|
||
species transferred here to
|
||
<taxonomicName id="AD011EB6E3F497B4282CDDB4BA6F3407" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
(i.e.,
|
||
<taxonomicName id="4D5C0A23276BEA97A380328B1F577CFD" lsidName="O. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">O. buetschlii</taxonomicName>
|
||
,
|
||
<taxonomicName id="8E8D26D1AB853A2B4C7EF20388AB2809" lsidName="O. nangkauriense" pageId="0" pageNumber="1" rank="species" species="nangkauriense">O. nangkauriense</taxonomicName>
|
||
, and
|
||
<taxonomicName id="31286A61E8CF6FAD9AD01CE13BBC53D3" lsidName="O. ovale" pageId="0" pageNumber="1" rank="species" species="ovale">O. ovale</taxonomicName>
|
||
), but he did not comment on the similarity between
|
||
<taxonomicName id="73911474DB2A8A1370FF228A9AAE2239" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
and these species. However, in his identification key,
|
||
<bibRefCitation id="902D9F4019B5130BC5AE93FC4596F45A" author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher (Jena)" pageId="0" pageNumber="1" pagination="29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
|
||
: 109-110) listed
|
||
<taxonomicName id="0BA100E3129F3E7E8B45C931B225EA19" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
in the same part of the key as the
|
||
<taxonomicName id="9611B962CBD7BD69BDFF62BEA2274346" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidium</taxonomicName>
|
||
species transferred here to
|
||
<taxonomicName id="9ACA57BD15B39D701AA14C56E6E9BAA5" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
, using exactly the same characters: no accessory penial gland, no rectal gland, and intestinal loops of type I. So, even though Hoffmann showed in his identification key that those species are similar, he did not create or name any group for them. Also, in that same part of the key, Hoffmann listed
|
||
<taxonomicName id="B8B86D689FDF012071629B60409C1F23" lsidName="O. papuanum" pageId="0" pageNumber="1" rank="species" species="papuanum">O. papuanum</taxonomicName>
|
||
and
|
||
<taxonomicName id="EEC17F56F76573F50F62F7954E3F6A58" lsidName="O. palaense" pageId="0" pageNumber="1" rank="species" species="palaense">O. palaense</taxonomicName>
|
||
. He must have assumed that the intestinal loops in these two species are of type I because there is no indication in the original descriptions that their intestinal loops are actually of type I and Hoffmann did not comment on the type material (which is lost for both species). However, the position of the male opening in the original descriptions of
|
||
<taxonomicName id="718FDAC94F7F3EE5A91D27C26FA5788F" lsidName="O. papuanum" pageId="0" pageNumber="1" rank="species" species="papuanum">O. papuanum</taxonomicName>
|
||
and
|
||
<taxonomicName id="E457DD35C3140F0E9FE9576BF50686D5" lsidName="O. palaense" pageId="0" pageNumber="1" rank="species" species="palaense">O. palaense</taxonomicName>
|
||
(to the left of right tentacle) indicates that these two names do not apply to
|
||
<taxonomicName id="3B0C44C4E3A13D142061667268C5C171" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
<paragraph id="695185FC6185788604C7CCFB8DE0E0DC" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="C83FC26634193370F7754C484EEFA7B2" author="Hoffmann, H" journalOrPublisher="Videnskabelige Meddelelser Dansk Naturhistorisk Forening" pageId="0" pageNumber="1" pagination="15 - 36" title="Die Nacktschnecken der Daenischen Expedition nach den Kei-Inseln." volume="81" year="1926">Hoffmann (1926</bibRefCitation>
|
||
:23) wrote that
|
||
<taxonomicName id="3CD494048F00CCF23A64C6215FE0DC97" lsidName="W. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">W. keiense</taxonomicName>
|
||
seemed to be like
|
||
<taxonomicName id="BC5D317EE9AAE24D542252099CC86560" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium cinereum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cinereum">Onchidium cinereum</taxonomicName>
|
||
Quoy & Gaimard, 1832, but his comparison was based on
|
||
<bibRefCitation id="0A6028BB1D3FAC6F3EA413034CA9D2C7" pageId="0" pageNumber="1">
|
||
<normalizedToken id="4873DF0930B25055BF8C9BAD405664D5" originalValue="Semper’s">Semper's</normalizedToken>
|
||
(1880
|
||
</bibRefCitation>
|
||
: pl. 20, fig. 11, pl. 23, fig. 13; 1882:286) description of new specimens from Tonga-tabu [Tongatapu Island, Tonga] which he received from the Museum Godeffroy, instead of Quoy and
|
||
<normalizedToken id="9231E017629722E76B207E8D580D6C4A" originalValue="Gaimard’s">Gaimard's</normalizedToken>
|
||
original type material (from Tonga). The type material of
|
||
<taxonomicName id="C9DF4F0C2944F105B8A12CC6B29DA953" lsidName="O. cinereum" pageId="0" pageNumber="1" rank="species" species="cinereum">O. cinereum</taxonomicName>
|
||
could not be located.
