treatments-xml/data/14/7B/C4/147BC44BC44D2D5BFF69FB46C70C79AB.xml
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<document id="D14664A963E89BD370C9496DBCA76CFE" ID-CLB-Dataset="9446" ID-DOI="10.11646/zootaxa.4885.4.6" ID-GBIF-Dataset="c7579c32-46c4-4a34-86ee-cd06de94a790" ID-ISSN="1175-5326" ID-Zenodo-Dep="4296955" ID-ZooBank="42DEF6F1-9C16-48AD-BD61-8B156111C20D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606684780813" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2020" docId="147BC44BC44D2D5BFF69FB46C70C79AB" docLanguage="en" docName="zootaxa.4885.4.6.pdf" docOrigin="Zootaxa 4885 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Saphobranchia ilys Salazar-Vallejo 2020, n. sp." docType="treatment" docVersion="9" lastPageNumber="571" masterDocId="E842BC33C4472D50FFFEFFB8C4647F32" masterDocTitle="Four new deep-water flabelligerid species from Pacific Costa Rica (Annelida Sedentaria, Flabelligeridae)" masterLastPageNumber="578" masterPageNumber="560" pageNumber="570" updateTime="1698846847864" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5AA82A90D108F4C5E3EBE9F838C3023A">Four new deep-water flabelligerid species from Pacific Costa Rica (Annelida Sedentaria, Flabelligeridae)</mods:title>
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<mods:namePart id="EE523817E6097B37DB3A628EF28928E5">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:date id="397E76A4141A7EC165A2D0ED79D97241">2020</mods:date>
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<mods:number id="DE4F66CB50F11280EC0326EBEE11627F">2020-11-27</mods:number>
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<treatment id="147BC44BC44D2D5BFF69FB46C70C79AB" ID-DOI="http://doi.org/10.5281/zenodo.4329954" ID-GBIF-Taxon="170618219" ID-Zenodo-Dep="4329954" LSID="urn:lsid:plazi:treatment:147BC44BC44D2D5BFF69FB46C70C79AB" httpUri="http://treatment.plazi.org/id/147BC44BC44D2D5BFF69FB46C70C79AB" lastPageId="11" lastPageNumber="571" pageId="10" pageNumber="570">
<subSubSection id="D4C826D6C44D2D5AFF69FB46C7B27A0F" pageId="10" pageNumber="570" type="nomenclature">
<paragraph id="9C6D755DC44D2D5AFF69FB46C5AF7A2A" blockId="10.[151,982,1278,1377]" box="[151,459,1278,1305]" pageId="10" pageNumber="570">
<heading id="C725C231C44D2D5AFF69FB46C5AF7A2A" bold="true" box="[151,459,1278,1305]" fontSize="11" level="1" pageId="10" pageNumber="570" reason="1">
<emphasis id="AEA6A94FC44D2D5AFF69FB46C5AF7A2A" bold="true" box="[151,459,1278,1305]" pageId="10" pageNumber="570">
<taxonomicName id="5BD20EDEC44D2D5AFF69FB46C5E57A2B" ID-CoL="85RYH" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[151,385,1278,1305]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="10" pageNumber="570" phylum="Annelida" rank="species" species="ilys" status="sp. nov.">
<emphasis id="AEA6A94FC44D2D5AFF69FB46C5E57A2B" bold="true" box="[151,385,1278,1305]" italics="true" pageId="10" pageNumber="570">Saphobranchia ilys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44D2D5AFE76FB46C5AF7A2A" box="[392,459,1278,1304]" pageId="10" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9C6D755DC44D2D5AFF69FA9AC7B27A0F" blockId="10.[151,982,1278,1377]" box="[151,982,1314,1341]" pageId="10" pageNumber="570">
<uri id="E843795FC44D2D5AFF69FA9AC7B27A0F" box="[151,982,1314,1341]" pageId="10" pageNumber="570">
urn:lsid:zoobank.org:act:
<uuid id="E8744F88C44D2D5AFE55FA9AC7B27A0F" box="[427,982,1314,1341]" pageId="10" pageNumber="570">2D7FC8D5-5368-4CCC-B5A6-909FBB2B0C2F</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44D2D5AFF69FAFEC5287A52" box="[151,332,1350,1377]" pageId="10" pageNumber="570" type="description">
<paragraph id="9C6D755DC44D2D5AFF69FAFEC5287A52" blockId="10.[151,982,1278,1377]" box="[151,332,1350,1377]" pageId="10" pageNumber="570">
<figureCitation id="04E969D8C44D2D5AFF69FAFEC5667A53" box="[151,258,1350,1377]" captionStart="FIGURE 5" captionStartId="10.[151,250,1064,1088]" captionTargetBox="[172,1416,183,1038]" captionTargetId="figure-18@10.[170,1418,181,1040]" captionTargetPageId="10" captionText="FIGURE 5. Saphobranchia ilys n. sp., holotype (SIO A9605). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 14, right parapodium, posterior view. E. Same, notochaetal basal, medial and distal sections. F. Same, neurochaetal basal and distal sections (inset: neurochaetal tip). G. Posterior region, ventral view. Scale bars.- A: 2.1 mm, B: 1.1 mm, C: 1.5 mm, D: 0.65 mm, E, F: 17 μm, G: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296965" httpUri="https://zenodo.org/record/4296965/files/figure.png" pageId="10" pageNumber="570">Figures 5</figureCitation>
,
<figureCitation id="04E969D8C44D2D5AFEF0FAFEC5287A52" box="[270,332,1350,1376]" captionStart="FIGURE 8" captionStartId="14.[151,250,1797,1821]" captionTargetBox="[157,1430,849,1774]" captionTargetId="figure-328@14.[157,1430,849,1774]" captionTargetPageId="14" captionText="FIGURE 8. Living specimens. A. Lamispina polycerata n. sp., holotype (SIO A9842), anterior fragment, left lateral view. B. Same, posterior fragment, dorsal view. C. Saphobranchia canela n. sp., holotype (SIO A1332), dorsal view. D. Same, ventral view. E. Same, anterior end, frontal view (Br: branchia, Pa: palp). F. S. ilys n. sp., holotype (SIO A 9605), dorsal view. G. Same, ventral view. H. S. omorpha n. sp., holotype (SIO A9595), dorsal view. I. Same, ventral view. Scale bars: A, B: 1.0 mm; C, D: 2.2 mm; E: 0.4 mm; F, G: 3.5 mm; H, I: 3.7 mm (Photos: Greg Rouse)." figureDoi="http://doi.org/10.5281/zenodo.4296971" httpUri="https://zenodo.org/record/4296971/files/figure.png" pageId="10" pageNumber="570">8F, G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44D2D5AFF69FA36C5A37AC2" pageId="10" pageNumber="570" type="diagnosis">
<paragraph id="9C6D755DC44D2D5AFF69FA36C5A37AC2" blockId="10.[151,1437,1422,2025]" pageId="10" pageNumber="570">
<emphasis id="AEA6A94FC44D2D5AFF69FA36C56E7A9A" bold="true" box="[151,266,1422,1448]" pageId="10" pageNumber="570">Diagnosis</emphasis>
.
<taxonomicName id="5BD20EDEC44D2D5AFEE6FA36C5A77A9A" authorityName="Chamberlin" authorityYear="1919" box="[280,451,1422,1448]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="10" pageNumber="570" phylum="Annelida" rank="genus">
<emphasis id="AEA6A94FC44D2D5AFEE6FA36C5A77A9A" box="[280,451,1422,1448]" italics="true" pageId="10" pageNumber="570">Saphobranchia</emphasis>
</taxonomicName>
with tunic without sand particles; median chaetigers with notochaetae longer than body width; neurochaetae with tips falcate, basal anchylosed region ½-1/3 chaetal length, articles 45 times longer than wide; no gonopodial lobes.
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44D2D5AFF39FA42C03979B2" pageId="10" pageNumber="570" type="materials_examined">
<paragraph id="9C6D755DC44D2D5AFF39FA42C03979B2" blockId="10.[151,1437,1422,2025]" pageId="10" pageNumber="570">
<emphasis id="AEA6A94FC44D2D5AFF39FA42C5107926" bold="true" box="[199,372,1530,1556]" pageId="10" pageNumber="570">Type material</emphasis>
.
<materialsCitation id="2CBA7F00C44D2D5AFE79FA42C003790A" ID-GBIF-Occurrence="2988754301" accessionNumber="MW172262" collectingDate="2018-10-19" collectionCode="SIO, MW" collectorName="V. Orphan &amp; N. Gallo" elevation="1795" latitude="9.118" longLatPrecision="1" longitude="-84.840004" pageId="10" pageNumber="570" specimenCode="A9605" specimenCount="1" typeStatus="holotype">
<typeStatus id="4369CBFFC44D2D5AFE79FA42C58B7926" box="[391,495,1530,1556]" pageId="10" pageNumber="570" type="holotype">Holotype</typeStatus>
(
<collectionCode id="FAC3ED98C44D2D5AFDFAFA42C6577926" box="[516,563,1530,1556]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="10" pageNumber="570" type="University or college">SIO</collectionCode>
<specimenCode id="CC74DD26C44D2D5AFDC0FA42C6EA7927" box="[574,654,1530,1557]" collectionCode="SIO" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="10" pageNumber="570" type="University or college">A9605</specimenCode>
), CR18-0028,
<collectorName id="3127108BC44D2D5AFCC2FA42C7187926" box="[828,892,1530,1556]" pageId="10" pageNumber="570">Alvin</collectorName>
dive 4973 slurp,
<collectorName id="3127108BC44D2D5AFBA8FA43C0A37926" box="[1110,1223,1530,1557]" pageId="10" pageNumber="570">Jaco Scar</collectorName>
(
<geoCoordinate id="F9E6139AC44D2D5AFB25FA42C1FC7927" box="[1243,1432,1530,1557]" degrees="09" direction="north" minutes="07" orientation="latitude" pageId="10" pageNumber="570" precision="1" seconds="04.80" value="9.118">09°0704.80” N</geoCoordinate>
,
<geoCoordinate id="F9E6139AC44D2D5AFF69F9A6C536790B" box="[151,338,1566,1593]" degrees="84" direction="west" minutes="50" orientation="longitude" pageId="10" pageNumber="570" precision="1" seconds="24.00" value="-84.840004">84°5024.00” W</geoCoordinate>
),
<quantity id="5B2AD8B8C44D2D5AFE93F9A6C5A2790B" box="[365,454,1566,1593]" metricMagnitude="3" metricUnit="m" metricValue="1.795" pageId="10" pageNumber="570" unit="m" value="1795.0">
<elevation id="17FF926EC44D2D5AFE93F9A6C5A2790B" box="[365,454,1566,1593]" metricMagnitude="3" metricUnit="m" metricValue="1.795" pageId="10" pageNumber="570" unit="m" value="1795.0">1795 m</elevation>