|
||
<taxonomicName id="F9B88F9D433D038EA4E2220F5AE3DA1E" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium cinereum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cinereum">Onchidium cinereum</taxonomicName>
|
||
should likely be regarded as a nomen dubium because and it is unclear to which genus it belongs. Hoffmann also found an enlarged oviduct in
|
||
<taxonomicName id="0FC5E36C6D30A6DE8F6208B0793EDE49" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium aberrans" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aberrans">Onchidium aberrans</taxonomicName>
|
||
Semper, 1882 similar to the oviduct found in
|
||
<taxonomicName id="29680EB0DE04EB5E43F1D9EA5CD6C46E" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
, but
|
||
<taxonomicName id="1D1323B94A1A6C4CF18DE486615FCC5C" lsidName="O. aberrans" pageId="0" pageNumber="1" rank="species" species="aberrans">O. aberrans</taxonomicName>
|
||
does not belong to
|
||
<taxonomicName id="4C1CFC0DED667A5ACDF065A2705545BC" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
as it lacks the diagnostic characters of
|
||
<taxonomicName id="2FE0A29BB5D059518F8A2827189F51B3" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DD2E2B329E8F987E9E3ECA3380055DD6" pageId="0" pageNumber="1">
|
||
Hoffmann did not illustrate the coiled penis of
|
||
<taxonomicName id="A8765A6002F0055D40F730EA4A8F81F2" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
(the penial complex was illustrated with the vestibule and did not show the loops inside), which made it difficult for others to identify the diagnostic character of
|
||
<taxonomicName id="F7DF0D0152781F679A5A80521B8A5291" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
.
|
||
<normalizedToken id="5F29ABE653D76134BBCA123A4203839A" originalValue="Labbé">Labbe</normalizedToken>
|
||
did not recognize the similarity between the syntypes of
|
||
<taxonomicName id="CCD77627D4DBCB7630B33DCCA58BA4B6" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
which he examined and
|
||
<normalizedToken id="FA0F32DD6D8FFB4008ECB1B36B60E3C7" originalValue="Hoffmann’s">Hoffmann's</normalizedToken>
|
||
description of
|
||
<taxonomicName id="65178134088009813F4C8575ADEDAF79" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
.