</quantity>
,
<date id="E86C539DC44D2D5AFE2BF9A6C60B790A" box="[469,623,1566,1592]" pageId="10" pageNumber="570" value="2018-10-19">
<collectingDate id="F828AA75C44D2D5AFE2BF9A6C60B790A" box="[469,623,1566,1592]" pageId="10" pageNumber="570" value="2018-10-19">19 Oct. 2018</collectingDate>
</date>
,
<collectorName id="3127108BC44D2D5AFD82F9A7C695790A" box="[636,753,1566,1593]" pageId="10" pageNumber="570">V. Orphan</collectorName>
&amp;
<collectorName id="3127108BC44D2D5AFCE4F9A7C7E4790A" box="[794,896,1566,1593]" pageId="10" pageNumber="570">N. Gallo</collectorName>
, coll. [
<accessionNumber id="8381E8BEC44D2D5AFC31F9A7C03B790A" box="[975,1119,1565,1593]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW172262" pageId="10" pageNumber="570">
<collectionCode id="FAC3ED98C44D2D5AFC31F9A7C061790B" box="[975,1029,1567,1593]" country="Netherlands" name="Museum Wasmann" pageId="10" pageNumber="570">MW</collectionCode>
172262
</accessionNumber>
]
</materialsCitation>
.
<materialsCitation id="2CBA7F00C44D2D5AFB8DF9A7C03979B2" ID-GBIF-Occurrence="2988754304" collectingDate="2018-10-23" collectionCode="SIO" collectorName="E. Cordes &amp; J. Klein" elevation="1783" latitude="9.118" longLatPrecision="1" longitude="-84.840004" pageId="10" pageNumber="570" specimenCode="A9725" specimenCount="1" typeStatus="paratype">
<typeStatus id="4369CBFFC44D2D5AFB8DF9A7C080790B" box="[1139,1252,1567,1593]" pageId="10" pageNumber="570" type="paratype">Paratypes</typeStatus>
: Two fragments (
<collectionCode id="FAC3ED98C44D2D5AFF61F9FAC4AA796E" box="[159,206,1602,1628]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="10" pageNumber="570" type="University or college">SIO</collectionCode>
<specimenCode id="CC74DD26C44D2D5AFF2AF9FAC540796F" box="[212,292,1602,1629]" collectionCode="SIO" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="10" pageNumber="570" type="University or college">A9725</specimenCode>
), one anterior, one posterior, probably from same specimen, Alvin dive 4977, mussel pot 2, Jaco Scar (
<geoCoordinate id="F9E6139AC44D2D5AFF61F9DEC53279B3" box="[159,342,1638,1665]" degrees="09" direction="north" minutes="07" orientation="latitude" pageId="10" pageNumber="570" precision="1" seconds="04.80" value="9.118">09°0704.80” N</geoCoordinate>
,
<geoCoordinate id="F9E6139AC44D2D5AFE9CF9DEC67E79B3" box="[354,538,1638,1665]" degrees="84" direction="west" minutes="50" orientation="longitude" pageId="10" pageNumber="570" precision="1" seconds="24.00" value="-84.840004">84°5024.00” W</geoCoordinate>
),
<quantity id="5B2AD8B8C44D2D5AFDCCF9DEC6ED79B3" box="[562,649,1638,1665]" metricMagnitude="3" metricUnit="m" metricValue="1.783" pageId="10" pageNumber="570" unit="m" value="1783.0">
<elevation id="17FF926EC44D2D5AFDCCF9DEC6ED79B3" box="[562,649,1638,1665]" metricMagnitude="3" metricUnit="m" metricValue="1.783" pageId="10" pageNumber="570" unit="m" value="1783.0">1783 m</elevation>
</quantity>
,
<date id="E86C539DC44D2D5AFD6BF9DEC74D79B2" box="[661,809,1638,1664]" pageId="10" pageNumber="570" value="2018-10-23">
<collectingDate id="F828AA75C44D2D5AFD6BF9DEC74D79B2" box="[661,809,1638,1664]" pageId="10" pageNumber="570" value="2018-10-23">23 Oct. 2018</collectingDate>
</date>
,
<collectorName id="3127108BC44D2D5AFCCAF9DFC7C679B2" box="[820,930,1638,1665]" pageId="10" pageNumber="570">E. Cordes</collectorName>
&amp;
<collectorName id="3127108BC44D2D5AFC38F9DFC04479B2" box="[966,1056,1638,1665]" pageId="10" pageNumber="570">J. Klein</collectorName>
, coll.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44D2D5BFF39F932C7E57C6F" lastPageId="11" lastPageNumber="571" pageId="10" pageNumber="570" type="description">
<paragraph id="9C6D755DC44D2D5AFF39F932C6F879DE" blockId="10.[151,1437,1422,2025]" pageId="10" pageNumber="570">
<emphasis id="AEA6A94FC44D2D5AFF39F932C53D7996" bold="true" box="[199,345,1674,1700]" pageId="10" pageNumber="570">Description.</emphasis>
<typeStatus id="4369CBFFC44D2D5AFE9AF932C5A87996" box="[356,460,1674,1700]" pageId="10" pageNumber="570" type="holotype">Holotype</typeStatus>
(SIO A9605) mature female, without pygidium, greenish. Body anteriorly swollen, slightly bent laterally, tapered posteriorly (
<figureCitation id="04E969D8C44D2D5AFD82F916C6BF79FB" box="[636,731,1710,1737]" captionStart="FIGURE 5" captionStartId="10.[151,250,1064,1088]" captionTargetBox="[172,1416,183,1038]" captionTargetId="figure-18@10.[170,1418,181,1040]" captionTargetPageId="10" captionText="FIGURE 5. Saphobranchia ilys n. sp., holotype (SIO A9605). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 14, right parapodium, posterior view. E. Same, notochaetal basal, medial and distal sections. F. Same, neurochaetal basal and distal sections (inset: neurochaetal tip). G. Posterior region, ventral view. Scale bars.- A: 2.1 mm, B: 1.1 mm, C: 1.5 mm, D: 0.65 mm, E, F: 17 μm, G: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296965" httpUri="https://zenodo.org/record/4296965/files/figure.png" pageId="10" pageNumber="570">Figs 5A</figureCitation>
;
<figureCitation id="04E969D8C44D2D5AFD17F916C76E79FA" box="[745,778,1710,1736]" captionStart="FIGURE 8" captionStartId="14.[151,250,1797,1821]" captionTargetBox="[157,1430,849,1774]" captionTargetId="figure-328@14.[157,1430,849,1774]" captionTargetPageId="14" captionText="FIGURE 8. Living specimens. A. Lamispina polycerata n. sp., holotype (SIO A9842), anterior fragment, left lateral view. B. Same, posterior fragment, dorsal view. C. Saphobranchia canela n. sp., holotype (SIO A1332), dorsal view. D. Same, ventral view. E. Same, anterior end, frontal view (Br: branchia, Pa: palp). F. S. ilys n. sp., holotype (SIO A 9605), dorsal view. G. Same, ventral view. H. S. omorpha n. sp., holotype (SIO A9595), dorsal view. I. Same, ventral view. Scale bars: A, B: 1.0 mm; C, D: 2.2 mm; E: 0.4 mm; F, G: 3.5 mm; H, I: 3.7 mm (Photos: Greg Rouse)." figureDoi="http://doi.org/10.5281/zenodo.4296971" httpUri="https://zenodo.org/record/4296971/files/figure.png" pageId="10" pageNumber="570">8F</figureCitation>
);
<quantity id="5B2AD8B8C44D2D5AFCDEF916C71579FB" box="[800,881,1710,1737]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="10" pageNumber="570" unit="mm" value="31.0">31 mm</quantity>
long,
<quantity id="5B2AD8B8C44D2D5AFC45F916C07779FB" box="[955,1043,1710,1737]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="570" unit="mm" value="2.5">2.5 mm</quantity>
wide (widest by chaetigers 56, 4 mm), cephalic cage
<quantity id="5B2AD8B8C44D2D5AFE87F96AC5DE79DF" box="[377,442,1746,1773]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="570" unit="mm" value="8.0">8 mm</quantity>
long, 31 chaetigers.
</paragraph>
<paragraph id="9C6D755DC44D2D5AFF39F94EC6307892" blockId="10.[151,1437,1422,2025]" pageId="10" pageNumber="570">
Tunic papillated, sediment particles mostly fine, adherent along body, and few larger sand particles, especially dorsally, and some dorsal papillae basally swollen (
<figureCitation id="04E969D8C44D2D5AFD2AF8A2C74F7807" box="[724,811,1818,1845]" captionStart="FIGURE 5" captionStartId="10.[151,250,1064,1088]" captionTargetBox="[172,1416,183,1038]" captionTargetId="figure-18@10.[170,1418,181,1040]" captionTargetPageId="10" captionText="FIGURE 5. Saphobranchia ilys n. sp., holotype (SIO A9605). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 14, right parapodium, posterior view. E. Same, notochaetal basal, medial and distal sections. F. Same, neurochaetal basal and distal sections (inset: neurochaetal tip). G. Posterior region, ventral view. Scale bars.- A: 2.1 mm, B: 1.1 mm, C: 1.5 mm, D: 0.65 mm, E, F: 17 μm, G: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296965" httpUri="https://zenodo.org/record/4296965/files/figure.png" pageId="10" pageNumber="570">Fig. 5B</figureCitation>
). Venter with digitate to conical papillae, 34 times longer than wide. Papillae short, about 1/51/10 as long as notochaetae, about five rows per chaetiger, especially visible in median and posterior chaetigers, anterior dorsal surface slightly eroded. Venter with smaller papillae, digitate, about five rows per chaetiger (
<figureCitation id="04E969D8C44D2D5AFE13F83EC6237892" box="[493,583,1926,1952]" captionStart="FIGURE 8" captionStartId="14.[151,250,1797,1821]" captionTargetBox="[157,1430,849,1774]" captionTargetId="figure-328@14.[157,1430,849,1774]" captionTargetPageId="14" captionText="FIGURE 8. Living specimens. A. Lamispina polycerata n. sp., holotype (SIO A9842), anterior fragment, left lateral view. B. Same, posterior fragment, dorsal view. C. Saphobranchia canela n. sp., holotype (SIO A1332), dorsal view. D. Same, ventral view. E. Same, anterior end, frontal view (Br: branchia, Pa: palp). F. S. ilys n. sp., holotype (SIO A 9605), dorsal view. G. Same, ventral view. H. S. omorpha n. sp., holotype (SIO A9595), dorsal view. I. Same, ventral view. Scale bars: A, B: 1.0 mm; C, D: 2.2 mm; E: 0.4 mm; F, G: 3.5 mm; H, I: 3.7 mm (Photos: Greg Rouse)." figureDoi="http://doi.org/10.5281/zenodo.4296971" httpUri="https://zenodo.org/record/4296971/files/figure.png" pageId="10" pageNumber="570">Fig. 8G</figureCitation>
).