|
||
<normalizedToken id="889B9437B9DAC2277EC1B6B516F1BB50" originalValue="Labbé">Labbe</normalizedToken>
|
||
moved
|
||
<taxonomicName id="1E0E492F058987987BB4152971C56B92" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
to his genus
|
||
<taxonomicName id="9433B4D53C5C2E7144BA763DF98221EF" class="Gastropoda" family="Onchidiidae" genus="Paraoncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraoncidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Paraoncidium</taxonomicName>
|
||
<normalizedToken id="8158BAA128DBE9D0549F790FB7C33CCC" originalValue="Labbé">Labbe</normalizedToken>
|
||
, 1934, which he created for a series of
|
||
<taxonomicName id="CA9EFA0F3FC8E11F0F8CABBB3CF9573A" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidium</taxonomicName>
|
||
species with no accessory gland and no rectal gland, but he did not examine any new material for
|
||
<taxonomicName id="708FC8CEDCC6A38C5D9483CF94CB5C6C" lsidName="O. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">O. keiense</taxonomicName>
|
||
. Note that
|
||
<taxonomicName id="3DB94B5859ED8303C3A425E359DBD626" class="Gastropoda" family="Onchidiidae" genus="Paraoncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraoncidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Paraoncidium</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="E6B00E864660B4FF6595C545DA68DDFE" class="Gastropoda" family="Onchidiidae" genus="Onchidina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidina" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidina</taxonomicName>
|
||
(
|
||
<bibRefCitation id="63271A3AAD715C9106C5103DD9A614D2" author="Dayrat, B" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="121 - 133" title="Systematics of the onchidiid slug Onchidinaaustralis (Mollusca: Gastropoda: Pulmonata)." url="https://doi.org/10.1127/arch.moll/146/121-133" volume="146" year="2017">Dayrat and Goulding 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="29EA3121F17F24477FB6E2CE4B02B6D5" pageId="0" pageNumber="1">
|
||
Another species,
|
||
<taxonomicName id="1A292C13544B560054F10FF9249E90FC" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium ferrugineum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ferrugineum">Onchidium ferrugineum</taxonomicName>
|
||
Lesson, 1830 was described from the same type locality as
|
||
<taxonomicName id="CCED0C5A2E0BE4E5600D36E3B97362E8" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
(Manokwari, West Papua, Indonesia). The number of syntypes was not included in
|
||
<normalizedToken id="7E9C949761219EA360E21E4B5F99BE01" originalValue="Lesson’s">Lesson's</normalizedToken>
|
||
original description, but
|
||
<bibRefCitation id="A7EF7DA10DCB4B2D6D6D64960FE069E8" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="FFA96182BC79B848BB79D28C024D1084" originalValue="Labbé">Labbe</normalizedToken>
|
||
(1934
|
||
</bibRefCitation>
|
||
: 214) indicates that he examined four syntypes, one of which was "completely emptied of its viscera." There are now only three syntypes left in the jar (36/25 mm, 35/19 mm, and 30/18 mm), suggesting that one of the syntypes was lost (MNHN 22951).
|
||
<bibRefCitation id="67B71F3DAA29E77B460D84203D200CEF" pageId="0" pageNumber="1">
|
||
<normalizedToken id="82DC8F9C1F0C31227BE25826C54AF40D" originalValue="Lesson’s">Lesson's</normalizedToken>
|
||
(1830
|
||
</bibRefCitation>
|
||
: 301) description of the penis of
|
||
<taxonomicName id="999938826D0670E22396453D52E45022" lsidName="O. ferrugineum" pageId="0" pageNumber="1" rank="species" species="ferrugineum">O. ferrugineum</taxonomicName>
|
||
as cylindrical and very twisted is remarkably similar to his description of the penis of
|
||
<taxonomicName id="38FFCE268945059F1FC6A1BB7F2C8A49" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