</paragraph>
<paragraph id="9C6D755DC44D2D5AFF39F812C78978F6" blockId="10.[151,1437,1422,2025]" box="[199,1005,1962,1989]" pageId="10" pageNumber="570">Anterior end features not visible; not dissected to avoid further damage.</paragraph>
<paragraph id="9C6D755DC44D2D5BFF39F876C0B37FE7" blockId="10.[151,1437,1422,2025]" lastBlockId="11.[151,1437,151,1689]" lastPageId="11" lastPageNumber="571" pageId="10" pageNumber="570">Cephalic cage chaetae 23 times longer than body width. Chaetigers 12 forming cephalic cage, chaetae direct-ed anteriorly; chaetiger 3 with less chaetae, notochaetae about half as long as those of chaetiger 2. Chaetae arranged in short rows, dorsolateral thin fascicles along chaetigers 47, in fan-shaped fascicles posteriorly.</paragraph>
<paragraph id="9C6D755DC44C2D5BFF39FF67C0B27EBB" blockId="11.[151,1437,151,1689]" pageId="11" pageNumber="571">Chaetiger 1 with 78 notochaetae, 34 neurochaetae, longest ones basally anchylosed, medially and distally articulated, shorter ones completely articulated, articles 34 times longer than wide medially, progressively longer distally. Chaetiger 2 with three thinner notochaetae, 2/3 as long as those of chaetiger 1, anchylosed section progressively shorter in smaller chaetae; neurochaetae shorter, 1/3 as long as those of chaetiger 1, articles 34 times longer than wide medially, progressively longer distally. Chaetigers 37 damaged, most chaetae broken.</paragraph>
<paragraph id="9C6D755DC44C2D5BFF39FE2BC1F77EC7" blockId="11.[151,1437,151,1689]" pageId="11" pageNumber="571">
Anterior dorsal margin of chaetiger 1 with 2 larger, conical papillae. Chaetigers 14 progressively longer. No chaetal transition from cephalic cage to body chaetae; all neurochaetae multiarticulate. Gonopodial lobes not seen (
<figureCitation id="04E969D8C44C2D5BFF5EFE63C49C7EC7" box="[160,248,474,501]" captionStart="FIGURE 5" captionStartId="10.[151,250,1064,1088]" captionTargetBox="[172,1416,183,1038]" captionTargetId="figure-18@10.[170,1418,181,1040]" captionTargetPageId="10" captionText="FIGURE 5. Saphobranchia ilys n. sp., holotype (SIO A9605). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 14, right parapodium, posterior view. E. Same, notochaetal basal, medial and distal sections. F. Same, neurochaetal basal and distal sections (inset: neurochaetal tip). G. Posterior region, ventral view. Scale bars.- A: 2.1 mm, B: 1.1 mm, C: 1.5 mm, D: 0.65 mm, E, F: 17 μm, G: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296965" httpUri="https://zenodo.org/record/4296965/files/figure.png" pageId="11" pageNumber="571">Fig. 5C</figureCitation>
). Oocytes seen in an ovary fragment, with oil droplets between them, each about 80100 μm in diameter.
</paragraph>
<paragraph id="9C6D755DC44C2D5BFF39FE47C1767D53" blockId="11.[151,1437,151,1689]" pageId="11" pageNumber="571">
Parapodia poorly developed, chaetae emerge from body wall. Parapodia lateral; median neuropodia ventrolateral. Notopodia with a larger, digitate postchaetal conical to digitate papillae along chaetigers 38; other notopodia and neuropodia with a few short papillae, each about as long as 1/51/10 notochaetal length (
<figureCitation id="04E969D8C44C2D5BFB55FDFFC1617D53" box="[1195,1285,582,609]" captionStart="FIGURE 5" captionStartId="10.[151,250,1064,1088]" captionTargetBox="[172,1416,183,1038]" captionTargetId="figure-18@10.[170,1418,181,1040]" captionTargetPageId="10" captionText="FIGURE 5. Saphobranchia ilys n. sp., holotype (SIO A9605). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 14, right parapodium, posterior view. E. Same, notochaetal basal, medial and distal sections. F. Same, neurochaetal basal and distal sections (inset: neurochaetal tip). G. Posterior region, ventral view. Scale bars.- A: 2.1 mm, B: 1.1 mm, C: 1.5 mm, D: 0.65 mm, E, F: 17 μm, G: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296965" httpUri="https://zenodo.org/record/4296965/files/figure.png" pageId="11" pageNumber="571">Fig. 5D</figureCitation>
).
</paragraph>
<paragraph id="9C6D755DC44C2D5BFF39FDD3C5A07C08" blockId="11.[151,1437,151,1689]" pageId="11" pageNumber="571">
Median notochaetae arranged in short transverse rows, chaetal fascicles fan-shaped, directed dorsally, up to twice longer than body width. All notochaetae multiarticulate capillaries; 910 per bundle, central notochaetae with basally anchylosed region up to half total length, basal chaetae fully articulated; articles 45 times longer than wide, becoming longer distally; tips straight (
<figureCitation id="04E969D8C44C2D5BFDB2FD6FC6C67DC3" box="[588,674,726,753]" captionStart="FIGURE 5" captionStartId="10.[151,250,1064,1088]" captionTargetBox="[172,1416,183,1038]" captionTargetId="figure-18@10.[170,1418,181,1040]" captionTargetPageId="10" captionText="FIGURE 5. Saphobranchia ilys n. sp., holotype (SIO A9605). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 14, right parapodium, posterior view. E. Same, notochaetal basal, medial and distal sections. F. Same, neurochaetal basal and distal sections (inset: neurochaetal tip). G. Posterior region, ventral view. Scale bars.- A: 2.1 mm, B: 1.1 mm, C: 1.5 mm, D: 0.65 mm, E, F: 17 μm, G: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296965" httpUri="https://zenodo.org/record/4296965/files/figure.png" pageId="11" pageNumber="571">Fig. 5E</figureCitation>
). Neurochaetae about as long as body width; 78 per bundle, anchylosed region 1/22/5 total length, medially with articles 34 times longer than wide, progressively shorter distally; tips falcate (
<figureCitation id="04E969D8C44C2D5BFEDFFCA7C5177C0B" box="[289,371,798,825]" captionStart="FIGURE 5" captionStartId="10.[151,250,1064,1088]" captionTargetBox="[172,1416,183,1038]" captionTargetId="figure-18@10.[170,1418,181,1040]" captionTargetPageId="10" captionText="FIGURE 5. Saphobranchia ilys n. sp., holotype (SIO A9605). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 14, right parapodium, posterior view. E. Same, notochaetal basal, medial and distal sections. F. Same, neurochaetal basal and distal sections (inset: neurochaetal tip). G. Posterior region, ventral view. Scale bars.- A: 2.1 mm, B: 1.1 mm, C: 1.5 mm, D: 0.65 mm, E, F: 17 μm, G: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296965" httpUri="https://zenodo.org/record/4296965/files/figure.png" pageId="11" pageNumber="571">Fig. 5F</figureCitation>
, inset).
</paragraph>
<paragraph id="9C6D755DC44C2D5BFF39FCFBC7E57C6F" blockId="11.[151,1437,151,1689]" box="[199,897,834,861]" pageId="11" pageNumber="571">
Posterior region cylindrical (
<figureCitation id="04E969D8C44C2D5BFDF7FCFBC6067C6F" box="[521,610,834,861]" captionStart="FIGURE 5" captionStartId="10.[151,250,1064,1088]" captionTargetBox="[172,1416,183,1038]" captionTargetId="figure-18@10.[170,1418,181,1040]" captionTargetPageId="10" captionText="FIGURE 5. Saphobranchia ilys n. sp., holotype (SIO A9605). A. Dorsal view. B. Anterior region, dorsal view. C. Anterior region, ventral view. D. Chaetiger 14, right parapodium, posterior view. E. Same, notochaetal basal, medial and distal sections. F. Same, neurochaetal basal and distal sections (inset: neurochaetal tip). G. Posterior region, ventral view. Scale bars.- A: 2.1 mm, B: 1.1 mm, C: 1.5 mm, D: 0.65 mm, E, F: 17 μm, G: 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4296965" httpUri="https://zenodo.org/record/4296965/files/figure.png" pageId="11" pageNumber="571">Fig. 5G</figureCitation>
), posterior end unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44C2D5BFF39FCDFC7717CFB" pageId="11" pageNumber="571" type="etymology">
<paragraph id="9C6D755DC44C2D5BFF39FCDFC7717CFB" blockId="11.[151,1437,151,1689]" pageId="11" pageNumber="571">
<emphasis id="AEA6A94FC44C2D5BFF39FCDFC52A7CB3" bold="true" box="[199,334,871,897]" pageId="11" pageNumber="571">Etymology.</emphasis>
The specific name is the Greek word for mud, feminine, is because of the presence of fine sediment particles in the tunic and covering papillae, as opposed to having sand particles on them. The specific name is regarded as a noun in apposition (
<bibRefCitation id="F84308ACC44C2D5BFDEFFC17C6F27CFB" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[529,662,943,969]" pageId="11" pageNumber="571" refId="ref15696" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature in association with the British Museum (Natural History), London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
, Art. 31.2).