. However, Lesson also clearly specifies that gills are present on the dorsal notum. A re-examination of the three syntypes of
|
||
<taxonomicName id="00455A31FC067483C12BD2197A0D994E" lsidName="O. ferrugineum" pageId="0" pageNumber="1" rank="species" species="ferrugineum">O. ferrugineum</taxonomicName>
|
||
shows that they are part of two different genera: the largest syntype (36/25 mm) bears gills (only found in species of the genus
|
||
<taxonomicName id="3CB82087D17F122A7BF0370A2C6066D5" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
|
||
) while the two other syntypes do not. To clarify the application of
|
||
<taxonomicName id="E3C009D6D7421794EE2FDD1A0C30E17E" lsidName="O. ferrugineum" pageId="0" pageNumber="1" rank="species" species="ferrugineum">O. ferrugineum</taxonomicName>
|
||
, the largest syntype with gills is designated here as the lectotype (and now clearly labeled in the jar). Therefore, the name-bearing type of
|
||
<taxonomicName id="F031E892194A44ABFE2A2DFE695B802A" lsidName="O. ferrugineum" pageId="0" pageNumber="1" rank="species" species="ferrugineum">O. ferrugineum</taxonomicName>
|
||
is part of a
|
||
<taxonomicName id="E9EBCE23DD867E661C85F52E0014C330" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
|
||
species. Both paralectotypes display the diagnostic characters of
|
||
<taxonomicName id="3DF98338235291A71D809B2BCA23EF4A" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
, and the penis left in one of the paralectotypes is identical to the large, coiled penis of
|
||
<taxonomicName id="631967C35C2AEF6930CD00B51C199A51" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
and
|
||
<taxonomicName id="CF822FA7FEAF55BFF84855E85074769F" lsidName="W. keiense" pageId="0" pageNumber="1" rank="species" species="keiense">W. keiense</taxonomicName>
|
||
. However, because the paralectotypes no longer have any name-bearing function,
|
||
<taxonomicName id="E3527D9BEDE85E7D2356FA199850E2B1" lsidName="O. ferrugineum" pageId="0" pageNumber="1" rank="species" species="ferrugineum">O. ferrugineum</taxonomicName>
|
||
applies to a
|
||
<taxonomicName id="AC5FC208CE43839EE48948D6674AC939" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
|
||
species and cannot apply to a
|
||
<taxonomicName id="D0D22F82850180184572B686EB553C14" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species. Because
|
||
<normalizedToken id="006ABFFBA31F67F7A00B882D9FB78E04" originalValue="Labbé’s">Labbe's</normalizedToken>
|
||
re-description is based on specimens from two different species (and two different genera), only
|
||
<bibRefCitation id="C7B3D0149A813B8DD19D66DAAB43FB9B" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="4A4E25D8FD138F433A2A64E361FDD7D9" originalValue="Labbé’s">Labbe's</normalizedToken>
|
||
(1934
|
||
</bibRefCitation>
|
||
: fig. 50) illustration of the penis of a paralectotype of
|
||
<taxonomicName id="A25C03D5B499A47FA0B3CC1653A25AFE" lsidName="O. ferrugineum" pageId="0" pageNumber="1" rank="species" species="ferrugineum">O. ferrugineum</taxonomicName>
|
||
is cited as a description of
|
||
<taxonomicName id="9E251B04A8FC1C6C5E32CD4AFB7D1E0F" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
in the list of references above.
|
||
</paragraph>
|
||
<paragraph id="4545FB011AE5419D79750EE799806341" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="87278E121EBAC025B1C20D96C0451184" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919)</bibRefCitation>
|
||
translated
|
||
<normalizedToken id="8A6952F86AC8F264933041B3632D91BB" originalValue="Lesson’s">Lesson's</normalizedToken>
|
||
description of
|
||
<taxonomicName id="8488303E7A218DDB69FADF555EAB2FBF" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
but did not examine any new material.