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44C2D5BFF39FC6AC6937B93" pageId="11" pageNumber="571" type="description">
<paragraph id="9C6D755DC44C2D5BFF39FC6AC6937B93" blockId="11.[151,1437,151,1689]" pageId="11" pageNumber="571">
<emphasis id="AEA6A94FC44C2D5BFF39FC6AC55C7CDE" bold="true" box="[199,312,978,1004]" pageId="11" pageNumber="571">Variation</emphasis>
.
<materialsCitation id="2CBA7F00C44C2D5BFEB8FC6BC6977B93" ID-GBIF-Occurrence="2988754309" collectionCode="SIO" pageId="11" pageNumber="571" specimenCode="A 9725" specimenCount="1" typeStatus="paratype">
A
<typeStatus id="4369CBFFC44C2D5BFE9CFC6BC5A67CDF" box="[354,450,979,1005]" pageId="11" pageNumber="571" type="paratype">paratype</typeStatus>
(
<collectionCode id="FAC3ED98C44C2D5BFE2DFC6BC6667CDF" box="[467,514,979,1005]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="11" pageNumber="571" type="University or college">SIO</collectionCode>
<specimenCode id="CC74DD26C44C2D5BFDF7FC6BC63B7CDF" box="[521,607,978,1005]" collectionCode="SIO" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" lsid="urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="11" pageNumber="571" type="University or college">A 9725</specimenCode>
) is
<quantity id="5B2AD8B8C44C2D5BFD75FC6BC6AA7CDF" box="[651,718,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="571" unit="mm" value="8.0">8 mm</quantity>
long,
<quantity id="5B2AD8B8C44C2D5BFCE6FC6BC7147CDF" box="[792,880,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="571" unit="mm" value="2.8">2.8 mm</quantity>
wide, 10 chaetigers; anteriormost left chaetigers removed, integument with fine sediment particles making dorsal papillae look digitate to conical; epizoic organisms on chaetae making them look pilose. Cephalic cage chaetae
<quantity id="5B2AD8B8C44C2D5BFC95FBA3C7CF7B07" box="[875,939,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="571" unit="mm" value="6.0">6 mm</quantity>
long; chaetiger 1 with eight notochaetae and seven neurochaetae; chaetiger 2 with eight notochaetae, 4/5 as long as those of chaetiger 1, and seven neurochaetae, half as long as those of chaetiger 1. Chaetiger 3 with smaller chaetae, neurochaetae directed laterally. Posterior fragment with seven chaetigers,
<quantity id="5B2AD8B8C44C2D5BFE2BFB3FC6727B93" box="[469,534,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="571" unit="mm" value="4.0">4 mm</quantity>
long,
<quantity id="5B2AD8B8C44C2D5BFDA3FB3FC6D77B93" box="[605,691,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="571" unit="mm" value="1.4">1.4 mm</quantity>
wide
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44C2D5BFF39FB13C79A7947" pageId="11" pageNumber="571" type="discussion">
<paragraph id="9C6D755DC44C2D5BFF39FB13C14A7A4B" blockId="11.[151,1437,151,1689]" pageId="11" pageNumber="571">
<emphasis id="AEA6A94FC44C2D5BFF39FB13C5507BF7" bold="true" box="[199,308,1195,1221]" pageId="11" pageNumber="571">Remarks</emphasis>
.
<taxonomicName id="5BD20EDEC44C2D5BFEBCFB13C67F7BF7" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[322,539,1195,1221]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="ilys" status="sp. nov.">
<emphasis id="AEA6A94FC44C2D5BFEBCFB13C67F7BF7" box="[322,539,1195,1221]" italics="true" pageId="11" pageNumber="571">Saphobranchia ilys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44C2D5BFDDCFB13C63A7BF7" box="[546,606,1195,1221]" pageId="11" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="5BD20EDEC44C2D5BFD22FB13C06A7BF7" authority="(Fauchald, 1972)" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Fauchald" baseAuthorityYear="1972" box="[732,1038,1195,1221]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="micans" status="comb. nov.">
<emphasis id="AEA6A94FC44C2D5BFD22FB13C7237BF6" box="[732,839,1195,1221]" italics="true" pageId="11" pageNumber="571">S. micans</emphasis>
(
<bibRefCitation id="F84308ACC44C2D5BFCA8FB13C0617BF7" author="Fauchald, K." box="[854,1029,1195,1221]" pageId="11" pageNumber="571" pagination="1 - 575" refId="ref15175" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep waters off Western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575." type="journal article" year="1972">Fauchald, 1972</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="B5951434C44C2D5BFBEBFB13C0137BF7" box="[1045,1143,1195,1221]" pageId="11" pageNumber="571" rank="species">n. comb.</taxonomicNameLabel>
from Western
<collectingRegion id="5E16BBBFC44C2D5BFAE2FB13C1167BF7" box="[1308,1394,1195,1221]" country="Mexico" name="Mexico" pageId="11" pageNumber="571">Mexico</collectingRegion>
because they have long papillae, giving integument an hirsute appearance, by having a few sand particles along body, notochaetae far longer than body width in median chaetigers, and neurochaetae with articles longer than wide. They differ in neurochaetal features in median chaetigers. In
<taxonomicName id="5BD20EDEC44C2D5BFCF0FAAFC7367A03" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[782,850,1303,1329]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="ilys" status="sp. nov.">
<emphasis id="AEA6A94FC44C2D5BFCF0FAAFC7367A03" box="[782,850,1303,1329]" italics="true" pageId="11" pageNumber="571">S. ilys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44C2D5BFCA5FAAFC7FC7A03" box="[859,920,1303,1329]" pageId="11" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
articles are 45 times longer than wide, with a very slight reduction distally, and the anchylosed region is ½1/3 chaetal length, whereas in
<taxonomicName id="5BD20EDEC44C2D5BFB32FA83C15C7A66" authorityName="Salazar-Vallejo" authorityYear="2020" baseAuthorityName="Fauchald" baseAuthorityYear="1972" box="[1228,1336,1339,1365]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="micans">
<emphasis id="AEA6A94FC44C2D5BFB32FA83C15C7A66" box="[1228,1336,1339,1365]" italics="true" pageId="11" pageNumber="571">S. micans</emphasis>
</taxonomicName>
they are progressively smaller, about twice longer than wide, and the anchylosed region is 1/51/6 chaetal length.
</paragraph>
<paragraph id="9C6D755DC44C2D5BFF39FA3BC79A7947" blockId="11.[151,1437,151,1689]" pageId="11" pageNumber="571">
On the other hand,
<taxonomicName id="5BD20EDEC44C2D5BFE5CFA3BC5817AAF" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[418,485,1411,1437]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="ilys" status="sp. nov.">
<emphasis id="AEA6A94FC44C2D5BFE5CFA3BC5817AAF" box="[418,485,1411,1437]" italics="true" pageId="11" pageNumber="571">S. ilys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44C2D5BFE13FA3BC64E7AAF" box="[493,554,1411,1437]" pageId="11" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="5BD20EDEC44C2D5BFD57FA3BC74F7AAF" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[681,811,1411,1437]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC44C2D5BFD57FA3BC74F7AAF" box="[681,811,1411,1437]" italics="true" pageId="11" pageNumber="571">S. omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44C2D5BFCCDFA3BC7147AAF" box="[819,880,1411,1437]" pageId="11" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
especially because both have pale body wall and brownish chaetae. They differ in the development of tunic papillae, associated sediment particles, parapodial development, and chaetal size in median chaetigers. In
<taxonomicName id="5BD20EDEC44C2D5BFD33FA73C7757AD7" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[717,785,1483,1509]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="ilys" status="sp. nov.">
<emphasis id="AEA6A94FC44C2D5BFD33FA73C7757AD7" box="[717,785,1483,1509]" italics="true" pageId="11" pageNumber="571">S. ilys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44C2D5BFCE7FA73C7317AD7" box="[793,853,1483,1509]" pageId="11" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
body wall is rugose because the dorsal papillae are short, blunt mainly with fine sediment and a few sand particles, parapodia are barely projected from the body wall, and chaetae are as long as body width. On the contrary, in
<taxonomicName id="5BD20EDEC44C2D5BFCD5F9ABC7C8791F" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[811,940,1555,1581]" class="Polychaeta" family="Flabelligeridae" genus="Saphobranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="571" phylum="Annelida" rank="species" species="omorpha" status="sp. nov.">
<emphasis id="AEA6A94FC44C2D5BFCD5F9ABC7C8791F" box="[811,940,1555,1581]" italics="true" pageId="11" pageNumber="571">S. omorpha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5951434C44C2D5BFC4AF9ABC794791F" box="[948,1008,1555,1581]" pageId="11" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
body wall is pilose because the fewer dorsal papillae are without sediment particles, or fine particles adhering forming a thin cover, parapodia are markedly projected laterally, and chaetae are up to 4 times longer than body width.
</paragraph>
</subSubSection>
<subSubSection id="D4C826D6C44C2D5BFF39F9C6C70C79AB" box="[199,872,1662,1689]" pageId="11" pageNumber="571" type="distribution">
<paragraph id="9C6D755DC44C2D5BFF39F9C6C70C79AB" blockId="11.[151,1437,151,1689]" box="[199,872,1662,1689]" pageId="11" pageNumber="571">
<emphasis id="AEA6A94FC44C2D5BFF39F9C6C53D79AA" bold="true" box="[199,345,1662,1688]" pageId="11" pageNumber="571">Distribution</emphasis>
. Off Pacific
<collectingCountry id="E4C535CDC44C2D5BFE14F9C7C60279AB" box="[490,614,1663,1689]" name="Costa Rica" pageId="11" pageNumber="571">Costa Rica</collectingCountry>
, in
<quantity id="5B2AD8B8C44C2D5BFD70F9C7C77F79AB" box="[654,795,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.39" metricValueMax="1.784" metricValueMin="0.9959999999999999" pageId="11" pageNumber="571" unit="m" value="1390.0" valueMax="1784.0" valueMin="996.0">9961784 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>