|
||
<bibRefCitation id="8289D1A248C7401071AE5B202FF29FA0" author="Gray, JE" journalOrPublisher="Longman, Brown, Green and Longmans, London. Vol. 4" pageId="0" pageNumber="1" title="Figures of molluscous animals selected from various authors. Etched for the use of students by M. E. Gray." year="1850">Gray (1850)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="CAD148F542305B39317B8D7F40E5BDC2" pageId="0" pageNumber="1">Adams and Adams (1858)</bibRefCitation>
|
||
mentioned the name
|
||
<taxonomicName id="2D3179305442F0B4F5B1E268C5BD5224" class="Gastropoda" family="Onchidiidae" genus="Oncidiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncidiella nigra" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nigra">Oncidiella nigra</taxonomicName>
|
||
Lesson, but, as
|
||
<bibRefCitation id="D62E74B14E5F10838EF6461029A038C5" author="Tapparone-Canefri, C" journalOrPublisher="Annali del Museo Civico di Storia Naturale, Genova" pageId="0" pageNumber="1" pagination="1 - 313" title="Fauna malacologica delle Nuova Guinea e delle Isole adiacenti." volume="19" year="1883">Tapparone-Canefri (1883)</bibRefCitation>
|
||
first suggested, Lesson did not describe a species with this name, and these authors likely made the error of using the name
|
||
<taxonomicName id="60E7B64F40E2F38BDAA18D5BA9B27C6D" lsidName="O. nigra" pageId="0" pageNumber="1" rank="species" species="nigra">O. nigra</taxonomicName>
|
||
for
|
||
<taxonomicName id="09EB172A0F07591E96FBB7BF47577A9E" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
(both
|
||
<taxonomicName id="CBDD17DDFC8D00F9871CA94E3B9E3826" lsidName="ater" pageId="0" pageNumber="1" rank="species" species="ater">ater</taxonomicName>
|
||
and
|
||
<taxonomicName id="C2E124A44F30925F22C1D2F292418AEB" lsidName="nigra" pageId="0" pageNumber="1" rank="species" species="nigra">nigra</taxonomicName>
|
||
mean black in Latin).
|
||
<bibRefCitation id="F702873F27289E485CFA7B1E73D6922A" author="Gray, JE" journalOrPublisher="Longman, Brown, Green and Longmans, London. Vol. 4" pageId="0" pageNumber="1" title="Figures of molluscous animals selected from various authors. Etched for the use of students by M. E. Gray." year="1850">Gray (1850)</bibRefCitation>
|
||
did not include a description to accompany the name
|
||
<taxonomicName id="E9EBA9CD948534C950B89DD69918957B" lsidName="O. nigra" pageId="0" pageNumber="1" rank="species" species="nigra">O. nigra</taxonomicName>
|
||
, but included an illustration (pl. 181, fig. 7) of the dorsal notum of an animal which could be a
|
||
<taxonomicName id="51DF35306C284894D356FEE5DE02EC5F" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species but cannot be positively identified.
|
||
<bibRefCitation id="D9CCE822891403A621390DE85EBC35E4" pageId="0" pageNumber="1">Adams and Adams (1858)</bibRefCitation>
|
||
only listed the name
|
||
<taxonomicName id="CF504C92950ABCDC6E6611A93195C26D" lsidName="O. nigra" pageId="0" pageNumber="1" rank="species" species="nigra">O. nigra</taxonomicName>
|
||
without commenting on it. Finally,
|
||
<bibRefCitation id="CF379D7504E0D24C075C383FD3E86853" author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher (Jena)" pageId="0" pageNumber="1" pagination="29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="1A84FAC6C44310AD8ECEF8E4A1E512A6" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="3738BF74F5E70B88D5723E50D0FCA268" originalValue="Labbé">Labbe</normalizedToken>
|
||
(1934)
|
||
</bibRefCitation>
|
||
adopted
|
||
<normalizedToken id="AA13C0FB77CA6288085C4DC0BE3DFFE9" originalValue="Tapparone-Canefri’s">Tapparone-Canefri's</normalizedToken>
|
||
conclusion that
|
||
<taxonomicName id="95F773FE1C75B3D429429F149350F24D" lsidName="O. nigra" pageId="0" pageNumber="1" rank="species" species="nigra">O. nigra</taxonomicName>
|
||
was mistakenly used to refer to
|
||
<taxonomicName id="5298E9A5AF64D713B34B78608D567587" lsidName="O. ater" pageId="0" pageNumber="1" rank="species" species="ater">O. ater</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |